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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.5.e21947" ID-PMC="PMC5740419" ID-Pensoft-Pub="1314-2828--21947" ID-PubMed="29308046" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1314-2828--e21947" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="Three new minute leaf litter beetles discovered by citizen scientists in Maliau Basin, Malaysian Borneo (Coleoptera: Leiodidae, Chrysomelidae)" checkinTime="1585357048129" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Seip, Lilian A, Otani, Sean, Suhaimi, Jadda &amp; Njunjic, Iva" docDate="2017" docId="D18D2EAA005405372FA457620E332EEF" docLanguage="en" docName="BiodivDatJour 5: e21947" docOrigin="Biodiversity Data Journal 5" docSource="http://dx.doi.org/10.3897/BDJ.5.e21947" docTitle="Clavicornaltica sabahensis Schilthuizen, Seip &amp; Otani, sp. n." docType="treatment" docVersion="3" lastPageNumber="21947" masterDocId="FFA15F5BED47FFA6FF8AC40DFFF91260" masterDocTitle="Three new minute leaf litter beetles discovered by citizen scientists in Maliau Basin, Malaysian Borneo (Coleoptera: Leiodidae, Chrysomelidae)" masterLastPageNumber="21947" masterPageNumber="21947" pageNumber="21947" updateTime="1668124493693" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new minute leaf litter beetles discovered by citizen scientists in Maliau Basin, Malaysian Borneo (Coleoptera: Leiodidae, Chrysomelidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seip, Lilian A</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Otani, Sean</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suhaimi, Jadda</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Njunjic, Iva</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>5</mods:number>
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<mods:start>21947</mods:start>
<mods:end>21947</mods:end>
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<mods:url>http://dx.doi.org/10.3897/BDJ.5.e21947</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.5.e21947</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828--21947</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:D18D2EAA005405372FA457620E332EEF" httpUri="http://treatment.plazi.org/id/D18D2EAA005405372FA457620E332EEF" lastPageNumber="21947" pageId="0" pageNumber="21947">
<subSubSection pageId="0" pageNumber="21947" type="nomenclature">
<paragraph pageId="0" pageNumber="21947">
<taxonomicName LSID="837800D6-5C3C-490F-9262-11603AA591BE" authority="Schilthuizen, Seip &amp; Otani" class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica sabahensis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="sabahensis">Clavicornaltica sabahensis Schilthuizen, Seip &amp; Otani</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="21947">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="materials_examined">
<paragraph pageId="0" pageNumber="21947">Materials</paragraph>
<paragraph pageId="0" pageNumber="21947">
<materialsCitation collectingDate="2017-09-27" collectingMethod="Winkler, litter from basis of trees" collectionCode="UMS, BORN" collectorName="Iva Njunjic; Menno Schilthuizen; Taxon Expeditions" country="Malaysia" latitude="4.7389" location="Malaysia" longitude="116.9696" pageId="0" pageNumber="21947" specimenCode="BOR / COL / 14089" specimenCount="1" specimenCount-female="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="21947">Holotype</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="21947">BOR/COL/14089</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="21947">
Iva
<normalizedToken originalValue="Njunjić">Njunjic</normalizedToken>
; Menno Schilthuizen; Taxon Expeditions
</collectorName>
; sex:
<specimenCount type="female">1 female</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="21947">adult</specimenType>
; preparations: card-mounted; disposition: in collection; Taxon: scientificName: Clavicornalticasabahensis; order: Coleoptera; family: Chrysomelidae; genus: Clavicornaltica; specificEpithet: sabahensis; taxonRank: species; scientificNameAuthorship: Schilthuizen, Seip &amp; Otani; Location: continent: Asia; island: Borneo; country:
<collectingCountry pageId="0" pageNumber="21947">Malaysia</collectingCountry>
; stateProvince: Sabah; municipality: Tongod; verbatimLocality: Maliau Basin, near Studies Centre, along Seraya Trail, where stream enters the river; verbatimElevation:
<elevation pageId="0" pageNumber="21947">260 m</elevation>
; verbatimCoordinateSystem: decimal degrees; decimalLatitude:
<geoCoordinate orientation="latitude" pageId="0" pageNumber="21947" value="4.7389">4.7389</geoCoordinate>
; decimalLongitude:
<geoCoordinate orientation="longitude" pageId="0" pageNumber="21947" value="116.9696">116.9696</geoCoordinate>
; Event: samplingProtocol:
<collectingMethod pageId="0" pageNumber="21947">Winkler, litter from basis of trees</collectingMethod>
; samplingEffort: 15 l of leaf litter; eventDate:
<collectingDate pageId="0" pageNumber="21947" value="2017-09-27">2017-09-27</collectingDate>
; habitat: lowland dipterocarp forest; fieldNumber: TxEx-MBSC1wb; Record Level: institutionID: Universiti Malaysia Sabah; collectionID: Institute for Tropical Biology and Conservation, BORNEENSIS; institutionCode:
<collectionCode pageId="0" pageNumber="21947">UMS</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="21947">BORN</collectionCode>
; basisOfRecord: PreservedSpecimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="description">
<paragraph pageId="0" pageNumber="21947">Description</paragraph>
<paragraph pageId="0" pageNumber="21947">Body dark reddish brown, small, oval and convex, ca. 0.75 mm long and ca. 0.58 mm wide (Fig. 10a). Eyes ca. 1/7 the width of the head. Antennae yellowish brown; clava long and moderately robust. Female wingless. Tibia and tarsus yellowish brown, femur dark brown and robust. Male unknown.</paragraph>
<paragraph pageId="0" pageNumber="21947">Head (Fig. 10b): Rectangular, shallowly and sparsely punctate; vertex smooth; frontal tubercles present, frontal carina absent. Eyes ca. 1/7 the width of the head in dorsal view. Antennae: clava long and moderately robust (Fig. 11).</paragraph>
<paragraph pageId="0" pageNumber="21947">
Pronotum: Lenticular in dorsal view, convex, four angles angular with a deep furrow along the length of the edge, which itself is somewhat angular in the middle with two deep seta-bearing pores: one at
<normalizedToken originalValue="¼">1/4</normalizedToken>
of the
<normalizedToken originalValue="margins">margin's</normalizedToken>
length, the other in the posterior angle. Punctuation covers the entire surface in an irregular pattern and is the same strength as the dorsal surface of the elytra.
</paragraph>
<paragraph pageId="0" pageNumber="21947">Hind wings: Absent.</paragraph>
<paragraph pageId="0" pageNumber="21947">Elytra: Striae punctiform, punctures shallower dorsally, more deeply impressed laterally and becoming less visible towards the very apex. A deep but narrow punctuated groove runs along the entire margin continuing to the apex; apex itself slightly drawn out.</paragraph>
<paragraph pageId="0" pageNumber="21947">Legs: Metafemur robust, oval, covered in fine white setae. Metatibia bearing eight minute setae which cover the terminal one-quarter along the external edge and one long spine, slightly shorter than the first tarsomere, provided with three minute teeth (Fig. 12).</paragraph>
<paragraph pageId="0" pageNumber="21947">Abdomen: Carina on the first abdominal sternite sharp and narrow.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="diagnosis">
<paragraph pageId="0" pageNumber="21947">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="21947">
Differential comparisons were made with all known species of the genus.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica sabahensis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="sabahensis">Clavicornaltica sabahensis</taxonomicName>
sp. n. differs from these in the following respects.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica buechei" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="buechei">Clavicornaltica buechei</taxonomicName>
Medvedev, 2008 (Sulawesi): is larger (1.4 mm) and the ridge on the first abdominal segment is widened posteriorly (
<bibRefCitation author="Medvedev, LN" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="0" pageNumber="21947" pagination="243 - 261" title="New species of Chrysomelidae (Coleoptera) from Sulawesi." volume="30" year="2008">Medvedev 2008</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica mizusawai" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="mizusawai">Clavicornaltica mizusawai</taxonomicName>
Suenaga &amp; Yoshida, 2016 (Taiwan) has impunctate elytra and larger eyes (
<bibRefCitation author="Suenaga, H" journalOrPublisher="Elytra, Tokyo, New Series" pageId="0" pageNumber="21947" pagination="1 - 9" title="Two New Species of the Genus Clavicornaltica (Coleoptera, Chrysomelidae, Galerucinae) from Taiwan and Ishigaki-jima Island, Japan" volume="6" year="2016">Suenaga and Yoshida 2016</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica sakishimana" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="sakishimana">Clavicornaltica sakishimana</taxonomicName>
Suenaga &amp; Yoshida, 2016 (Japan) has the carina on the first abdominal sternite widened caudally (
<bibRefCitation author="Suenaga, H" journalOrPublisher="Elytra, Tokyo, New Series" pageId="0" pageNumber="21947" pagination="1 - 9" title="Two New Species of the Genus Clavicornaltica (Coleoptera, Chrysomelidae, Galerucinae) from Taiwan and Ishigaki-jima Island, Japan" volume="6" year="2016">Suenaga and Yoshida 2016</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica pusilla" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="pusilla">Clavicornaltica pusilla</taxonomicName>
Scherer, 1974 and
<taxonomicName lsidName="C. loebli" pageId="0" pageNumber="21947" rank="species" species="loebli">C. loebli</taxonomicName>
Scherer, 1974 (Sri Lanka) have impunctate elytra (
<bibRefCitation author="Scherer, G" journalOrPublisher="Revue Suisse de Zoologie." pageId="0" pageNumber="21947" pagination="57 - 68" title="Clavicornaltica a new genus from Ceylon (Coleoptera-Chrysomelidae-Alticinae)" volume="81" year="1974">Scherer 1974</bibRefCitation>
).
<taxonomicName lsidName="C. besucheti" pageId="0" pageNumber="21947" rank="species" species="besucheti">C. besucheti</taxonomicName>
Scherer, 1974 (Sri Lanka) is larger, 1.5-2.2 mm (
<bibRefCitation author="Scherer, G" journalOrPublisher="Revue Suisse de Zoologie." pageId="0" pageNumber="21947" pagination="57 - 68" title="Clavicornaltica a new genus from Ceylon (Coleoptera-Chrysomelidae-Alticinae)" volume="81" year="1974">Scherer 1974</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica malayana" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="malayana">Clavicornaltica malayana</taxonomicName>
Medvedev, 1996 (Malaysia) has the pronotum impunctate (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica iriana" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="subspecies" species="iriana" subspecies="iriana">Clavicornaltica iriana iriana</taxonomicName>
Medvedev, 1996 (New Guinea) is larger (1.2-1.4 mm), has larger eyes, only 4-5 rows of punctures on the elytra and these do not continue to the apex (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica iriana" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="subspecies" species="iriana" subspecies="sarawacensis">Clavicornaltica iriana sarawacensis</taxonomicName>
Medvedev, 1996 (Borneo) is larger (1.2-1.4 mm) and the elytral punctures are only visible laterally (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica takizawai" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="takizawai">Clavicornaltica takizawai</taxonomicName>
Doeberl, 2009 (Nepal) is larger (1.45 mm) and its frons and vertex are densely punctuated (
<bibRefCitation author="Doeberl, M." journalOrPublisher="VERNATE" pageId="0" pageNumber="21947" pagination="413 - 420" title="Contribution to the knowledge of Alticinae from Nepal (Insecta: Coleoptera: Chrysomelidae)" volume="28" year="2009">
<normalizedToken originalValue="Döberl">Doeberl</normalizedToken>
2009
</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica mussardi" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="mussardi">Clavicornaltica mussardi</taxonomicName>
Scherer 1974 (Sri Lanka) is larger (1.3-1.5 mm) (
<bibRefCitation author="Scherer, G" journalOrPublisher="Revue Suisse de Zoologie." pageId="0" pageNumber="21947" pagination="57 - 68" title="Clavicornaltica a new genus from Ceylon (Coleoptera-Chrysomelidae-Alticinae)" volume="81" year="1974">Scherer 1974</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica rileyi" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="rileyi">Clavicornaltica rileyi</taxonomicName>
<normalizedToken originalValue="Döberl">Doeberl</normalizedToken>
, 2002 (India) is larger (1.5 mm) and its eyes are larger (
<bibRefCitation author="Doeberl, M." journalOrPublisher="Bonner zoologische Beitra ̈ ge" pageId="0" pageNumber="21947" pagination="297 - 304" title="Alticinae from India and Pakistan Stored in the Collection of the Texas A. &amp; M. University, U. S. A. (Coleoptera, Chrysomelidae)" volume="51" year="2002">
<normalizedToken originalValue="Döberl">Doeberl</normalizedToken>
2002
</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica dali" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="dali">Clavicornaltica dali</taxonomicName>
Konstantinov &amp; Duckett, 2005 (Yunnan): punctures on the head are stronger and those on the pronotum weaker than in
<taxonomicName lsidName="C. sabahensis" pageId="0" pageNumber="21947" rank="species" species="sabahensis">C. sabahensis</taxonomicName>
sp. n. (
<bibRefCitation author="Konstantinov, Alexander S." journalOrPublisher="Zootaxa" pageId="0" pageNumber="21947" pagination="49 - 64" title="New species of Clavicornaltica Scherer (Coleoptera: Chrysomelidae) from continental Asia" volume="1037" year="2005">Konstantinov and Duckett 2005</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica tarsalis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="tarsalis">Clavicornaltica tarsalis</taxonomicName>
Medvedev, 1996 (Irian Jaya) is larger, 1.6 mm and the ridge on the first abdominal sternite is anteriorly broadly widened (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica australis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="australis">Clavicornaltica australis</taxonomicName>
Konstantinov, 1996 (Queensland) lacks the long seta in the middle of the pronotal edge (
<bibRefCitation author="Konstantinov, Alexander S." journalOrPublisher="Zootaxa" pageId="0" pageNumber="21947" pagination="49 - 64" title="New species of Clavicornaltica Scherer (Coleoptera: Chrysomelidae) from continental Asia" volume="1037" year="2005">Konstantinov and Duckett 2005</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica fortepunctata" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="fortepunctata">Clavicornaltica fortepunctata</taxonomicName>
Scherer, 1974 (Sri Lanka): elytral punctuation fades abruptly before the apex (
<bibRefCitation author="Scherer, G" journalOrPublisher="Revue Suisse de Zoologie." pageId="0" pageNumber="21947" pagination="57 - 68" title="Clavicornaltica a new genus from Ceylon (Coleoptera-Chrysomelidae-Alticinae)" volume="81" year="1974">Scherer 1974</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica trautneri" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="trautneri">Clavicornaltica trautneri</taxonomicName>
Medvedev, 1993 (Philippines) is much larger (2.1 mm) (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica takimotoi" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="takimotoi">Clavicornaltica takimotoi</taxonomicName>
Lesage, 1997 (Taiwan) is more globular and the punctures on the dorsum of the elytra are nearly invisible (
<bibRefCitation author="Suenaga, H" journalOrPublisher="Elytra, Tokyo, New Series" pageId="0" pageNumber="21947" pagination="1 - 9" title="Two New Species of the Genus Clavicornaltica (Coleoptera, Chrysomelidae, Galerucinae) from Taiwan and Ishigaki-jima Island, Japan" volume="6" year="2016">Suenaga and Yoshida 2016</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica philippinensis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="philippinensis">Clavicornaltica philippinensis</taxonomicName>
Scherer, 1979 (Philippines) is larger (1.3 mm) (
<bibRefCitation author="Medvedev, L. N." journalOrPublisher="Entomofauna, Zeitschrift fuer Entomologie" pageId="0" pageNumber="21947" pagination="137 - 148" title="Review of the flea-beetle genus Clavicornaltica (Coleoptera: Chrysomelidae, Alticinae)" volume="17" year="1996">Medvedev 1996</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica tamdao" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="tamdao">Clavicornaltica tamdao</taxonomicName>
Konstantinov &amp; Duckett, 2005 (Vietnam): terminal setae on the metatibia are more numerous (12 in
<taxonomicName lsidName="C. tamdao" pageId="0" pageNumber="21947" rank="species" species="tamdao">C. tamdao</taxonomicName>
versus 8 in
<taxonomicName lsidName="C. sabahensis" pageId="0" pageNumber="21947" rank="species" species="sabahensis">C. sabahensis</taxonomicName>
sp. n.) and cover a larger section of the tibia (the terminal one third) (
<bibRefCitation author="Konstantinov, Alexander S." journalOrPublisher="Zootaxa" pageId="0" pageNumber="21947" pagination="49 - 64" title="New species of Clavicornaltica Scherer (Coleoptera: Chrysomelidae) from continental Asia" volume="1037" year="2005">Konstantinov and Duckett 2005</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica vietnamensis" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="species" species="vietnamensis">Clavicornaltica vietnamensis</taxonomicName>
Konstantinov &amp; Duckett, 2005 (Vietnam): apex of the elytra is less extended (
<bibRefCitation author="Konstantinov, Alexander S." journalOrPublisher="Zootaxa" pageId="0" pageNumber="21947" pagination="49 - 64" title="New species of Clavicornaltica Scherer (Coleoptera: Chrysomelidae) from continental Asia" volume="1037" year="2005">Konstantinov and Duckett 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="etymology">
<paragraph pageId="0" pageNumber="21947">Etymology</paragraph>
<paragraph pageId="0" pageNumber="21947">
Since this is the first species of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="genus">Clavicornaltica</taxonomicName>
found in Sabah, the specific epithet
<taxonomicName lsidName="sabahensis" pageId="0" pageNumber="21947" rank="species" species="sabahensis">sabahensis</taxonomicName>
(&quot;inhabitant of Sabah&quot;) was chosen. This was one of several names suggested during a naming ceremony in Maliau Basin Studies Centre on 6 October 2017, in which expedition participants as well as a large number of field centre staff and porters took part.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="distribution">
<paragraph pageId="0" pageNumber="21947">Distribution</paragraph>
<paragraph pageId="0" pageNumber="21947">Known only from one location in the valley where the Maliau river flows out of Maliau Basin, at 260 m elevation (Maliau Basin Studies Centre).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="21947" type="taxon discussion">
<paragraph pageId="0" pageNumber="21947">Taxon discussion</paragraph>
<paragraph pageId="0" pageNumber="21947">
Only a single female was at the authors' disposal. Nonetheless, the authors felt confident that this specimen represents an undescribed species. First of all, given their small size, apterism/brachypterism, and habitat (leaf litter in forests), it is unlikely that
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Clavicornaltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clavicornaltica" order="Coleoptera" pageId="0" pageNumber="21947" phylum="Arthropoda" rank="genus">Clavicornaltica</taxonomicName>
species have wide ranges (
<bibRefCitation author="Konstantinov, Alexander S." journalOrPublisher="Zootaxa" pageId="0" pageNumber="21947" pagination="49 - 64" title="New species of Clavicornaltica Scherer (Coleoptera: Chrysomelidae) from continental Asia" volume="1037" year="2005">Konstantinov and Duckett 2005</bibRefCitation>
) and the only species known from Borneo and its immediate vicinity have external characters that do not match
<taxonomicName lsidName="C. sabahensis" pageId="0" pageNumber="21947" rank="species" species="sabahensis">C. sabahensis</taxonomicName>
sp. n. (see under Diagnosis). Moreover,
<taxonomicName lsidName="C. sabahensis" pageId="0" pageNumber="21947" rank="species" species="sabahensis">C. sabahensis</taxonomicName>
sp. n. carries a combination of traits that make it easily recognisable. Specifically, these are its extremely small size, highly vaulted shape, dark colouration, narrow ridge on the first abdominal sternite, angular, thick pronotal margin, and strong punctuation on the entire elytra.
</paragraph>
</subSubSection>
</treatment>
</document>