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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.235.99140" ID-Pensoft-Pub="1314-2003-235-137" ID-Pensoft-UUID="5C03056135C05D05BC1818F226325D31" ModsDocID="1314-2003-235-137" checkinTime="1700246633984" checkinUser="pensoft" docAuthor="Yahara, Tetsukazu, Hirota, Shun K., Fujii, Seiko, Kokami, Yasushi, Fuse, Kengo, Sato, Hiroyuki, Tagane, Shuichiro &amp; Suyama, Yoshihisa" docDate="2023" docId="78121A0F0DA55E8A81A3FDD7731470E3" docLanguage="en" docName="PhytoKeys 235: 137-187" docOrigin="PhytoKeys 235" docPubDate="2023-11-17" docSource="http://dx.doi.org/10.3897/phytokeys.235.99140" docTitle="Hosta takiminazukiflora Se. Fujii &amp; Yahara 2023, sp. nov." docType="treatment" docVersion="1" id="5C03056135C05D05BC1818F226325D31" lastPageNumber="137" masterDocId="5C03056135C05D05BC1818F226325D31" masterDocTitle="Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments" masterLastPageNumber="187" masterPageNumber="137" pageNumber="137" updateTime="1700246633984" updateUser="pensoft">
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<mods:title id="4D4BAF25052510FC14B99A24C4870900">Molecular phylogeny and taxonomy of Hosta (Asparagaceae) on Shikoku Island, Japan, including five new species, one new subspecies, and two new status assignments</mods:title>
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<mods:name id="B8796961BF93BF6B68A8D73FCE973733" type="personal">
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<mods:namePart id="C18DA5B49042C0D66B431E65989B519A">Yahara, Tetsukazu</mods:namePart>
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<mods:affiliation id="B938A76B9BBD2ADFD5FDD6DC334DC0B1">Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
<mods:nameIdentifier id="70CDCE4CA1D2D6C800E8B0D8DFDC1CBD" type="email">tet.yahara@gmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="44DAE8805380134272556120499CF996">Hirota, Shun K.</mods:namePart>
<mods:nameIdentifier id="3088D61918CA7E5A03DEB523096923BC" type="ORCID">https://orcid.org/0000-0002-6104-1119</mods:nameIdentifier>
<mods:affiliation id="2F3B29682B3F676CC240662B707FD883">Field Science Center, Graduate School of Agricultural Science, Tohoku University, 232 - 3 Yomogida, Naruko-onsen, Osaki, Miyagi 989 - 6711, Japan &amp; Botanical Gardens, Osaka Metropolitan University, 2000 Kisaichi, Katano, Osaki 576 - 0004, Japan</mods:affiliation>
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<mods:namePart id="334D284CFE33768B81E64E4258098DA6">Fujii, Seiko</mods:namePart>
<mods:affiliation id="1C44E19776DAE0E680D178839E077845">The Kochi Prefectural Makino Botanical Garden, Godai-san 4200 - 6, Kochi 781 - 8125, Japan</mods:affiliation>
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<mods:namePart id="05B39F6E387C98CE5E25991787DC309C">Kokami, Yasushi</mods:namePart>
<mods:affiliation id="13859929F023A658CB992D2324B91D2E">The Kochi Prefectural Makino Botanical Garden, Godai-san 4200 - 6, Kochi 781 - 8125, Japan</mods:affiliation>
</mods:name>
<mods:name id="4A524643992681A8FA5CCD7ABD3A2F26" type="personal">
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<mods:namePart id="7C2E38A7C2DAA44A3230193AF261EC92">Fuse, Kengo</mods:namePart>
<mods:affiliation id="446D6EE18070B520D78E0E6127D277F5">Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="5962BF99783C4F0FEE6B816D3D92B541">Sato, Hiroyuki</mods:namePart>
<mods:nameIdentifier id="3C1369407059A38F53D8B9A480AB3F5F" type="ORCID">https://orcid.org/0000-0001-6003-6492</mods:nameIdentifier>
<mods:affiliation id="9722573AC4FC134C5985F86835A0B053">Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="3445C53C22600FDBE4D0611124111D34">Tagane, Shuichiro</mods:namePart>
<mods:nameIdentifier id="8F22F8555B2A50B0D5495F695390A000" type="ORCID">https://orcid.org/0000-0002-1974-7329</mods:nameIdentifier>
<mods:affiliation id="56B14444E3BB09253D54BB83D27751CE">The Kagoshima University Museum, Kagoshima University, 1 - 21 - 30 Korimoto, Kagoshima, 890 - 0065, Japan</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B88F528D483E06D2B47E4521A6EA19F2">Suyama, Yoshihisa</mods:namePart>
<mods:nameIdentifier id="C3359DD0378D573C1E9E020B7EE14FE1" type="ORCID">https://orcid.org/0000-0002-3136-5489</mods:nameIdentifier>
<mods:affiliation id="BEB2618EC21BCBD50B2295DD2EFA0006">Field Science Center, Graduate School of Agricultural Science, Tohoku University, 232 - 3 Yomogida, Naruko-onsen, Osaki, Miyagi 989 - 6711, Japan</mods:affiliation>
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<mods:typeOfResource id="6EE29BD747ABCC25EEDBDFBE9B0A7F50">text</mods:typeOfResource>
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<mods:title id="1BD6EECCCCB61E553CB9A3C281514A25">PhytoKeys</mods:title>
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<mods:part id="EC67E62831EF18776E5CEF24678637BD">
<mods:date id="6DAABA35F712AE3DFA43C023A149AD39">2023</mods:date>
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<mods:number id="44FA6300AF21713799504DB669D73B56">2023-11-17</mods:number>
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<mods:number id="D6DF65AAD48ACDBAFA3D155CF72379C2">235</mods:number>
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<mods:start id="1546E2594E7462211DDEFF280AEF9973">137</mods:start>
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<mods:url id="16A38CE2D0E914159F188A550B6B53A2">http://dx.doi.org/10.3897/phytokeys.235.99140</mods:url>
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<mods:classification id="5A2E5A3D81D2CCB525AEEF33DC76FEED">journal article</mods:classification>
<mods:identifier id="6A452B2C2CA707EEC60BD5951ED63A2A" type="DOI">http://dx.doi.org/10.3897/phytokeys.235.99140</mods:identifier>
<mods:identifier id="3593E94DF8E5132EC8C4D57191B574D1" type="Pensoft-Pub">1314-2003-235-137</mods:identifier>
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<treatment id="78121A0F0DA55E8A81A3FDD7731470E3" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3" httpUri="http://treatment.plazi.org/id/78121A0F0DA55E8A81A3FDD7731470E3" lastPageNumber="137" pageId="0" pageNumber="137">
<subSubSection id="9FF844D46C794D1701AE96C8AAE35A11" pageId="0" pageNumber="137" type="nomenclature">
<paragraph id="93674AD8D1952C1AED251BD2A0E504C3" pageId="0" pageNumber="137">
6.
<taxonomicName id="098EA66C0CC515017DD4EC98DA3358B6" LSID="78121A0F-0DA5-5E8A-81A3-FDD7731470E3" authority="Se. Fujii &amp; Yahara" authorityName="Se. Fujii &amp; Yahara" authorityYear="2023" class="Liliopsida" family="Asparagaceae" genus="Hosta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hosta takiminazukiflora" order="Asparagales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="takiminazukiflora" status="sp. nov.">Hosta takiminazukiflora Se.Fujii &amp; Yahara</taxonomicName>
<taxonomicNameLabel id="8BA6C062159F466D7F2F57724D21561A" pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B1DE19799416E745E4A896779A10EC3B" pageId="0" pageNumber="137" type="description">
<paragraph id="10DBC8931E27960B03E47D9B3E248184" pageId="0" pageNumber="137">
<figureCitation id="732737F6B41026031E0DC079F14F2AEB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The type specimen of H. takiminazukiflora: N. Inagaki et al. FOK- 080097 (holotype MBK)." figureDoi="10.3897/phytokeys.235.99140.figure14" httpUri="https://binary.pensoft.net/fig/938515" pageId="0" pageNumber="137">Figs 14</figureCitation>
<figureCitation id="337705C3F8A68652B927778E320EEF4A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. A flowering plant of H. takiminazukiflora subsp. takiminazukiflora (JPN 6481, 24 Jun. 2021). Scale bars: 5 cm (A); 2 cm (B)." figureDoi="10.3897/phytokeys.235.99140.figure15" httpUri="https://binary.pensoft.net/fig/938516" pageId="0" pageNumber="137">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="80F99A2355420F7D2E1BCFF852E96DBD" pageId="0" pageNumber="137" type="diagnosis">
<paragraph id="63BCEC991483F7383AB5F6DE294E3A7D" pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph id="F4E4EDE8CFA58790D93E1CF91312DBBC" pageId="0" pageNumber="137">
<taxonomicName id="EAFFE4894EE6C5DE217ABD85FFB4DA83" authorityName="Se. Fujii &amp; Yahara" authorityYear="2023" class="Liliopsida" family="Asparagaceae" genus="Hosta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hosta takiminazukiflora" order="Asparagales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="takiminazukiflora">
<emphasis id="7CEFAB37E3C845A32A914ED614CB7979" italics="true" pageId="0" pageNumber="137">Hosta takiminazukiflora</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="E626AE2E0BDC53FAB0D130C3BBF58F39" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. longipedicellata" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="longipedicellata">
<emphasis id="29B6393E93FF7345DCCB357BFA03617A" italics="true" pageId="0" pageNumber="137">H. longipedicellata</emphasis>
</taxonomicName>
and
<taxonomicName id="9C1BBA64203271E9FFEDE24312F1A4C4" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. minazukiflora" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="minazukiflora">
<emphasis id="8EA20D746D870524ADA4B862678460EE" italics="true" pageId="0" pageNumber="137">H. minazukiflora</emphasis>
</taxonomicName>
. It is distinguished from
<taxonomicName id="EDF37DAEC6A7F77EE9283C8E5BC982CF" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. longipedicellata" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="longipedicellata">
<emphasis id="610B860F1434C22C1D0E82EF3B211058" italics="true" pageId="0" pageNumber="137">H. longipedicellata</emphasis>
</taxonomicName>
by leaves smooth on the lower surface (vs. papillose), pedicels 1.4-2.5 cm long (compared to 2.5-3.3 cm long), and flower bracts being fresh during flowering (in contrast to being withering), and from
<taxonomicName id="D748DF3DDD98FB4930A67B415391A56F" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. minazukiflora" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="minazukiflora">
<emphasis id="9FB23619C428F211A21F5CDA9C828985" italics="true" pageId="0" pageNumber="137">H. minazukiflora</emphasis>
</taxonomicName>
by perianths with a distinct midvein (as opposed to three distinct veins) and pedicels 1.4-2.5 cm long (compared to 1.1-1.2 cm long).
</paragraph>
<caption id="0A5E3F6644F83EAA809D5F2C28617FD2" doi="10.3897/phytokeys.235.99140.figure14" httpUri="https://binary.pensoft.net/fig/938515" pageId="0" pageNumber="137" start="Figure 14" startId="F14">
<paragraph id="949FBEADEB83BA18A9141D0C3FF586F8" pageId="0" pageNumber="137">
<emphasis id="0591861A735A8290641198A94A0CCD42" bold="true" pageId="0" pageNumber="137">Figure 14.</emphasis>
The type specimen of
<emphasis id="1B500568FC4CC95A769AC9FF5E013ADE" italics="true" pageId="0" pageNumber="137">
<taxonomicName id="7002CD308518A6E0BA50117A7A43804F" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. takiminazukiflora" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="takiminazukiflora">H. takiminazukiflora</taxonomicName>
: N. Inagaki et al. FOK-080097
</emphasis>
(holotype MBK).
</paragraph>
</caption>
<caption id="FC1CBE88B419E32F4A645BFBA57E43B5" doi="10.3897/phytokeys.235.99140.figure15" httpUri="https://binary.pensoft.net/fig/938516" pageId="0" pageNumber="137" start="Figure 15" startId="F15">
<paragraph id="70BECFB426D0657D65EEE8052F82E28D" pageId="0" pageNumber="137">
<emphasis id="DFEFE475675982E1C99AFF62AFF01707" bold="true" pageId="0" pageNumber="137">Figure 15.</emphasis>
A flowering plant of
<taxonomicName id="D4B7A37FEF393B5834607BB541F23BC8" authority="subsp. takiminazukiflora" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. takiminazukiflora subsp. takiminazukiflora" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="subSpecies" species="takiminazukiflora" subSpecies="takiminazukiflora">H. takiminazukiflora subsp. takiminazukiflora</taxonomicName>
(JPN6481, 24 Jun. 2021). Scale bars: 5 cm (
<emphasis id="85A5A1132401931558E220ED533C10BF" bold="true" pageId="0" pageNumber="137">A</emphasis>
); 2 cm (
<emphasis id="37EF8143964E0FF1D52F7E2DE568157A" bold="true" pageId="0" pageNumber="137">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="79F7D7981DC82381F7B8F226AB24F7E6" pageId="0" pageNumber="137" type="materials_examined">
<paragraph id="F6B585E639178156E3DE156816ED704A" pageId="0" pageNumber="137">
<typeStatus id="23C2AEDA0D2C931D2D5FFA9BB7D35786">Type</typeStatus>
.
</paragraph>
<paragraph id="45DA9966E0CF67B276D9CE318D2D268A" pageId="0" pageNumber="137">
<materialsCitation id="F3FF7E227D2324E5E2494DFECDBAC7BA" collectingDate="2007-07-22" collectorName="N. Inagaki" country="Japan" county="Tosa County" location="Mt. Higashikado-yama" municipality="Ookawa village" specimenCount="1" stateProvince="Kochi Pref." typeStatus="holotype">
<emphasis id="2F543D0F611ACFC4E3EB66B1C217AB90" bold="true" pageId="0" pageNumber="137">
<collectingCountry id="1B0100DDEB70F0B7B18872ECF9890D37" name="Japan">Japan</collectingCountry>
.
<collectingRegion id="F2ADF050BB474BB91F5628B1A9F2E86D" country="Japan" name="Kochi">Kochi Pref.</collectingRegion>
</emphasis>
:
<collectingCounty id="6E33B123589B9E2E7287AABD979738B2">Tosa County</collectingCounty>
,
<collectingMunicipality id="E302B2D2B2940253B148DB6E70D10047">Ookawa village</collectingMunicipality>
,
<location id="52FB914F74E2074217C6A6E163208A29" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3:52FB914F74E2074217C6A6E163208A29" country="Japan" county="Tosa County" municipality="Ookawa village" name="Mt. Higashikado-yama" stateProvince="Kochi Pref.">Mt. Higashikado-yama</location>
,
<collectingDate id="C3579A9B4A5D2CDBF35936689ECFFF7C" value="2007-07-22">22 Jul. 2007</collectingDate>
, with flowers,
<emphasis id="4819214EDBD0C3F8C72D26B99CDD316A" italics="true" pageId="0" pageNumber="137">
<collectorName id="88F3119BA2C64F5E3ABA1EFAA41432B7">N. Inagaki</collectorName>
et al. FOK-080097
</emphasis>
with flowers (
<typeStatus id="9F0D2BBA58A72F623ACAAD4C1D0E9756">holotype</typeStatus>
MBK0189902!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0E6221A76AF6DA73D5380713E55B96C8" pageId="0" pageNumber="137" type="description">
<paragraph id="8DBCC5E827CCC8E7141EFDB9D18E85BD" pageId="0" pageNumber="137">Description.</paragraph>
<paragraph id="7D8979F7EEE6A672D88A33A02053B8F9" pageId="0" pageNumber="137">Herbs perennial. Leaves basal, spiral, long petiolate, 3-12 per ramet; blades oblong-ovate, 11-26.5 cm long, 3.6-10.5 cm, 2.1-3.3 times longer than wide, thinly papery, glabrous on both surfaces, base cuneate to obtuse, often decurrent, apex long acuminate, margin entire, veins in 6-10 pairs, smooth on the lower surface; petioles 3.4-22 cm long, narrowly winged, wing 0.1-0.6 cm wide, glabrous. Scape 8.6-28.5 cm long, terete. Raceme 5.9-18.5 cm long, 4-13-flowered; flower bracts vivid (not withering) in anthesis, erect, purplish light green, light bluish purple, ovate oblong-lanceolate, boat-shaped, 2-4.7 cm long, 0.2-0.7 cm wide, papery, glabrous, apex acute to acuminate. Flowers not fragrant; pedicels 1.4-2.5 cm long, glabrous; perianths 4-5.7 cm long, funnel-form, pale white-purple to light bluish-purple outside, almost white to light pale purple inside, midveins more or less purplish, glabrous, 6-lobed; tube dilated from apical 1/2, lobes triangular, 1.2-1.8 cm long, 0.5-1 cm wide, apex acute. Stamens 6, same or ca. 0.5 cm exerted from perianths; filaments 3.9-5.8 cm long, upwardly curved at the distal part, white, free, glabrous, anthers purple when fresh, dark blue-grey when dried, 3 mm long. Ovary ellipsoid, 0.6-0.7 cm long, style 4.5-6.5 cm long, upwardly curved at the distal part, up to 1 cm exerted from perianth, glabrous, stigma capitate. Young capsules 2.8 cm long (for MBK0087737).</paragraph>
</subSubSection>
<subSubSection id="BED4FC6A95F8B72997FE2DE1CF61F5DA" pageId="0" pageNumber="137" type="phenology">
<paragraph id="18A491696C4D8C75FF669560A270ED01" pageId="0" pageNumber="137">Phenology.</paragraph>
<paragraph id="1424FC914B23DC85A39ED14CA1F4FE5A" pageId="0" pageNumber="137">Flowering in late June to early August.</paragraph>
</subSubSection>
<subSubSection id="E44521D92F44B2540F8F1A106FF6D0C0" pageId="0" pageNumber="137" type="distribution">
<paragraph id="56BE72A151B07F1CE9FD038CD1DBC89E" pageId="0" pageNumber="137">Distribution and habitat.</paragraph>
<paragraph id="E693F981F92E253C6B34BEB91635AC46" pageId="0" pageNumber="137">Japan (Kochi Prefecture: Tosa County, endemic to Mt. Inamura, Mt. Higashikado, and the surrounding area). It grows on cliffs.</paragraph>
</subSubSection>
<subSubSection id="2A72EC74CB074BD361257C070DE02FD9" pageId="0" pageNumber="137" type="etymology">
<paragraph id="039A2A72355229A08EB26917280DB5F2" pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph id="242AEEA1A98E7CF84920BDAAE1054D84" pageId="0" pageNumber="137">
A specific epithet is derived from its habit of growing on rock cliffs (called
<normalizedToken id="6533D78B277BB6E913D3245A98BC5904" originalValue="taki">'taki'</normalizedToken>
in Kochi dialect) and flowering in June (Minazuki).
</paragraph>
</subSubSection>
<subSubSection id="BDF150A3C27295CCD5062CD71BB1D881" pageId="0" pageNumber="137" type="conservation status">
<paragraph id="6D2EDEB61C292E182A1FD8F12E4B524B" pageId="0" pageNumber="137">Conservation status.</paragraph>
<paragraph id="BCD3BB999A6F4C3B73E199A662B16010" pageId="0" pageNumber="137">
Using criterion D1 for IUCN Red List categories (
<bibRefCitation id="F4412F85143A9209DABBFD416B3FB06D" author="Huson, DH" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="137" publicationUrl="https://portals.iucn.org/library/node/10315" refId="B13" refString="2012. . https://portals.iucn.org/library/node/10315" url="https://portals.iucn.org/library/node/10315" year="2012">IUCN 2012</bibRefCitation>
,
<bibRefCitation id="388B8898E0EA2A9AE059EF19D541331D" author="Huson, DH" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="137" publicationUrl="https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" refId="B14" refString="2022. . https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" url="https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" year="2022">IUCN 2022</bibRefCitation>
), we recommend that this species be classified as VU (Vulnerable) because its population is estimated to be between 250 and 1000.
</paragraph>
</subSubSection>
<subSubSection id="6DAD7C878F280028DF5A5698D242DC8B" pageId="0" pageNumber="137" type="japanese name">
<paragraph id="AE57FA7721ED027F0D475362C9D47C6F" pageId="0" pageNumber="137">Japanese name.</paragraph>
<paragraph id="A32B81528C660DCCB3C4F48C05A329F4" pageId="0" pageNumber="137">Taki-minazuki-giboshi (new).</paragraph>
</subSubSection>
<subSubSection id="7C9B9BF9C99670D6344569E071E4EAF8" pageId="0" pageNumber="137" type="materials_examined">
<paragraph id="D4B31A21D90328EBCFC7B7A8DC6F8760" pageId="0" pageNumber="137">Additional specimens examined.</paragraph>
<paragraph id="488BF9C51DC5A9031A4CE67F5516ECAA" pageId="0" pageNumber="137">
<materialsCitation id="3E383CD5044B84524FC5EB3418A197DA" accessionNumber="JPN15229" collectingDate="2004-07-27" collectingDateMax="2022-07-23" collectingDateMin="2004-07-27" collectorName="N. Inagaki, T. Yahara, Agawa-gun, Ino-cho, Y. Oohira, Se. Fujii" country="Japan" county="Tosa County" elevation="900" location="Tosa County" municipality="Ookawa village" specimenCount="1" stateProvince="Kochi Pref.">
<emphasis id="2B2FBA7319BB68F6781782125212211B" bold="true" pageId="0" pageNumber="137">
<collectingCountry id="0B3B51769948C31646E0138E0628BAB8" name="Japan">Japan</collectingCountry>
.
<collectingRegion id="C1FA73F941FAA75733003BE660AFF9B7" country="Japan" name="Kochi">Kochi Pref.</collectingRegion>
</emphasis>
:
<collectingCounty id="2B07C292DA903EC3B057806752BA6CD5">Tosa County</collectingCounty>
,
<collectingMunicipality id="38E124E3128F99FE2F30C8D8D1E65474">Ookawa village</collectingMunicipality>
,
<quantity id="9128973E8AE292BCD442FD635EB9A6C6" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation id="FF61AF67F9AE024D533F4D0E0B81D69C" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<collectingDate id="EA43F9DBB1AFF8101421440F2BD3A57F" value="2004-07-27">27 Jul. 2004</collectingDate>
,
<emphasis id="4E2A00EA42D188B400FFFAEA038F8942" italics="true" pageId="0" pageNumber="137">
<collectorName id="FBBD810CC87E48995660EB29DBDEBBE9">N. Inagaki</collectorName>
et al. FOK-067742
</emphasis>
with flowers and young fruits (MBK0087737!);
<location id="2FC1A7850E242F7992C082D627A7AEE1" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3:2FC1A7850E242F7992C082D627A7AEE1" country="Japan" county="Tosa County" municipality="Ookawa village" name="Tosa County" stateProvince="Kochi Pref.">Tosa County</location>
,
<location id="D115E2DF7F316980B038DCBE6C8215F4" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3:D115E2DF7F316980B038DCBE6C8215F4" country="Japan" county="Tosa County" municipality="Ookawa village" name="Tosa Town" stateProvince="Kochi Pref.">Tosa Town</location>
,
<quantity id="C22F8DB07DEC85B18C65303B94CA3EB2" metricMagnitude="2" metricUnit="m" metricValue="6.2" unit="m" value="620.0">
<elevation id="5F2F72B20ACC12D5DD3E01EAD18642AA" metricMagnitude="2" metricUnit="m" metricValue="6.2" unit="m" value="620.0">620 m</elevation>
</quantity>
elev.,
<collectingDate id="7732D08DDDE1378D9262D05295AABB97" value="2021-06-24">24 Jun. 2021</collectingDate>
,
<emphasis id="7E1106BDBC8B8AE6AC3E288BD1AF2DAE" italics="true" pageId="0" pageNumber="137">
<collectorName id="B5773D74C7B8814A4DFC7F564756FA5B">T. Yahara</collectorName>
et al. JPN6478
</emphasis>
-
<emphasis id="FBB54F001442896AB593BA985897C206" italics="true" pageId="0" pageNumber="137">6486</emphasis>
,
<emphasis id="ECD27E2FBACE4D81C2B9E0C2564A69F1" italics="true" pageId="0" pageNumber="137">6490</emphasis>
with flowers (FU!);
<collectorName id="23FDF4D5A39469D5C0B13CB3DDFE5E79">Agawa-gun</collectorName>
,
<collectorName id="C5F4917345468740C5860E56E891DF24">Ino-cho</collectorName>
,
<location id="3063925DB770C4F91C65032E72F7766E" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3:3063925DB770C4F91C65032E72F7766E" country="Japan" county="Tosa County" municipality="Ookawa village" name="Mt. Inamura" stateProvince="Kochi Pref.">Mt. Inamura</location>
,
<quantity id="31FEF7B41A185CCF07A36150087B2F3B" metricMagnitude="3" metricUnit="m" metricValue="1.39" unit="m" value="1390.0">
<elevation id="AA0A479129FE65F3ABE636113D2A9F0E" metricMagnitude="3" metricUnit="m" metricValue="1.39" unit="m" value="1390.0">1390 m</elevation>
</quantity>
elev.,
<collectingDate id="4E368414AC58BA8B84C1655D02BEA350" value="2019-08-02">2 Aug. 2019</collectingDate>
,
<emphasis id="797200D2DA8864B66E5DD2A8E67E9B71" italics="true" pageId="0" pageNumber="137">
<collectorName id="F8E4AD602A71967979F59CE0715504E5">Y. Oohira</collectorName>
14695
</emphasis>
with flowers (MBK0314655!);
<collectorName id="D721465AB5B260571D683FEA8F11C6AA">Agawa-gun</collectorName>
,
<collectorName id="2A798927BBD01F879E3E1E8ED517768E">Ino-cho</collectorName>
, north cliff of
<location id="B1EBE41F222D6289D5E717210ADC768F" LSID="urn:lsid:plazi:treatment:78121A0F0DA55E8A81A3FDD7731470E3:B1EBE41F222D6289D5E717210ADC768F" country="Japan" county="Tosa County" municipality="Ookawa village" name="Mt. Inamura" stateProvince="Kochi Pref.">Mt. Inamura</location>
,
<collectingDate id="A353D26284432428CD71725321C0466E" value="2022-07-23">23 Jul. 2022</collectingDate>
,
<emphasis id="0D4C618F54D7A2CB843C29FC18A5099C" italics="true" pageId="0" pageNumber="137">
<collectorName id="90CDA7E99489197925ADF0106C802657">Se. Fujii</collectorName>
<accessionNumber id="CEE84A5F5BAA828CCDB61B4816B0ACDF" httpUri="http://www.ncbi.nlm.nih.gov/protein/JPN15229">JPN15229</accessionNumber>
</emphasis>
,
<emphasis id="FEDDCCA826BD80296804328D64C535E0" italics="true" pageId="0" pageNumber="137">15230</emphasis>
with flowers (FU!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AFFF442530136BB5D7146D0BC3F52140" pageId="0" pageNumber="137" type="notes">
<paragraph id="164075195D6875F87356829C39899A25" pageId="0" pageNumber="137">Note.</paragraph>
<paragraph id="7B3BF3C080B2B2FB4BADD0A46A49CEBF" pageId="0" pageNumber="137">
The clade comprising this species and
<taxonomicName id="DE24A4C727CB9C331B40FEF9B1BFA533" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. longipedicellata" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="longipedicellata">
<emphasis id="2FAEE179B9464F54E4C5855A77351F17" italics="true" pageId="0" pageNumber="137">H. longipedicellata</emphasis>
</taxonomicName>
is sister to all the other species in Clade 1 (Fig.
<figureCitation id="8B5A1CA3DB5B1E65EE430BF0FC6D6471" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A rooted Maximum Likelihood tree for Hosta Clade 1 reconstructed using MIG-seq data. Bootstrap values are indicated on internodes. Based on the Maximum Likelihood tree of 28 Japanese species (Fig. 1), H. gracillima is used as an outgroup." figureDoi="10.3897/phytokeys.235.99140.figure2" httpUri="https://binary.pensoft.net/fig/938503" pageId="0" pageNumber="137">2</figureCitation>
), and its morphological similarities to
<taxonomicName id="00D31FAC634F7A5E17A3DF99C13FEBF0" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. longipedicellata" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="longipedicellata">
<emphasis id="75566AD14AF9CEFDDB41594A45622B05" italics="true" pageId="0" pageNumber="137">H. longipedicellata</emphasis>
</taxonomicName>
and
<taxonomicName id="34840148F83CCD4B021D7C1D297B3C5C" class="Dicotyledoneae" family="Plantaginaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. minazukiflora" order="Plantaginales" pageId="0" pageNumber="137" phylum="Angiospermae" rank="species" species="minazukiflora">
<emphasis id="3471E9C6AB53477FD5482D8ECEC071A6" italics="true" pageId="0" pageNumber="137">H. minazukiflora</emphasis>
</taxonomicName>
are considered to be derived from a common ancestor.
</paragraph>
</subSubSection>
</treatment>
</document>