402 lines
56 KiB
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402 lines
56 KiB
XML
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<mods:title id="80534EF8D23FAEE10768A13039E9E001">Cutting the Gordian knot: a historical and taxonomic revision of the Jurassic crocodylomorph Metriorhynchus</mods:title>
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<mods:namePart id="AF3E62EEFDC423DE00307C64FC81A5B9">Kitson, James J. N.</mods:namePart>
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<treatment id="5E272C7FFF85FFD71AE2FCCF7137F888" ID-DOI="http://doi.org/10.5281/zenodo.7017013" ID-GBIF-Taxon="199723522" ID-Zenodo-Dep="7017013" LSID="urn:lsid:plazi:treatment:5E272C7FFF85FFD71AE2FCCF7137F888" httpUri="http://treatment.plazi.org/id/5E272C7FFF85FFD71AE2FCCF7137F888" lastPageNumber="543" pageId="33" pageNumber="543">
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<subSubSection id="9E94CEE2FF85FFD71AE2FCCF768FFC6F" pageId="33" pageNumber="543" type="nomenclature">
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<heading id="8D792A05FF85FFD71AE2FCCF768FFC6F" centered="true" fontSize="9" level="2" pageId="33" pageNumber="543" reason="2">
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COMPARISONS BETWEEN
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<taxonomicName id="118EE6EAFF85FFD71BCEFCF07527FC6F" baseAuthorityName="Holl" baseAuthorityYear="1829" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
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<emphasis id="E4FA417BFF85FFD71BCEFCF07527FC6F" italics="true" pageId="33" pageNumber="543">METRIORHYNCHUS BREVIROSTRIS</emphasis>
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</taxonomicName>
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AND OTHER METRIORHYNCHIDS
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</heading>
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</paragraph>
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<subSubSection id="9E94CEE2FF85FFD71AB8FC3675EAF9B9" pageId="33" pageNumber="543" type="description">
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<paragraph id="D6319D69FF85FFD71AB8FC3675EAF9B9" blockId="33.[145,762,934,1907]" pageId="33" pageNumber="543">
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While the
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<typeStatus id="093523CBFF85FFD71B3AFC36753FFC45" box="[275,378,934,956]" pageId="33" pageNumber="543" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="118EE6EAFF85FFD71B8DFC3676BCFC42" box="[420,761,934,955]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
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<emphasis id="E4FA417BFF85FFD71B8DFC3676BCFC42" box="[420,761,934,955]" italics="true" pageId="33" pageNumber="543">Metriorhynchus brevirostris</emphasis>
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</taxonomicName>
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(MHNG
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<date id="A230BBA9FF85FFD71AD2FC557515FC22" box="[251,336,965,987]" pageId="33" pageNumber="543" value="2232-05">V-2232</date>
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) lacks autapomorphic characters, it is readily distinguishable from better-known metriorhynchid species. It has a unique combination of metriorhynchid symplesiomorphies and a slightly posteriorly retracted narial fossa (see:
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<bibRefCitation id="B21FE098FF85FFD7184DFBD0749DFB8D" author="Young" etAl="et al." firstAuthor="Young" pageId="33" pageNumber="543" pagination="494 - 520" refId="ref37488" refString="Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJN. 2020 a. Convergent evolution and possible constraint in the posterodorsal retractions of the external nares in pelagic crocodylomorphs. Zoological Journal of the Linnean Society 189: 494 - 520." type="journal article" year="2020" yearSuffix="a">
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Young
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<emphasis id="E4FA417BFF85FFD7189FFBD076B7FBAC" box="[694,754,1088,1109]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
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, 2020a
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||
</bibRefCitation>
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||
). Furthermore, it lacks the autapomorphies seen in the rostra of the Late Jurassic genera
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<taxonomicName id="118EE6EAFF85FFD71AB8FB0C7559FB48" authorityName=", Hulke" authorityYear="1869" box="[145,284,1180,1201]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
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||
<emphasis id="E4FA417BFF85FFD71AB8FB0C7559FB48" box="[145,284,1180,1201]" italics="true" pageId="33" pageNumber="543">Dakosaurus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="118EE6EAFF85FFD71B00FB0C75F9FB48" authorityName="Owen" authorityYear="1884" box="[297,444,1180,1201]" class="Reptilia" family="Metriorhynchidae" genus="Plesiosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
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<emphasis id="E4FA417BFF85FFD71B00FB0C75F9FB48" box="[297,444,1180,1201]" italics="true" pageId="33" pageNumber="543">Plesiosuchus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="118EE6EAFF85FFD71BE3FB0C761BFB48" box="[458,606,1180,1201]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
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<emphasis id="E4FA417BFF85FFD71BE3FB0C761BFB48" box="[458,606,1180,1201]" italics="true" pageId="33" pageNumber="543">Torvoneustes</emphasis>
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</taxonomicName>
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, the ‘E-clade’ taxa,
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<taxonomicName id="118EE6EAFF85FFD71AE7FB2B7512FB29" authorityName="Wagner" authorityYear="1858" box="[206,343,1211,1232]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
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<emphasis id="E4FA417BFF85FFD71AE7FB2B7512FB29" box="[206,343,1211,1232]" italics="true" pageId="33" pageNumber="543">Cricosaurus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="118EE6EAFF85FFD71BA7FB2A7679FB36" box="[398,572,1210,1231]" class="Reptilia" family="Metriorhynchidae" genus="Rhacheosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
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<emphasis id="E4FA417BFF85FFD71BA7FB2A7679FB36" box="[398,572,1210,1231]" italics="true" pageId="33" pageNumber="543">Rhacheosaurus</emphasis>
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</taxonomicName>
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(see:
|
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<bibRefCitation id="B21FE098FF85FFD7185EFB2B748DFB17" author="Young" etAl="et al." firstAuthor="Young" pageId="33" pageNumber="543" pagination="44985" refId="ref37244" refString="Young MT, Brusatte SL, Andrade MB, Desojo JB, Beatty BL, Steel L, Fernandez MS, Sakamoto M, Ruiz- Omenaca JI, Schoch RR. 2012. The cranial osteology and feeding ecology of the metriorhynchid crocodylomorph genera Dakosaurus and Plesiosuchus from the Late Jurassic of Europe. PLoS One 7: e 44985." type="journal article" year="2012">
|
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Young
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||
<emphasis id="E4FA417BFF85FFD718EAFB2B76B7FB36" box="[707,754,1210,1232]" italics="true" pageId="33" pageNumber="543">et al</emphasis>
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. 2012
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</bibRefCitation>
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,
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<bibRefCitation id="B21FE098FF85FFD71AFBFB497552FB17" author="Young MT & Sachs S & Abel P & Foffa D & Herrera Y & Kitson JJN" box="[210,279,1241,1262]" pageId="33" pageNumber="543" pagination="494 - 520" refId="ref37488" refString="Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJN. 2020 a. Convergent evolution and possible constraint in the posterodorsal retractions of the external nares in pelagic crocodylomorphs. Zoological Journal of the Linnean Society 189: 494 - 520." type="journal article" year="2020">2020a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B21FE098FF85FFD71B08FB497595FB17" author="Abel" box="[289,464,1241,1263]" etAl="et al." firstAuthor="Abel" pageId="33" pageNumber="543" refId="ref31725" refString="Abel P, Sachs S, Young MT. 2020. Metriorhynchid crocodylomorphs from the lower Kimmeridgian of southern Germany: evidence for a new large-bodied geosaurin lineage in Europe. Alcheringa. Doi: 10.1080 / 03115518.2019.1701079." type="book" year="2020">
|
||
Abel
|
||
<emphasis id="E4FA417BFF85FFD71B70FB4A75CAFB17" box="[345,399,1241,1263]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2020
|
||
</bibRefCitation>
|
||
). These include the rostral shortening and resultant reduction in tooth count seen in Geosaurini, the amblygnathous rostrum and lateral plates seen in
|
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<taxonomicName id="118EE6EAFF85FFD71B63FAA57599FAB3" authorityName=", Hulke" authorityYear="1869" box="[330,476,1333,1354]" class="Reptilia" family="Metriorhynchidae" genus="Dakosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
|
||
<emphasis id="E4FA417BFF85FFD71B63FAA57599FAB3" box="[330,476,1333,1354]" italics="true" pageId="33" pageNumber="543">Dakosaurus</emphasis>
|
||
</taxonomicName>
|
||
, the robust triangular rostrum and elongated palatines of
|
||
<taxonomicName id="118EE6EAFF85FFD71807FAC47685FA90" authorityName="Owen" authorityYear="1884" box="[558,704,1364,1385]" class="Reptilia" family="Metriorhynchidae" genus="Plesiosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
|
||
<emphasis id="E4FA417BFF85FFD71807FAC47685FA90" box="[558,704,1364,1385]" italics="true" pageId="33" pageNumber="543">Plesiosuchus</emphasis>
|
||
</taxonomicName>
|
||
, and the mediolaterally expanded maxillary alveoli seen in the ‘E-clade’ (see:
|
||
<bibRefCitation id="B21FE098FF85FFD71BBBFA0176DAFA5F" author="Wilkinson" box="[402,671,1425,1447]" etAl="et al." firstAuthor="Wilkinson" pageId="33" pageNumber="543" pagination="1307 - 1333" refId="ref36933" refString="Wilkinson LE, Young MT, Benton MJ. 2008. A new metriorhynchid crocodilian (Mesoeucrocodylia: Thalattosuchia) from the Kimmeridgian (Upper Jurassic) of Wiltshire, UK. Palaeontology 51: 1307 - 1333." type="journal article" year="2008">
|
||
Wilkinson
|
||
<emphasis id="E4FA417BFF85FFD7183FFA027611FA5F" box="[534,596,1425,1447]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B21FE098FF85FFD71886FA027553FA3C" author="Young" etAl="et al." firstAuthor="Young" pageId="33" pageNumber="543" pagination="44985" refId="ref37244" refString="Young MT, Brusatte SL, Andrade MB, Desojo JB, Beatty BL, Steel L, Fernandez MS, Sakamoto M, Ruiz- Omenaca JI, Schoch RR. 2012. The cranial osteology and feeding ecology of the metriorhynchid crocodylomorph genera Dakosaurus and Plesiosuchus from the Late Jurassic of Europe. PLoS One 7: e 44985." type="journal article" year="2012">
|
||
Young
|
||
<emphasis id="E4FA417BFF85FFD71AB8FA207480FA3C" box="[145,197,1456,1477]" italics="true" pageId="33" pageNumber="543">et al</emphasis>
|
||
., 2012
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B21FE098FF85FFD71B0CFA207529FA3C" author="Young MT & Sachs S & Abel P & Foffa D & Herrera Y & Kitson JJN" box="[293,364,1456,1477]" pageId="33" pageNumber="543" pagination="494 - 520" refId="ref37488" refString="Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJN. 2020 a. Convergent evolution and possible constraint in the posterodorsal retractions of the external nares in pelagic crocodylomorphs. Zoological Journal of the Linnean Society 189: 494 - 520." type="journal article" year="2020">2020a</bibRefCitation>
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;
|
||
<bibRefCitation id="B21FE098FF85FFD71B55FA207605FA3C" author="Abel" box="[380,576,1456,1478]" etAl="et al." firstAuthor="Abel" pageId="33" pageNumber="543" refId="ref31725" refString="Abel P, Sachs S, Young MT. 2020. Metriorhynchid crocodylomorphs from the lower Kimmeridgian of southern Germany: evidence for a new large-bodied geosaurin lineage in Europe. Alcheringa. Doi: 10.1080 / 03115518.2019.1701079." type="book" year="2020">
|
||
Abel
|
||
<emphasis id="E4FA417BFF85FFD71B92FA2075B2FA3C" box="[443,503,1456,1477]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2020
|
||
</bibRefCitation>
|
||
). Furthermore,
|
||
<emphasis id="E4FA417BFF85FFD71AB8FA5F7501FA1A" box="[145,324,1486,1508]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD71AE9FA5E7501FA1A" box="[192,324,1486,1507]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
lacks the ossified internarial bar and more noticeable narial fossa retraction seen in the rhacheosaurin genera
|
||
<taxonomicName id="118EE6EAFF85FFD71BA4F99C7653F9D8" authorityName="Wagner" authorityYear="1858" box="[397,534,1548,1569]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
|
||
<emphasis id="E4FA417BFF85FFD71BA4F99C7653F9D8" box="[397,534,1548,1569]" italics="true" pageId="33" pageNumber="543">Cricosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="118EE6EAFF85FFD71862F99C76BCF9D8" box="[587,761,1548,1569]" class="Reptilia" family="Metriorhynchidae" genus="Rhacheosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="genus">
|
||
<emphasis id="E4FA417BFF85FFD71862F99C76BCF9D8" box="[587,761,1548,1569]" italics="true" pageId="33" pageNumber="543">Rhacheosaurus</emphasis>
|
||
</taxonomicName>
|
||
(see:
|
||
<bibRefCitation id="B21FE098FF85FFD71AE0F9BB75E5F9B9" author="Young" box="[201,416,1578,1600]" etAl="et al." firstAuthor="Young" pageId="33" pageNumber="543" pagination="494 - 520" refId="ref37488" refString="Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJN. 2020 a. Convergent evolution and possible constraint in the posterodorsal retractions of the external nares in pelagic crocodylomorphs. Zoological Journal of the Linnean Society 189: 494 - 520." type="journal article" year="2020" yearSuffix="a">
|
||
Young
|
||
<emphasis id="E4FA417BFF85FFD71B3FF9BB750BF9C6" box="[278,334,1578,1600]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2020a
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9E94CEE2FF85FFD71A80F9D97137F888" pageId="33" pageNumber="543" type="reference_group">
|
||
<paragraph id="D6319D69FF85FFD71A80F9D97036FD1D" blockId="33.[145,762,934,1907]" lastBlockId="33.[809,1426,197,1905]" pageId="33" pageNumber="543">
|
||
<taxonomicName id="118EE6EAFF85FFD71A80F9D975B2F9A7" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[169,503,1609,1630]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="E4FA417BFF85FFD71A80F9D975B2F9A7" box="[169,503,1609,1630]" italics="true" pageId="33" pageNumber="543">Metriorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all known Middle Jurassic metriorhynchids in that the narial fossa anterior margin is slightly retracted, being level to the P2 alveoli rather than the P1 alveoli (see:
|
||
<bibRefCitation id="B21FE098FF85FFD71B56F9547620F920" author="Young" box="[383,613,1732,1753]" etAl="et al." firstAuthor="Young" pageId="33" pageNumber="543" pagination="494 - 520" refId="ref37488" refString="Young MT, Sachs S, Abel P, Foffa D, Herrera Y, Kitson JJN. 2020 a. Convergent evolution and possible constraint in the posterodorsal retractions of the external nares in pelagic crocodylomorphs. Zoological Journal of the Linnean Society 189: 494 - 520." type="journal article" year="2020" yearSuffix="a">
|
||
Young
|
||
<emphasis id="E4FA417BFF85FFD71BFBF954764BF920" box="[466,526,1732,1753]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2020a
|
||
</bibRefCitation>
|
||
). Compared to the Callovian geosaurines, the presence of at least 13 maxillary alveoli anterior to the palatines is higher than the dental formulae of ‘
|
||
<emphasis id="E4FA417BFF85FFD71813F8B076B6F8CC" box="[570,755,1824,1845]" italics="true" pageId="33" pageNumber="543">Metriorhynchus</emphasis>
|
||
’
|
||
<emphasis id="E4FA417BFF85FFD71AB8F8AE750AF8AA" box="[145,335,1854,1875]" italics="true" pageId="33" pageNumber="543">brachyrhynchus</emphasis>
|
||
and
|
||
<taxonomicName id="118EE6EAFF85FFD71BA7F8AF76BCF8AA" baseAuthorityName="Foffa & Young" baseAuthorityYear="2014" box="[398,761,1854,1876]" class="Reptilia" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="E4FA417BFF85FFD71BA7F8AF76BCF8AA" box="[398,761,1854,1876]" italics="true" pageId="33" pageNumber="543">Tyrannoneustes lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="9B0CA8D2FF85FFD71AB0F8CD74ABF88A" box="[153,238,1885,1907]" captionStart="Table 6" captionStartId="29.[145,209,196,217]" captionTargetPageId="29" captionText="Table 6. Dental formulae of metriorhynchid species from the Callovian of Europe" httpUri="http://table.plazi.org/id/82F1CDE1FFB9FFEB1AB8FF5477AFFF23" pageId="33" pageNumber="543" tableUuid="82F1CDE1FFB9FFEB1AB8FF5477AFFF23">Table 6</tableCitation>
|
||
;
|
||
<figureCitation id="4EB581ECFF85FFD71AD2F8CD7579F88B" box="[251,316,1885,1907]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">Fig. 7</figureCitation>
|
||
). Moreover, the nasal terminates level to the M8 alveoli in
|
||
<emphasis id="E4FA417BFF85FFD71E21FF5670FDFF23" box="[1032,1208,197,219]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD71E1FFF5570FDFF23" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[1078,1208,197,218]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
, whereas the nasal terminates level to the M
|
||
<quantity id="1176308CFF85FFD71E76FF7470C8FF03" box="[1119,1165,228,250]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="33" pageNumber="543" unit="in" value="6.0">6 in</quantity>
|
||
‘
|
||
<emphasis id="E4FA417BFF85FFD71EB3FF747081FF00" box="[1178,1220,228,249]" italics="true" pageId="33" pageNumber="543">Me.</emphasis>
|
||
’
|
||
<emphasis id="E4FA417BFF85FFD71EFBFF7471D5FF00" box="[1234,1424,228,249]" italics="true" pageId="33" pageNumber="543">brachyrhynchus</emphasis>
|
||
(NHMUK PV R 3804) and
|
||
<taxonomicName id="118EE6EAFF85FFD71E75FE937166FEEE" baseAuthorityName="Foffa & Young" baseAuthorityYear="2014" box="[1116,1315,258,280]" class="Reptilia" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="E4FA417BFF85FFD71E75FE937166FEEE" box="[1116,1315,258,280]" italics="true" pageId="33" pageNumber="543">Ty. lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
(PETMG R176) (see
|
||
<figureCitation id="4EB581ECFF85FFD71993FEB17046FECF" box="[954,1027,289,311]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">Fig. 7</figureCitation>
|
||
). The fact that the premaxilla posterodorsal processes terminate level to the M3 alveoli in
|
||
<emphasis id="E4FA417BFF85FFD719B6FECF7012FE8A" box="[927,1111,350,372]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD719F9FECE7012FE8A" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[976,1111,350,371]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
also differentiates it from
|
||
<taxonomicName id="118EE6EAFF85FFD71900FEEE77BCFE6B" baseAuthorityName="Foffa & Young" baseAuthorityYear="2014" box="[809,1017,381,403]" class="Reptilia" family="Metriorhynchidae" genus="Tyrannoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="lythrodectikos">
|
||
<emphasis id="E4FA417BFF85FFD71900FEEE77BCFE6B" box="[809,1017,381,403]" italics="true" pageId="33" pageNumber="543">Ty. lythrodectikos</emphasis>
|
||
</taxonomicName>
|
||
, where these processes terminate level to the M5 alveoli (PETMG R176, see
|
||
<figureCitation id="4EB581ECFF85FFD71F22FE0C710EFE48" box="[1291,1355,412,434]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">Fig. 7</figureCitation>
|
||
;
|
||
<bibRefCitation id="B21FE098FF85FFD71F7EFE0C7794FE36" author="Foffa & Young" firstAuthor="Foffa" pageId="33" pageNumber="543" pagination="608" refId="ref33398" refString="Foffa D, Young MT. 2014. The cranial osteology of Tyrannoneustes lythrodectikos (Crocodylomorpha: Metriorhynchidae) from the Middle Jurassic of Europe. PeerJ 2: e 608." type="journal article" year="2014">Foffa & Young, 2014</bibRefCitation>
|
||
).
|
||
<taxonomicName id="118EE6EAFF85FFD719CEFE2A716EFE36" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[999,1323,442,463]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="E4FA417BFF85FFD719CEFE2A716EFE36" box="[999,1323,442,463]" italics="true" pageId="33" pageNumber="543">Metriorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
can also be distinguished from the Callovian metriorhynchine
|
||
<taxonomicName id="118EE6EAFF85FFD71900FE6870C4FDF4" authority="(Parrilla-Bel et al., 2013)" baseAuthorityName="Parrilla-Bel" baseAuthorityYear="2013" box="[809,1153,504,525]" class="Reptilia" family="Metriorhynchidae" genus="Maledictosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="riclaensis">
|
||
<emphasis id="E4FA417BFF85FFD71900FE6870C4FDF4" box="[809,1153,504,525]" italics="true" pageId="33" pageNumber="543">Maledictosuchus riclaensis</emphasis>
|
||
</taxonomicName>
|
||
as it lacks the welldeveloped reception pits on the lateral margins of the anterior maxilla seen in
|
||
<taxonomicName id="118EE6EAFF85FFD71E62FDA577EEFD90" authority="(Parrilla-Bel et al., 2013)" baseAuthorityName="Parrilla-Bel" baseAuthorityYear="2013" class="Reptilia" family="Metriorhynchidae" genus="Maledictosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="riclaensis">
|
||
<emphasis id="E4FA417BFF85FFD71E62FDA570B3FDB3" box="[1099,1270,565,586]" italics="true" pageId="33" pageNumber="543">Mal. riclaensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B21FE098FF85FFD71F2EFDA577E5FD90" author="Parrilla-Bel" etAl="et al." firstAuthor="Parrilla-Bel" pageId="33" pageNumber="543" pagination="54275" refId="ref35759" refString="Parrilla-Bel J, Young MT, Moreno-Azanza M, Canudo JI. 2013. The first metriorhynchid crocodyliform from the Middle Jurassic of Spain, with implications for evolution of the subclade Rhacheosaurini. PLoS One 8: e 54275." type="journal article" year="2013">
|
||
Parrilla-Bel
|
||
<emphasis id="E4FA417BFF85FFD71900FDC4771BFD90" box="[809,862,596,617]" italics="true" pageId="33" pageNumber="543">et al.</emphasis>
|
||
, 2013
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="118EE6EAFF85FFD7199BFDC470A9FD90" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[946,1260,596,617]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="E4FA417BFF85FFD7199BFDC470A9FD90" box="[946,1260,596,617]" italics="true" pageId="33" pageNumber="543">Metriorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
is also distinct from
|
||
<taxonomicName id="118EE6EAFF85FFD7194DFDE27008FD7E" baseAuthorityName="Andrews" baseAuthorityYear="1913" box="[868,1101,626,647]" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="E4FA417BFF85FFD7194DFDE27008FD7E" box="[868,1101,626,647]" italics="true" pageId="33" pageNumber="543">Gracilineustes leedsi</emphasis>
|
||
</taxonomicName>
|
||
as it lacks: the exceptionally high tooth count, closely packed alveoli and the poorly developed-to-almost absent maxillary ornamentation of
|
||
<taxonomicName id="118EE6EAFF85FFD7196DFD5F702BFD1D" authority="(Andrews, 1913)" baseAuthorityName="Andrews" baseAuthorityYear="1913" box="[836,1134,718,740]" class="Reptilia" family="Metriorhynchidae" genus="Gracilineustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="E4FA417BFF85FFD7196DFD5F77E9FD1A" box="[836,940,718,740]" italics="true" pageId="33" pageNumber="543">Gr. leedsi</emphasis>
|
||
(
|
||
<bibRefCitation id="B21FE098FF85FFD71992FD5E7021FD1A" author="Andrews" box="[955,1124,718,740]" firstAuthor="Andrews" pageId="33" pageNumber="543" refId="ref31961" refString="Andrews CW. 1913. A descriptive catalogue of the marine reptiles of the Oxford Clay, Part Two. London: British Museum (Natural History)." type="book" year="1913">Andrews, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D6319D69FF85FFD71968FD7D7137F888" blockId="33.[809,1426,197,1905]" pageId="33" pageNumber="543">
|
||
There are notable differences between
|
||
<taxonomicName id="118EE6EAFF85FFD71900FC9C7039FCD8" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[809,1148,780,801]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="E4FA417BFF85FFD71900FC9C7039FCD8" box="[809,1148,780,801]" italics="true" pageId="33" pageNumber="543">Metriorhynchus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="E4FA417BFF85FFD71E95FC9C71CFFCD8" box="[1212,1418,780,801]" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71EC4FC9C71CFFCD8" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[1261,1418,780,801]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
. Although the rostrum of
|
||
<emphasis id="E4FA417BFF85FFD71EAAFCBB7113FCB9" box="[1155,1366,810,832]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD71E94FCBA7113FCB9" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[1213,1366,810,832]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
has encrustations, the maxillary ornamentation is noticeably not as pronounced as it is in
|
||
<emphasis id="E4FA417BFF85FFD71EFCFCF771D4FC85" box="[1237,1425,871,892]" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71ED7FCF771D4FC85" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[1278,1425,871,892]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
(compare
|
||
<figureCitation id="4EB581ECFF85FFD719B1FC1677A8FC65" box="[920,1005,902,924]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Figs 1A</figureCitation>
|
||
–
|
||
<figureCitation id="4EB581ECFF85FFD719D4FC16704FFC65" box="[1021,1034,902,924]" captionStart="Figure 6" captionStartId="31.[145,225,1298,1320]" captionTargetBox="[145,1425,195,1259]" captionTargetId="figure-254@31.[145,1425,195,1259]" captionTargetPageId="31" captionText="Figure 6. Thalattosuchus superciliosus lectotype MNHN.F RJN 256. A, dorsal view; B, palatal view; C, left lateral view. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.7017034" httpUri="https://zenodo.org/record/7017034/files/figure.png" pageId="33" pageNumber="543">6</figureCitation>
|
||
and
|
||
<figureCitation id="4EB581ECFF85FFD71E6AFC167027FC62" box="[1091,1122,902,923]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">7C</figureCitation>
|
||
;
|
||
<bibRefCitation id="B21FE098FF85FFD71E47FC167153FC62" author="Andrews" box="[1134,1302,902,924]" firstAuthor="Andrews" pageId="33" pageNumber="543" refId="ref31961" refString="Andrews CW. 1913. A descriptive catalogue of the marine reptiles of the Oxford Clay, Part Two. London: British Museum (Natural History)." type="book" year="1913">Andrews, 1913</bibRefCitation>
|
||
). In dorsal view, there is a constriction at the premaxilla–maxilla contact in
|
||
<emphasis id="E4FA417BFF85FFD71981FC547024FC21" box="[936,1121,963,985]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD719F0FC537024FC21" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[985,1121,963,984]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
, followed by a noticeable broadening of the premaxilla at approximately the level of the narial fossa posterior margin (
|
||
<figureCitation id="4EB581ECFF85FFD71F2FFB917113FBEF" box="[1286,1366,1025,1047]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Fig. 1A</figureCitation>
|
||
);
|
||
<emphasis id="E4FA417BFF85FFD71F42FB9177F9FBCD" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71900FB8F77F9FBCD" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[809,956,1055,1076]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
lacks both of features, with the posterior premaxilla and anterior maxilla having a largely constant width (PETMG R17 and MNHN.F RJN 256; see
|
||
<figureCitation id="4EB581ECFF85FFD7197EFBEB77DAFB68" box="[855,927,1147,1169]" captionStart="Figure 6" captionStartId="31.[145,225,1298,1320]" captionTargetBox="[145,1425,195,1259]" captionTargetId="figure-254@31.[145,1425,195,1259]" captionTargetPageId="31" captionText="Figure 6. Thalattosuchus superciliosus lectotype MNHN.F RJN 256. A, dorsal view; B, palatal view; C, left lateral view. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.7017034" httpUri="https://zenodo.org/record/7017034/files/figure.png" pageId="33" pageNumber="543">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="4EB581ECFF85FFD71987FBEC778BFB68" box="[942,974,1148,1169]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">7C</figureCitation>
|
||
). In
|
||
<emphasis id="E4FA417BFF85FFD71E21FBEC7085FB69" box="[1032,1216,1147,1169]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD71E10FBEB7085FB69" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[1081,1216,1147,1168]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
the premaxillary alveoli successively increase in size, with the P1 alveoli being circular in shape and the P3 alveoli being the most oval of the premaxillary tooth-row (
|
||
<figureCitation id="4EB581ECFF85FFD71EEEFB47715FFB14" box="[1223,1306,1239,1261]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Fig. 1B</figureCitation>
|
||
); whereas in
|
||
<emphasis id="E4FA417BFF85FFD71965FB667064FAF2" box="[844,1057,1270,1291]" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71956FB667064FAF2" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[895,1057,1270,1291]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
the premaxillary alveoli are largely comparable in size, with the P1 and P3 alveoli being oval in shape, and the P2 being the most circular (PETMG R17 and MNHN.F RJN 256; see
|
||
<figureCitation id="4EB581ECFF85FFD71F26FAC27110FA91" box="[1295,1365,1362,1384]" captionStart="Figure 6" captionStartId="31.[145,225,1298,1320]" captionTargetBox="[145,1425,195,1259]" captionTargetId="figure-254@31.[145,1425,195,1259]" captionTargetPageId="31" captionText="Figure 6. Thalattosuchus superciliosus lectotype MNHN.F RJN 256. A, dorsal view; B, palatal view; C, left lateral view. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.7017034" httpUri="https://zenodo.org/record/7017034/files/figure.png" pageId="33" pageNumber="543">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="4EB581ECFF85FFD71F4AFAC271C6FA9E" box="[1379,1411,1362,1383]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">7C</figureCitation>
|
||
). The anterior maxillary alveoli of
|
||
<emphasis id="E4FA417BFF85FFD71E8DFAE17120FA7F" box="[1188,1381,1393,1414]" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71EF8FAE17120FA7F" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[1233,1381,1393,1414]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
are fairly tightly packed, with the interalveolar spaces reduced to thin laminae (especially for the M2–M3 alveoli; PETMG R17 and MNHN.F RJN 256); whereas in
|
||
<emphasis id="E4FA417BFF85FFD71961FA7C77B9F9F9" box="[840,1020,1515,1537]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD7195EFA7B77B9F9F9" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[887,1020,1515,1536]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
the interalveolar spaces are much broader, even between the M1–M2 and M2–M3 alveoli (
|
||
<figureCitation id="4EB581ECFF85FFD7191BF9B97730F9C7" box="[818,885,1577,1599]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Fig. 1</figureCitation>
|
||
). In
|
||
<emphasis id="E4FA417BFF85FFD71999F9B9702CF9C7" box="[944,1129,1577,1598]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD719C8F9B9702CF9C7" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[993,1129,1577,1598]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
, the premaxilla–maxilla suture is almost straight in dorsal view (
|
||
<figureCitation id="4EB581ECFF85FFD71F00F9D771C4F9A4" box="[1321,1409,1607,1629]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Fig. 1A</figureCitation>
|
||
), whereas in
|
||
<emphasis id="E4FA417BFF85FFD7199EF9F670C6F982" box="[951,1155,1638,1659]" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD719CEF9F670C6F982" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[999,1155,1638,1659]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
it has a more ‘jagged’ interdigitating profile (PETMG R17 and MNHN.F RJN 256; see
|
||
<figureCitation id="4EB581ECFF85FFD719E0F9337055F940" box="[969,1040,1699,1721]" captionStart="Figure 6" captionStartId="31.[145,225,1298,1320]" captionTargetBox="[145,1425,195,1259]" captionTargetId="figure-254@31.[145,1425,195,1259]" captionTargetPageId="31" captionText="Figure 6. Thalattosuchus superciliosus lectotype MNHN.F RJN 256. A, dorsal view; B, palatal view; C, left lateral view. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.7017034" httpUri="https://zenodo.org/record/7017034/files/figure.png" pageId="33" pageNumber="543">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="4EB581ECFF85FFD71E34F9347078F940" box="[1053,1085,1700,1721]" captionStart="Figure 7" captionStartId="34.[162,240,986,1008]" captionTargetBox="[172,1443,198,945]" captionTargetId="figure-380@34.[163,1443,195,947]" captionTargetPageId="34" captionText="Figure 7. Comparative plate of metriorhynchid anterior cranial rostra. A, B, Metriorhynchus brevirostris, holotype MHNG V-2232 in dorsal and palatal views, respectively; C, D, Thalattosuchus superciliosus, referred specimen PETMG R17 in dorsal and palatal views, respectively; E, F, ‘Metriorhynchus’ brachyrhynchus, referred specimen NHMUK PV R 3804 (holotype of ‘Metriorhynchus’ cultridens) in dorsal and palatal views, respectively; G, H, ‘Metriorhynchus’ palpebrosus, holotype OUMNH J.29823 in dorsal and palatal views, respectively; I, J, Tyrannoneustes lythrodectikos, referred specimen PETMG R176 in dorsal and palatal views, respectively; K, L, Plesiosuchus manselii, holotype NHMUK PV OR 40103 in dorsal and palatal views, respectively. See text for anatomical abbreviations. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.7017037" httpUri="https://zenodo.org/record/7017037/files/figure.png" pageId="33" pageNumber="543">7C</figureCitation>
|
||
). The nasal proceeds slightly further anteriorly in
|
||
<emphasis id="E4FA417BFF85FFD71E1FF95370BDF92E" box="[1078,1272,1730,1752]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD71E43F95270BDF92E" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[1130,1272,1730,1751]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
than in
|
||
<emphasis id="E4FA417BFF85FFD71F40F95277F8F90F" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71900F97177F8F90F" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[809,957,1761,1782]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
, with seven maxillary alveoli anterior to the nasals in
|
||
<emphasis id="E4FA417BFF85FFD719EEF8907039F8ED" box="[967,1148,1791,1813]" italics="true" pageId="33" pageNumber="543">
|
||
Me.
|
||
<taxonomicName id="118EE6EAFF85FFD719DEF96F7039F8ED" baseAuthorityName="Holl" baseAuthorityYear="1829" box="[1015,1148,1791,1812]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="brevirostris">brevirostris</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="4EB581ECFF85FFD71EA5F96F7088F8ED" box="[1164,1229,1791,1813]" captionStart="Figure 1" captionStartId="23.[145,225,1678,1700]" captionTargetBox="[227,1345,197,1637]" captionTargetId="figure-74@23.[225,1345,195,1638]" captionTargetPageId="23" captionText="Figure 1. Metriorhynchus brevirostris holotype MHNG V-2232. A, dorsal view; B, palatal view. See text for anatomical abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7017017" httpUri="https://zenodo.org/record/7017017/files/figure.png" pageId="33" pageNumber="543">Fig. 1</figureCitation>
|
||
); whereas in
|
||
<emphasis id="E4FA417BFF85FFD71F42F96F7784F8CA" italics="true" pageId="33" pageNumber="543">
|
||
Th.
|
||
<taxonomicName id="118EE6EAFF85FFD71900F88E7784F8CA" authorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" authorityYear="2021" baseAuthorityName="Young & Brignon & Sachs & Hornung & Foffa & Kitson & Johnson & Steel" baseAuthorityYear="1977" box="[809,961,1822,1843]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="33" pageNumber="543" phylum="Chordata" rank="species" species="superciliosus">superciliosus</taxonomicName>
|
||
</emphasis>
|
||
there can be eight (MNHN.F RJN 256, NHMUK PV R 2051), nine (GLAHM V930, NHMUK PV R 2030, PETMG R17) or 10–11 (GLAHM V962).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |