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<document id="3706760A65D4CF3EB211AA1D676D209C" ID-DOI="10.5281/zenodo.193454" ID-GBIF-Dataset="9fd64c73-48dd-4434-bed0-16047e5da7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="193454" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460144990490" checkinUser="plazi" docAuthor="Ribera, Ignacio, Castro, Agustín &amp; Hernando, Carles" docDate="2010" docId="5E11432F3C24FF92FF7CFF2DFD5697B6" docLanguage="en" docName="zt02351p013.pdf" docOrigin="Zootaxa 2351" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ochthebius (Enicocerus) aguilerai Ribera, Castro &amp; Hernando, 2010, sp.n." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="A2283B573C20FF9BFFEBFFBAFFF8962C" masterDocTitle="Ochthebius (Enicocerus) aguilerai sp. n. from central Spain, with a molecular phylogeny of the Western Palaearctic species of Enicocerus (Coleoptera, Hydraenidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="5" updateTime="1698235395010" updateUser="plazi">
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<mods:title id="A07199797452B58CFF4E4F5D08053F9B">Ochthebius (Enicocerus) aguilerai sp. n. from central Spain, with a molecular phylogeny of the Western Palaearctic species of Enicocerus (Coleoptera, Hydraenidae)</mods:title>
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<mods:namePart id="32706298B9E6DD699CE45613CB5628C4">Ribera, Ignacio</mods:namePart>
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<mods:namePart id="F203849D197FD8ACA8D1658EB7D79003">Castro, Agustín</mods:namePart>
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<mods:namePart id="D62B75EBF875E50697F90560432140F0">Hernando, Carles</mods:namePart>
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<mods:date id="858C4C898EA14573C0C79DE5428E804D">2010</mods:date>
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<treatment id="5E11432F3C24FF92FF7CFF2DFD5697B6" ID-DOI="http://doi.org/10.5281/zenodo.5621341" ID-GBIF-Taxon="124955882" ID-Zenodo-Dep="5621341" LSID="urn:lsid:plazi:treatment:5E11432F3C24FF92FF7CFF2DFD5697B6" httpUri="http://treatment.plazi.org/id/5E11432F3C24FF92FF7CFF2DFD5697B6" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5">
<subSubSection id="9EA2A1B23C24FF9FFF7CFF2DFE9F96F8" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="D607F2393C24FF9FFF7CFF2DFD83969D" blockId="4.[151,635,151,212]" box="[151,635,151,177]" pageId="4" pageNumber="5">
<heading id="8D4F45553C24FF9FFF7CFF2DFD83969D" bold="true" box="[151,635,151,177]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="E4CC2E2B3C24FF9FFF7CFF2DFD83969D" bold="true" box="[151,635,151,177]" pageId="4" pageNumber="5">
<taxonomicName id="11B889BA3C24FF9FFF7CFF2DFDC1969D" ID-CoL="6SFRK" box="[151,569,151,177]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov." subGenus="Enicocerus">
<emphasis id="E4CC2E2B3C24FF9FFF7CFF2DFEDA969D" bold="true" box="[151,290,151,177]" italics="true" pageId="4" pageNumber="5">Ochthebius</emphasis>
(
<emphasis id="E4CC2E2B3C24FF9FFED9FF2DFE43969D" bold="true" box="[306,443,151,177]" italics="true" pageId="4" pageNumber="5">Enicocerus</emphasis>
)
<emphasis id="E4CC2E2B3C24FF9FFE27FF2DFDC1969D" bold="true" box="[460,569,151,177]" italics="true" pageId="4" pageNumber="5">aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C24FF9FFDABFF2DFD83969D" box="[576,635,151,177]" pageId="4" pageNumber="5" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D607F2393C24FF9FFF7CFF00FE9F96F8" blockId="4.[151,635,151,212]" box="[151,359,186,212]" pageId="4" pageNumber="5">
(
<figureCitation id="4E83EEBC3C24FF9FFF74FF00FEF196F8" box="[159,265,186,212]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 10" captionStartId-0="4.[151,269,1955,1979]" captionStartId-1="5.[151,275,1897,1921]" captionTargetBox-0="[157,1428,1088,1924]" captionTargetBox-1="[176,1428,186,1866]" captionTargetId-0="figure@4.[151,1436,1079,1930]" captionTargetId-1="figure@5.[164,1449,174,1875]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 2. Habitus of Ochthebius aguilerai sp. n., 1) male (paratype, NMW); 2) female (paratype, NMW)." captionText-1="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri-0="https://zenodo.org/record/193455/files/figure.png" httpUri-1="https://zenodo.org/record/193456/files/figure.png" pageId="4" pageNumber="5">
Figs 1
<emphasis id="E4CC2E2B3C24FF9FFF02FF00FF0096F8" bold="true" box="[233,248,186,212]" pageId="4" pageNumber="5"></emphasis>
4
</figureCitation>
,
<figureCitation id="4E83EEBC3C24FF9FFEFEFF00FEA696F8" box="[277,350,186,212]" captionStart="FIGURES 11 16" captionStartId="6.[151,270,765,789]" captionTargetBox="[404,1181,198,736]" captionTargetId="figure@6.[395,1191,194,741]" captionTargetPageId="6" captionText="FIGURES 11 16. Last abdominal tergites of O. aguilerai sp. n. 11) ventral, 12) lateral and 13) caudal views (paratype from type locality) and O. legionensis 14) ventral, 15) lateral and 16) caudal views (León, Rioscuro, 30.6. 1985 L. F. Valladares leg.). Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193457/files/figure.png" pageId="4" pageNumber="5">
11
<emphasis id="E4CC2E2B3C24FF9FFEDAFF00FEB896F8" bold="true" box="[305,320,186,212]" pageId="4" pageNumber="5"></emphasis>
13
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)
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C24FF93FF7CFEBDFE0396F4" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="materials_examined">
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<emphasis id="E4CC2E2B3C24FF9FFF7CFEBDFEC7970D" bold="true" box="[151,319,263,289]" pageId="4" pageNumber="5">
<typeStatus id="09034C9B3C24FF9FFF7CFEBDFF2D970D" box="[151,213,263,289]" pageId="4" pageNumber="5">Type</typeStatus>
locality.
</emphasis>
Madrid, Puerto de los Cotos, Arroyo de la Laguna Grande de Peñalara,
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,
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(
<figureCitation id="4E83EEBC3C24FF9FFEC4FE94FE4F9764" box="[303,439,302,328]" captionStart="FIGURES 18 19" captionStartId="7.[151,270,1974,1998]" captionTargetBox="[256,1331,180,1942]" captionTargetId="figure@7.[250,1337,172,1950]" captionTargetPageId="7" captionText="FIGURES 18 19. Type locality of O. aguilerai sp. n. (Arroyo de la Laguna Grande de Peñalara), 18) top, 2 June 2007, with D. T. Bilton and F. Bameul; 19) bottom, 20 July 2006, detail of the microhabitat (fotos, IR)." httpUri="https://zenodo.org/record/193459/files/figure.png" pageId="4" pageNumber="5">
Figs 18
<emphasis id="E4CC2E2B3C24FF9FFE63FE94FE6F9764" bold="true" box="[392,407,302,328]" pageId="4" pageNumber="5"></emphasis>
19
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).
</paragraph>
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<typeStatus id="09034C9B3C24FF9FFF2DFEEFFEF99743" box="[198,257,341,367]" pageId="4" pageNumber="5">Type</typeStatus>
material.
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<emphasis id="E4CC2E2B3C24FF9FFE63FEEFFE0C9742" box="[392,500,341,366]" italics="true" pageId="4" pageNumber="5">Holotype</emphasis>
</typeStatus>
male (
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): “ES Madrid, Puerto de los Cotos \ Arryo. Laguna Grande de Peñalara \
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,
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\ rd M604 pk 38
<date id="A206D4F93C24FF9FFC6DFEC6FC0797BA" box="[902,1023,380,406]" pageId="4" pageNumber="5" value="2006-07-20">20.7.2006</date>
I. Ribera leg.” and
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label (aedegus extracted and mounted on the same card).
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<emphasis id="E4CC2E2B3C24FF9FFD13FE18FC979797" box="[760,879,418,443]" italics="true" pageId="4" pageNumber="5">Paratypes</emphasis>
</typeStatus>
(31 exx):
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and
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(
<collectionCode id="B0A96AFC3C24FF9FFB0CFE18FAC09790" box="[1255,1336,418,444]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="5">NMW</collectionCode>
),
<specimenCount id="C0BE39B03C24FF9FFAA6FE18FA649790" box="[1357,1436,418,444]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFF74FE73FF2697CF" box="[159,222,457,483]" pageId="4" pageNumber="5">CVL</collectionCode>
) and 4 exx (
<collectionCode id="B0A96AFC3C24FF9FFE9DFE73FE5197CF" box="[374,425,457,483]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
) labelled as
<typeStatus id="09034C9B3C24FF9FFDA8FE73FD4897CF" box="[579,688,457,483]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
;
<specimenCount id="C0BE39B03C24FF9FFD56FE73FCE297CF" box="[701,794,457,483]" pageId="4" pageNumber="5" type="male">2 males</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFCC0FE73FCA697CF" box="[811,862,457,483]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
) and
<specimenCount id="C0BE39B03C24FF9FFC49FE73FBF497CF" box="[930,1036,457,483]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFBF7FE73FB8397CF" LSID="urn:lsid:biocol.org:col:35275" box="[1052,1147,457,483]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="4" pageNumber="5">MNCN</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFB6CFE73FB3F97CF" LSID="urn:lsid:biocol.org:col:15536" box="[1159,1223,457,483]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="5">DNA</collectionCode>
voucher MNCN- AI957) “2 ES Madrid Puerto de Cotos
<date id="A206D4F93C24FF9FFD65FE4AFD069426" box="[654,766,496,522]" pageId="4" pageNumber="5" value="2006-07-01">1.7.2006</date>
\ Arroyo de la Laguna Grande de Peñalara \
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\ rd M604 pk 38 IR [I. Ribera] &amp; AC [A. Cieslak] &amp; BR [Bernard Ribera] leg”;
<specimenCount id="C0BE39B03C24FF9FFA66FDACFF24947B" pageId="4" pageNumber="5" type="male">3 males</specimenCount>
<specimenCount id="C0BE39B03C24FF9FFF0FFD87FEAF947B" box="[228,343,573,599]" pageId="4" pageNumber="5" type="female">4 females</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFE8CFD87FE41947B" box="[359,441,573,599]" pageId="4" pageNumber="5">CDTB</collectionCode>
) “
<date id="A206D4F93C24FF9FFE3FFD87FDB6947B" box="[468,590,573,599]" pageId="4" pageNumber="5" value="2008-07">July 2008</date>
,
<collectingCountry id="AEAFB2A93C24FF9FFDB0FD87FD66947B" box="[603,670,573,599]" name="Spain" pageId="4" pageNumber="5">Spain</collectingCountry>
Madrid, Sierra de Guadarrama, Arroyo de la Laguna Grande de Peñalara below Puerto del Coto, D.T. Bilton leg.”;
<specimenCount id="C0BE39B03C24FF9FFD13FDDEFCB09452" box="[760,840,612,638]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFCB2FDDEFC4D9452" LSID="urn:lsid:biocol.org:col:34992" box="[857,949,612,638]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="5">RMNH</collectionCode>
) “ES: Rascafria, Rio Lozoya \ [WGS84] 30T VL257272 [= 30T 425.7 4527.2] M19W \
<date id="A206D4F93C24FF9FFD30FD30FCB69488" box="[731,846,650,676]" pageId="4" pageNumber="5" value="2007-06-05">5.vi.2007</date>
\ leg.
<collectingCountry id="AEAFB2A93C24FF9FFC75FD30FC4B9488" box="[926,947,650,676]" name="Iceland" pageId="4" pageNumber="5">O</collectingCountry>
.Vorst \ Bergriviertje”;
<specimenCount id="C0BE39B03C24FF9FFB24FD30FADD9488" box="[1231,1317,650,676]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
,
<specimenCount id="C0BE39B03C24FF9FFAD9FD30FA659488" box="[1330,1437,650,676]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFF74FD0BFF1994E7" LSID="urn:lsid:biocol.org:col:13223" box="[159,225,689,715]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13223" name="Herbert Art Gallery and Museum" pageId="4" pageNumber="5">COV</collectionCode>
) “ES: Rascafria, Rio Lozoya \ 30T VL253262 [=30T 425.3 4526.2] M6n \
<date id="A206D4F93C24FF9FFB9AFD0BFB1A94E7" box="[1137,1250,689,715]" pageId="4" pageNumber="5" value="2007-06-01">1.vi.2007</date>
\ leg.
<collectingCountry id="AEAFB2A93C24FF9FFADBFD0BFABC94E7" box="[1328,1348,689,715]" name="Iceland" pageId="4" pageNumber="5">O</collectingCountry>
.Vorst \ Bergbeek”;
<specimenCount id="C0BE39B03C24FF9FFEC8FD62FE8B94DE" box="[291,371,728,754]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFE68FD62FE1894DE" LSID="urn:lsid:biocol.org:col:35275" box="[387,480,728,754]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="4" pageNumber="5">MNCN</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFE07FD62FDD494DE" LSID="urn:lsid:biocol.org:col:15536" box="[492,556,728,754]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="5">DNA</collectionCode>
voucher MNCN-AI387) “10
<collectingCountry id="AEAFB2A93C24FF9FFC60FD62FC2594DE" box="[907,989,728,754]" name="Spain" pageId="4" pageNumber="5">SPAIN</collectingCountry>
Avila, Arenas de S. Pedro \ r. Pelayo, rd. AV924 \
<quantity id="11405FDC3C24FF9FFEC1FD44FE959534" box="[298,365,766,792]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="500.0">500m</quantity>
<geoCoordinate id="B38C94FE3C24FF9FFE9EFD44FDFB9534" box="[373,515,766,792]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="40.201668">N40º1206”</geoCoordinate>
<geoCoordinate id="B38C94FE3C24FF9FFDE0FD44FD689534" box="[523,656,766,792]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="15" value="-5.1116667">W5º0642”</geoCoordinate>
\ I.Ribera leg.
<date id="A206D4F93C24FF9FFCAFFD44FC389534" box="[836,960,766,792]" pageId="4" pageNumber="5" value="2005-05-15">15.5.2005</date>
”;
<specimenCount id="C0BE39B03C24FF9FFC3CFD44FBDF9534" box="[983,1063,766,792]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFBD3FD44FB929534" box="[1080,1130,766,792]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
) “2 ES Ávila, Sa. Gredos
<date id="A206D4F93C24FF9FFF7CFC9FFF179513" box="[151,239,805,831]" pageId="4" pageNumber="5" value="06-05-28">280506</date>
\ rv. Pelayo in Arenas de S. Pedro \ rd. AV924, ca. 2Km W Arenas \
<quantity id="11405FDC3C24FF9FFBF8FC9FFBAE9513" box="[1043,1110,805,831]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="500.0">500m</quantity>
<geoCoordinate id="B38C94FE3C24FF9FFBB6FC9FFB0E9513" box="[1117,1270,805,831]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="1" value="40.201557">N40º125,6</geoCoordinate>
<geoCoordinate id="B38C94FE3C24FF9FFB16FC9FFA639513" box="[1277,1435,805,831]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="1" value="-5.111528">W5º0641,5</geoCoordinate>
\ I. Ribera &amp; A. Cieslak leg.”;
<specimenCount id="C0BE39B03C24FF9FFE16FCF6FDAF954A" box="[509,599,844,870]" pageId="4" pageNumber="5" type="male">3 males</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFD8CFCF6FD50954A" box="[615,680,844,870]" pageId="4" pageNumber="5">CAC</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFD58FCF6FD1D954A" box="[691,741,844,870]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFD1AFCF6FCB1954A" LSID="urn:lsid:biocol.org:col:35275" box="[753,841,844,870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="4" pageNumber="5">MNCN</collectionCode>
<collectionCode id="B0A96AFC3C24FF9FFCBBFCF6FC77954A" LSID="urn:lsid:biocol.org:col:15536" box="[848,911,844,870]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="5">DNA</collectionCode>
voucher MNCN-AI648) and
<specimenCount id="C0BE39B03C24FF9FFB07FCF6FAAA954A" box="[1260,1362,844,870]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFA8AFCF6FA6B954A" box="[1377,1427,844,870]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
) “CÁCERES. Jarandilla \ Gta.
<collectingCountry id="AEAFB2A93C24FF9FFE14FCC8FDDA95A0" box="[511,546,882,908]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="5">De</collectingCountry>
Jaranda \ 8.2002 A. Castro leg.”;
<specimenCount id="C0BE39B03C24FF9FFC51FCC8FBD595A0" box="[954,1069,882,908]" pageId="4" pageNumber="5" type="female">5 females</specimenCount>
(
<collectionCode id="B0A96AFC3C24FF9FFBD5FCC8FB8495A0" LSID="urn:lsid:biocol.org:col:15840" box="[1086,1148,882,908]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15840" name="Charles Darwin Research Station" pageId="4" pageNumber="5">CDS</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFB62FCC8FB4595A0" box="[1161,1213,882,908]" pageId="4" pageNumber="5">CJV</collectionCode>
,
<collectionCode id="B0A96AFC3C24FF9FFB22FCC8FAFE95A0" box="[1225,1286,882,908]" pageId="4" pageNumber="5">CVL</collectionCode>
) “
<collectingCountry id="AEAFB2A93C24FF9FFACBFCC8FA6095A0" box="[1312,1432,882,908]" name="Spain" pageId="4" pageNumber="5">ESPAÑA</collectingCountry>
: Cáceres / Guijo de Sta. Bárbara /
<date id="A206D4F93C24FF9FFDD4FC23FD33959F" box="[575,715,921,947]" pageId="4" pageNumber="5" value="1991-06-26">26.06.1991</date>
/ leg. J.A. Régil &amp; J.A. Díaz” [specimens recorded as
<taxonomicName id="11B889BA3C24FF9FFA95FC23FEE395F5" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C24FF9FFA95FC23FEE395F5" italics="true" pageId="4" pageNumber="5">
<collectingCountry id="AEAFB2A93C24FF9FFA95FC23FA60959E" box="[1406,1432,921,946]" name="Iceland" pageId="4" pageNumber="5">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
in
<bibRefCitation id="B2298FC83C24FF9FFEABFC7AFD4995F6" box="[320,689,960,986]" pageId="4" pageNumber="12" refString="Garrido-Gonzalez, J., Diaz-Pazos, J. A. &amp; Regil-Cueto, J. A. (1994) Coleopteros acuaticos de Extremadura 2 (Hydraenidae, Hydrochidae, Helophoridae, Hydrophilidae, Dryopidae y Elmidae). Boletin de la Asociacion Espanola de Entomologia, 18, 113 - 133." type="journal article">
Garrido-González
<emphasis id="E4CC2E2B3C24FF9FFDF0FC7AFDB695F5" box="[539,590,960,985]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. (1994)
</bibRefCitation>
].
</paragraph>
<paragraph id="D607F2393C24FF9FFF2DFC5CFAC6922C" blockId="4.[151,1437,263,1024]" box="[198,1342,998,1024]" pageId="4" pageNumber="5">
Aliquots of the
<collectionCode id="B0A96AFC3C24FF9FFE96FC5CFE45922C" LSID="urn:lsid:biocol.org:col:15536" box="[381,445,998,1024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="5">DNA</collectionCode>
extractions are kept in the
<collectionCode id="B0A96AFC3C24FF9FFD11FC5CFCAA922C" LSID="urn:lsid:biocol.org:col:35275" box="[762,850,998,1024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="4" pageNumber="5">MNCN</collectionCode>
(
<collectionCode id="B0A96AFC3C24FF9FFC8AFC5CFC5B922C" LSID="urn:lsid:biocol.org:col:15536" box="[865,931,998,1024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="5">DNA</collectionCode>
and tissue collection) and the
<collectionCode id="B0A96AFC3C24FF9FFAECFC5CFAC2922C" box="[1287,1338,998,1024]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="4" pageNumber="5">IBE</collectionCode>
.
</paragraph>
<caption id="82C7A2B13C24FF9FFF7CF819FABA9196" box="[151,1346,1955,1979]" httpUri="https://zenodo.org/record/193455/files/figure.png" pageId="4" pageNumber="5" targetBox="[157,1428,1088,1924]" targetPageId="4">
<paragraph id="D607F2393C24FF9FFF7CF819FABA9196" blockId="4.[151,1346,1955,1979]" box="[151,1346,1955,1979]" pageId="4" pageNumber="5">
<emphasis id="E4CC2E2B3C24FF9FFF7CF819FEBA9197" bold="true" box="[151,322,1955,1979]" pageId="4" pageNumber="5">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="11B889BA3C24FF9FFE2BF819FD5A9196" box="[448,674,1955,1978]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C24FF9FFE2BF819FD5A9196" box="[448,674,1955,1978]" italics="true" pageId="4" pageNumber="5">Ochthebius aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C24FF9FFD43F819FD259196" box="[680,733,1955,1979]" pageId="4" pageNumber="5" rank="species">
<emphasis id="E4CC2E2B3C24FF9FFD43F819FD2E9197" bold="true" box="[680,726,1955,1979]" pageId="4" pageNumber="5">sp.n</emphasis>
.
</taxonomicNameLabel>
, 1) male (paratype, NMW); 2) female (paratype, NMW).
</paragraph>
</caption>
<caption id="82C7A2B13C25FF9EFF7CF8D3FBBB91C5" httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="5" pageNumber="6" targetBox="[176,1428,186,1866]" targetPageId="5">
<paragraph id="D607F2393C25FF9EFF7CF8D3FBBB91C5" blockId="5.[151,1436,1897,2025]" pageId="5" pageNumber="6">
<emphasis id="E4CC2E2B3C25FF9EFF7CF8D3FEA291AD" bold="true" box="[151,346,1897,1921]" pageId="5" pageNumber="6">FIGURES 310.</emphasis>
Aedeagus of
<taxonomicName id="11B889BA3C25FF9EFE12F8D0FD7D91AD" box="[505,645,1898,1921]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C25FF9EFE12F8D0FD7D91AD" box="[505,645,1898,1921]" italics="true" pageId="5" pageNumber="6">O. aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C25FF9EFD66F8D3FD3F91AD" box="[653,711,1897,1921]" pageId="5" pageNumber="6" rank="species">
<emphasis id="E4CC2E2B3C25FF9EFD66F8D3FD3F91AD" bold="true" box="[653,711,1897,1921]" pageId="5" pageNumber="6">sp.n.</emphasis>
</taxonomicNameLabel>
3) lateral and 4) ventral view (Paratype from type locality);
<taxonomicName id="11B889BA3C25FF9EFA6AF8D0FEEA9188" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C25FF9EFA6AF8D0FEEA9188" italics="true" pageId="5" pageNumber="6">O. legionensis</emphasis>
</taxonomicName>
5) lateral and 6) ventral view (voucher specimen MNCN-AI906);
<taxonomicName id="11B889BA3C25FF9EFC19F837FB7E9188" box="[1010,1158,1933,1956]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C25FF9EFC19F837FB7E9188" box="[1010,1158,1933,1956]" italics="true" pageId="5" pageNumber="6">O. exsculptus</emphasis>
</taxonomicName>
(northern clade, voucher specimen MNCN-AI1069) 7) lateral and 8) ventral view; and
<taxonomicName id="11B889BA3C25FF9EFCACF815FC2591EA" box="[839,989,1967,1990]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C25FF9EFCACF815FC2591EA" box="[839,989,1967,1990]" italics="true" pageId="5" pageNumber="6">O. exsculptus</emphasis>
</taxonomicName>
(southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm.
</paragraph>
</caption>
<caption id="82C7A2B13C26FF9DFF7CFD47FDE79576" httpUri="https://zenodo.org/record/193457/files/figure.png" pageId="6" pageNumber="7" targetBox="[404,1181,198,736]" targetPageId="6">
<paragraph id="D607F2393C26FF9DFF7CFD47FDE79576" blockId="6.[151,1437,765,858]" pageId="6" pageNumber="7">
<emphasis id="E4CC2E2B3C26FF9DFF7CFD47FEA69539" bold="true" box="[151,350,765,789]" pageId="6" pageNumber="7">FIGURES 1116.</emphasis>
Last abdominal tergites of
<taxonomicName id="11B889BA3C26FF9DFD63FD44FCF39539" box="[648,779,766,789]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C26FF9DFD63FD44FCF39539" box="[648,779,766,789]" italics="true" pageId="6" pageNumber="7">O. aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C26FF9DFCF8FD47FCB19539" box="[787,841,765,789]" pageId="6" pageNumber="7" rank="species">
<emphasis id="E4CC2E2B3C26FF9DFCF8FD47FCB19539" bold="true" box="[787,841,765,789]" pageId="6" pageNumber="7">sp.n.</emphasis>
</taxonomicNameLabel>
11) ventral, 12) lateral and 13) caudal views (paratype from type locality) and
<taxonomicName id="11B889BA3C26FF9DFE75FC9AFDC5951B" box="[414,573,800,823]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C26FF9DFE75FC9AFDC5951B" box="[414,573,800,823]" italics="true" pageId="6" pageNumber="7">O. legionensis</emphasis>
</taxonomicName>
14) ventral, 15) lateral and 16) caudal views (León, Rioscuro, 30.6.1985 L.F. Valladares leg.). Scale bar, 0.25 mm.
</paragraph>
</caption>
<caption id="82C7A2B13C26FF9DFF7CF838FC6F9190" httpUri="https://zenodo.org/record/193458/files/figure.png" pageId="6" pageNumber="7" targetBox="[155,1429,917,1894]" targetPageId="6">
<paragraph id="D607F2393C26FF9DFF7CF838FC6F9190" blockId="6.[151,1436,1922,1980]" pageId="6" pageNumber="7">
<emphasis id="E4CC2E2B3C26FF9DFF7CF838FED391B6" bold="true" box="[151,299,1922,1946]" pageId="6" pageNumber="7">FIGURE 17.</emphasis>
Distribution map of the Iberian species of the
<taxonomicName id="11B889BA3C26FF9DFCD1F838FB3391B5" box="[826,1227,1922,1945]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="exsculptus" subGenus="Enicocerus">
<emphasis id="E4CC2E2B3C26FF9DFCD1F838FC4091B5" box="[826,952,1922,1945]" italics="true" pageId="6" pageNumber="7">Ochthebius</emphasis>
(
<emphasis id="E4CC2E2B3C26FF9DFC22F838FBBD91B5" box="[969,1093,1922,1945]" italics="true" pageId="6" pageNumber="7">Enicocerus</emphasis>
)
<emphasis id="E4CC2E2B3C26FF9DFBBCF838FB3391B5" box="[1111,1227,1922,1945]" italics="true" pageId="6" pageNumber="7">exsculptus</emphasis>
</taxonomicName>
group. With black circles, localities from which at least one specimen has been sequenced.
</paragraph>
</caption>
<caption id="82C7A2B13C27FF9CFF7CF80CFB6091DC" httpUri="https://zenodo.org/record/193459/files/figure.png" pageId="7" pageNumber="8" targetBox="[256,1331,180,1942]" targetPageId="7">
<paragraph id="D607F2393C27FF9CFF7CF80CFB6091DC" blockId="7.[151,1436,1974,2032]" pageId="7" pageNumber="8">
<emphasis id="E4CC2E2B3C27FF9CFF7CF80CFEA691E2" bold="true" box="[151,350,1974,1998]" pageId="7" pageNumber="8">FIGURES 1819.</emphasis>
Type locality of
<taxonomicName id="11B889BA3C27FF9CFDFCF80DFD6291E2" box="[535,666,1975,1998]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C27FF9CFDFCF80DFD6291E2" box="[535,666,1975,1998]" italics="true" pageId="7" pageNumber="8">O. aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C27FF9CFD49F80CFD2F91E2" box="[674,727,1974,1998]" pageId="7" pageNumber="8" rank="species">
<emphasis id="E4CC2E2B3C27FF9CFD49F80CFD2F91E2" bold="true" box="[674,727,1974,1998]" pageId="7" pageNumber="8">sp.n.</emphasis>
</taxonomicNameLabel>
(Arroyo de la Laguna Grande de Peñalara), 18) top, 2 June 2007, with D.T. Bilton and F. Bameul; 19) bottom, 20 July 2006, detail of the microhabitat (fotos, IR).
</paragraph>
</caption>
<paragraph id="D607F2393C28FF93FF2DFF2DFE0396F4" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFF2DFE3F969D" bold="true" box="[198,455,151,177]" pageId="8" pageNumber="9">Additional material.</emphasis>
<specimenCount id="C0BE39B03C28FF93FE3BFF22FDDA969E" box="[464,546,152,178]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(
<collectionCode id="B0A96AFC3C28FF93FDDFFF22FD8D969E" box="[564,629,152,178]" pageId="8" pageNumber="9">CVL</collectionCode>
): “Madrid (Norte), 1978 \ E. Roy leg.” (not designated as
<typeStatus id="09034C9B3C28FF93FADCFF22FA64969E" box="[1335,1436,152,178]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
due to the imprecise location).
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C28FF93FF2DFF5FFAAE9532" pageId="8" pageNumber="9" type="description">
<paragraph id="D607F2393C28FF93FF2DFF5FFAAE9532" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFF5FFEA696D3" bold="true" box="[198,350,229,255]" pageId="8" pageNumber="9">Description.</emphasis>
Habitus as in
<figureCitation id="4E83EEBC3C28FF93FDF4FF5FFD7796D3" box="[543,655,229,255]" captionStart="FIGURES 1 2" captionStartId="4.[151,269,1955,1979]" captionTargetBox="[157,1428,1088,1924]" captionTargetId="figure@4.[151,1436,1079,1930]" captionTargetPageId="4" captionText="FIGURES 1 2. Habitus of Ochthebius aguilerai sp. n., 1) male (paratype, NMW); 2) female (paratype, NMW)." httpUri="https://zenodo.org/record/193455/files/figure.png" pageId="8" pageNumber="9">
Figs 1
<emphasis id="E4CC2E2B3C28FF93FD85FF5FFD8596D3" bold="true" box="[622,637,229,255]" pageId="8" pageNumber="9"></emphasis>
2
</figureCitation>
. Body length
<quantity id="11405FDC3C28FF93FCAEFF5FFC2396D3" box="[837,987,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="8" pageNumber="9" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">
1.9
<emphasis id="E4CC2E2B3C28FF93FC82FF5FFC8096D3" bold="true" box="[873,888,229,255]" pageId="8" pageNumber="9"></emphasis>
2.1 mm
</quantity>
. Colour black, with strong greenish metallic reflections. Body appendages yellowish brown; apical half of antennae, palpi, knees and tarsi darker. Upper surface very sparsely covered with short whitish adpressed pubescence. Anterior margin of the labrum emarginated; anterior lobes transverse, exterior angle frequently with a spine. Labrum and clypeus with large punctures; surface with a distinct microreticulation. Frons more densely punctate, with a shagreened appearance. Pronotum of female and some males not very convex, roughly trapezoidal, admedian fovea and sulcus well impressed, median sulcus reaching anterior and posterior margins; pronotal punctures strongly impressed, sparse, space between punctures usually smooth and glabrous. In some males pronotum strongly convex, with admedian foveas more elongated, surface with less impressed punctation (
<figureCitation id="4E83EEBC3C28FF93FB5BFDA6FB0E941A" box="[1200,1270,540,566]" captionStart="FIGURES 1 2" captionStartId="4.[151,269,1955,1979]" captionTargetBox="[157,1428,1088,1924]" captionTargetId="figure@4.[151,1436,1079,1930]" captionTargetPageId="4" captionText="FIGURES 1 2. Habitus of Ochthebius aguilerai sp. n., 1) male (paratype, NMW); 2) female (paratype, NMW)." httpUri="https://zenodo.org/record/193455/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). Elytra oval, strongly convex, with ten strongly impressed regular striae; second interval flat and smooth, specially in male with convex pronotum; third interval strongly raised near the basis. Suture very depressed anterior to and strongly raised posterior to middle of elytra, so that in lateral view each elytron show a distinct median depression. Aedeagus as in Figs (3, 4), long and slender. Main piece, in ventral view, distinctly narrowed behind insertion of parameres, in lateral view, apex narrow and tapered; parameres short, adjoining median lobe. Female tergite X (
<figureCitation id="4E83EEBC3C28FF93FE47FCBEFDC89532" box="[428,560,772,798]" captionStart="FIGURES 11 16" captionStartId="6.[151,270,765,789]" captionTargetBox="[404,1181,198,736]" captionTargetId="figure@6.[395,1191,194,741]" captionTargetPageId="6" captionText="FIGURES 11 16. Last abdominal tergites of O. aguilerai sp. n. 11) ventral, 12) lateral and 13) caudal views (paratype from type locality) and O. legionensis 14) ventral, 15) lateral and 16) caudal views (León, Rioscuro, 30.6. 1985 L. F. Valladares leg.). Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193457/files/figure.png" pageId="8" pageNumber="9">
Figs 11
<emphasis id="E4CC2E2B3C28FF93FDE8FCBEFDEA9532" bold="true" box="[515,530,772,798]" pageId="8" pageNumber="9"></emphasis>
13
</figureCitation>
) with a small medial protuberance visible in lateral and caudal view.
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C28FF93FF2DFC91FEE19279" pageId="8" pageNumber="9" type="distribution">
<paragraph id="D607F2393C28FF93FF2DFC91FEE19279" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFC91FEA79569" bold="true" box="[198,351,811,837]" pageId="8" pageNumber="9">Distribution</emphasis>
. Mountains of the Spanish Sistema Central, from north-east of the province of Cáceres to Sierra de Gredos and Guadarrama (
<figureCitation id="4E83EEBC3C28FF93FDD5FCE8FD6F9540" box="[574,663,850,876]" captionStart="FIGURE 17" captionStartId="6.[151,258,1922,1946]" captionTargetBox="[155,1429,917,1894]" captionTargetId="figure@6.[151,1436,912,1898]" captionTargetPageId="6" captionText="FIGURE 17. Distribution map of the Iberian species of the Ochthebius (Enicocerus) exsculptus group. With black circles, localities from which at least one specimen has been sequenced." httpUri="https://zenodo.org/record/193458/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
). The record of
<taxonomicName id="11B889BA3C28FF93FCB3FCE8FBFA9547" box="[856,1026,850,875]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C28FF93FCB3FCE8FBFA9547" box="[856,1026,850,875]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FCB3FCE8FC899547" box="[856,881,850,875]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
in
<bibRefCitation id="B2298FC83C28FF93FBC0FCE8FA639540" box="[1067,1435,850,876]" pageId="8" pageNumber="12" refString="Garrido-Gonzalez, J., Diaz-Pazos, J. A. &amp; Regil-Cueto, J. A. (1994) Coleopteros acuaticos de Extremadura 2 (Hydraenidae, Hydrochidae, Helophoridae, Hydrophilidae, Dryopidae y Elmidae). Boletin de la Asociacion Espanola de Entomologia, 18, 113 - 133." type="journal article">
Garrido-González
<emphasis id="E4CC2E2B3C28FF93FAE2FCE8FABD9547" box="[1289,1349,850,875]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(1994)
</bibRefCitation>
from NE Cáceres corresponds to this species (see
<typeStatus id="09034C9B3C28FF93FCFDFCC3FCAC95BF" box="[790,852,889,915]" pageId="8" pageNumber="9">Type</typeStatus>
Material). Although there are no genetic data available, according to the external morphology the populations of the north Iberian System (Burgos and
<collectingCountry id="AEAFB2A93C28FF93FF7CFC7CFF2495CC" box="[151,220,966,992]" name="Syria" pageId="8" pageNumber="9">Soria</collectingCountry>
,
<bibRefCitation id="B2298FC83C28FF93FF02FC7CFE0195CC" author="Valladares" box="[233,505,966,992]" pageId="8" pageNumber="13" refString="Valladares, L. F., Diaz, J. A. &amp; Garrido, J. (2000) Coleopteros acuaticos del Sistema Iberico Septentrional (Coleoptera: Haliplidae, Gyrinidae, Dytiscidae, Hydraenidae, Helophoridae, Hydrochidae, Hydrophilidae). Boletin de la Asociacion Espanola de Entomologia, 24 (3 - 4), 59 - 84." type="journal article" year="2000">
Valladares
<emphasis id="E4CC2E2B3C28FF93FE99FC7DFE4995CC" box="[370,433,967,992]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2000
</bibRefCitation>
) and Málaga (
<bibRefCitation id="B2298FC83C28FF93FD47FC7CFC8C95CC" author="Jach" box="[684,884,966,992]" pageId="8" pageNumber="12" refString="Jach, M. A., Diaz, J. A. &amp; Gayoso, A. (1999) &quot; Acciones Integradas &quot;: excursion to Andalucia (Spain: Malaga, Cadiz), October 1998 (Coleoptera: Hydroscaphidae, Haliplidae, Gyrinidae, Dytiscidae, Hydrochidae, Hydraenidae, Dryopidae, Elmidae). Koleopterologische Rundschau, 69, 171 - 181." type="journal article" year="1999">
Jäch
<emphasis id="E4CC2E2B3C28FF93FD06FC7DFCD595CC" box="[749,813,967,992]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
1999
</bibRefCitation>
) (
<figureCitation id="4E83EEBC3C28FF93FC64FC7CFC1295CC" box="[911,1002,966,992]" captionStart="FIGURE 17" captionStartId="6.[151,258,1922,1946]" captionTargetBox="[155,1429,917,1894]" captionTargetId="figure@6.[151,1436,912,1898]" captionTargetPageId="6" captionText="FIGURE 17. Distribution map of the Iberian species of the Ochthebius (Enicocerus) exsculptus group. With black circles, localities from which at least one specimen has been sequenced." httpUri="https://zenodo.org/record/193458/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
) would belong to
<taxonomicName id="11B889BA3C28FF93FB25FC7DFA8D95CC" box="[1230,1397,967,992]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FB25FC7DFA8D95CC" box="[1230,1397,967,992]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FB25FC7DFB1F95CC" box="[1230,1255,967,992]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
(J. Garrido, L.F. Valladares and M.A. Jäch, personal communication 2009). The records from Albarracín (Teruel) in
<bibRefCitation id="B2298FC83C28FF93FF5EFBAEFE519202" box="[181,425,1044,1070]" pageId="8" pageNumber="12" refString="d'Orchymont, A. (1941) Revision des Ochthebius Europeens du sous-genre Henicocerus Stephens, 1829 (Coleoptera Palpicornia). Bulletin du Musee royal d'Histoire naturelle de Belgique, 17 (12), 1 - 15." type="journal article">dOrchymont (1941)</bibRefCitation>
have been considered as
<taxonomicName id="11B889BA3C28FF93FD3FFBAEFC8B9201" box="[724,883,1044,1069]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FD3FFBAEFC8B9201" box="[724,883,1044,1069]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FD3FFBAEFD149201" box="[724,748,1044,1069]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
in
<figureCitation id="4E83EEBC3C28FF93FC72FBAEFC169202" box="[921,1006,1044,1070]" captionStart="FIGURE 17" captionStartId="6.[151,258,1922,1946]" captionTargetBox="[155,1429,917,1894]" captionTargetId="figure@6.[151,1436,912,1898]" captionTargetPageId="6" captionText="FIGURE 17. Distribution map of the Iberian species of the Ochthebius (Enicocerus) exsculptus group. With black circles, localities from which at least one specimen has been sequenced." httpUri="https://zenodo.org/record/193458/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
, although we have not examined the specimens.
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C28FF93FF2DFBDBFBAF93A7" pageId="8" pageNumber="9" type="etymology">
<paragraph id="D607F2393C28FF93FF2DFBDBFEB2928F" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFBDBFEB39257" bold="true" box="[198,331,1121,1147]" pageId="8" pageNumber="9">Etymology</emphasis>
. Named after our friend and water-beetler Pedro Aguilera, who died in an accident in
<date id="A206D4F93C28FF93FF7CFB33FEBE928F" box="[151,326,1161,1187]" pageId="8" pageNumber="9" value="2009-02">February 2009</date>
.
</paragraph>
<paragraph id="D607F2393C28FF93FF2DFB15FBAF93A7" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFB15FED092E5" bold="true" box="[198,296,1199,1225]" pageId="8" pageNumber="9">Ecology</emphasis>
. The new species is found in fast flowing, clean mountain streams in granitic substratum, usually on the surface of large semi-submerged rocks (
<figureCitation id="4E83EEBC3C28FF93FD2BFB6CFCB092DC" box="[704,840,1238,1264]" captionStart="FIGURES 18 19" captionStartId="7.[151,270,1974,1998]" captionTargetBox="[256,1331,180,1942]" captionTargetId="figure@7.[250,1337,172,1950]" captionTargetPageId="7" captionText="FIGURES 18 19. Type locality of O. aguilerai sp. n. (Arroyo de la Laguna Grande de Peñalara), 18) top, 2 June 2007, with D. T. Bilton and F. Bameul; 19) bottom, 20 July 2006, detail of the microhabitat (fotos, IR)." httpUri="https://zenodo.org/record/193459/files/figure.png" pageId="8" pageNumber="9">
Figs 18
<emphasis id="E4CC2E2B3C28FF93FCF2FB6CFCD092DC" bold="true" box="[793,808,1238,1264]" pageId="8" pageNumber="9"></emphasis>
19
</figureCitation>
). In the
<typeStatus id="09034C9B3C28FF93FC40FB6CFC2692DC" box="[939,990,1238,1264]" pageId="8" pageNumber="9">type</typeStatus>
locality (the one in which the species seems to be more abundant) it was collected in the company of an abundant and diverse fauna of
<taxonomicName id="11B889BA3C28FF93FAEDFB47FA64933B" box="[1286,1436,1277,1303]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
(
<taxonomicName id="11B889BA3C28FF93FF49FA9EFDDE9312" authority="d'Orchymont" authorityName="d'Orchymont" box="[162,550,1316,1342]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brachymera">
<emphasis id="E4CC2E2B3C28FF93FF49FA9EFF4F9311" box="[162,183,1316,1341]" italics="true" pageId="8" pageNumber="9">H</emphasis>
.
<emphasis id="E4CC2E2B3C28FF93FF27FA9EFE919311" box="[204,361,1316,1341]" italics="true" pageId="8" pageNumber="9">brachymera</emphasis>
dOrchymont
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FDDEFA9EFC7A9312" authority="d'Orchymont" authorityName="d'Orchymont" box="[565,898,1316,1342]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="corinna">
<emphasis id="E4CC2E2B3C28FF93FDDEFA9EFD3C9311" box="[565,708,1316,1341]" italics="true" pageId="8" pageNumber="9">H. corinna</emphasis>
dOrchymont
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FC7BFA9EFAF09312" authority="d'Orchymont" authorityName="d'Orchymont" box="[912,1288,1316,1342]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exasperata">
<emphasis id="E4CC2E2B3C28FF93FC7BFA9EFBB39311" box="[912,1099,1316,1341]" italics="true" pageId="8" pageNumber="9">H. exasperata</emphasis>
dOrchymont
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FAFDFA9EFEC09348" authority="d'Orchymont" authorityName="d'Orchymont" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="iberica">
<emphasis id="E4CC2E2B3C28FF93FAFDFA9EFA639311" box="[1302,1435,1316,1341]" italics="true" pageId="8" pageNumber="9">H. iberica</emphasis>
dOrchymont
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FEA8FAF1FD929348" authority="Ganglbauer" authorityName="Ganglbauer" box="[323,618,1354,1380]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="E4CC2E2B3C28FF93FEA8FAF1FE219348" box="[323,473,1355,1380]" italics="true" pageId="8" pageNumber="9">H. hispanica</emphasis>
Ganglbauer
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FD9CFAF1FC8D9348" authority="Rey" authorityName="Rey" box="[631,885,1354,1380]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sharpi">
<emphasis id="E4CC2E2B3C28FF93FD9CFAF1FCC69348" box="[631,830,1355,1380]" italics="true" pageId="8" pageNumber="9">Hydraena sharpi</emphasis>
Rey
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FC69FAF1FAF29348" authority="Thunberg" authorityName="Thunberg" box="[898,1290,1354,1380]" class="Insecta" family="Hydraenidae" genus="Limnebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="truncatellus">
<emphasis id="E4CC2E2B3C28FF93FC69FAF1FB779348" box="[898,1167,1355,1380]" italics="true" pageId="8" pageNumber="9">Limnebius truncatellus</emphasis>
Thunberg
</taxonomicName>
,
<taxonomicName id="11B889BA3C28FF93FAFDFAF1FEAE93A7" authority="Kuwert" authorityName="Kuwert" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="heydeni">
<emphasis id="E4CC2E2B3C28FF93FAFDFAF1FF0A93A6" italics="true" pageId="8" pageNumber="9">Ochthebius heydeni</emphasis>
Kuwert
</taxonomicName>
), all of them Iberian endemics except
<taxonomicName id="11B889BA3C28FF93FCFFFACBFBAB93A7" authority="Jach 2004" authorityName="Jach" authorityYear="2004" box="[788,1107,1393,1419]" class="Insecta" family="Hydraenidae" genus="Limnebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="truncatellus">
<emphasis id="E4CC2E2B3C28FF93FCFFFACBFC4793A6" box="[788,959,1393,1418]" italics="true" pageId="8" pageNumber="9">L. truncatellus</emphasis>
(
<bibRefCitation id="B2298FC83C28FF93FC25FACBFBB393A7" author="Jach" box="[974,1099,1393,1419]" pageId="8" pageNumber="12" refString="Jach, M. A. (2004) Hydraenidae. In: I. Lobl &amp; A. Smetana (Eds), Catalogue of Palaearctic Coleoptera. Volume 2: Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, pp. 102 - 122." type="journal article" year="2004">Jäch 2004</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C28FF92FF2DFA22FD6A96F4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="D607F2393C28FF93FF2DFA22FCFE90EE" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FF2DFA22FE3C939E" bold="true" box="[198,452,1432,1458]" pageId="8" pageNumber="9">Comparative notes.</emphasis>
<taxonomicName id="11B889BA3C28FF93FE3EFA22FD2F939D" box="[469,727,1432,1457]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FE3EFA22FD2F939D" box="[469,727,1432,1457]" italics="true" pageId="8" pageNumber="9">Ochthebius aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FD03FA22FCDB939E" box="[744,803,1432,1458]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FD03FA22FCDB939E" bold="true" box="[744,803,1432,1458]" pageId="8" pageNumber="9">sp.n.</emphasis>
</taxonomicNameLabel>
is very similar to
<taxonomicName id="11B889BA3C28FF93FBCCFA22FB20939D" box="[1063,1240,1432,1457]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C28FF93FBCCFA22FB20939D" box="[1063,1240,1432,1457]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FBCCFA22FBC7939D" box="[1063,1087,1432,1457]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
in its external morphology, but females of both species can easily distinguished by the shape of the last terguite, specially when observed in lateral and caudal view (
<figureCitation id="4E83EEBC3C28FF93FD45FA5CFCC7902C" box="[686,831,1509,1536]" captionStart="FIGURES 11 16" captionStartId="6.[151,270,765,789]" captionTargetBox="[404,1181,198,736]" captionTargetId="figure@6.[395,1191,194,741]" captionTargetPageId="6" captionText="FIGURES 11 16. Last abdominal tergites of O. aguilerai sp. n. 11) ventral, 12) lateral and 13) caudal views (paratype from type locality) and O. legionensis 14) ventral, 15) lateral and 16) caudal views (León, Rioscuro, 30.6. 1985 L. F. Valladares leg.). Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193457/files/figure.png" pageId="8" pageNumber="9">
Figs 11
<emphasis id="E4CC2E2B3C28FF93FCE7FA5FFCE393D3" bold="true" box="[780,795,1509,1535]" pageId="8" pageNumber="9"></emphasis>
16
</figureCitation>
). We have not found any other constant external character to separate these two species, although
<taxonomicName id="11B889BA3C28FF93FD07F9B7FC78900A" box="[748,896,1549,1574]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FD07F9B7FC78900A" box="[748,896,1549,1574]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FD07F9B7FCFD900A" box="[748,773,1549,1574]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. aguilerai
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FC62F9B6FC3D900A" box="[905,965,1548,1574]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FC62F9B6FC3D900A" bold="true" box="[905,965,1548,1574]" pageId="8" pageNumber="9">sp.n.</emphasis>
</taxonomicNameLabel>
tends to be slightly smaller and with a more strongly puncturated pronotum. The aedeagus of
<taxonomicName id="11B889BA3C28FF93FCA8F989FC0D9060" box="[835,1013,1587,1612]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C28FF93FCA8F989FC0D9060" box="[835,1013,1587,1612]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FCA8F989FCA49060" box="[835,860,1587,1612]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
(
<figureCitation id="4E83EEBC3C28FF93FBEDF989FB8C9061" box="[1030,1140,1587,1613]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="8" pageNumber="9">
Figs 5
<emphasis id="E4CC2E2B3C28FF93FBBEF989FB9C9061" bold="true" box="[1109,1124,1587,1613]" pageId="8" pageNumber="9"></emphasis>
6
</figureCitation>
) differs from that of
<taxonomicName id="11B889BA3C28FF93FA94F989FEF3905F" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<collectingCountry id="AEAFB2A93C28FF93FA94F989FA6C9060" box="[1407,1428,1587,1612]" name="Iceland" pageId="8" pageNumber="9">
<emphasis id="E4CC2E2B3C28FF93FA94F989FA6C9060" box="[1407,1428,1587,1612]" italics="true" pageId="8" pageNumber="9">O</emphasis>
</collectingCountry>
.
<emphasis id="E4CC2E2B3C28FF93FF7CF9E0FEF3905F" box="[151,267,1626,1651]" italics="true" pageId="8" pageNumber="9">aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FEF8F9E0FEAA9058" box="[275,338,1626,1652]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FEF8F9E0FEAA9058" bold="true" box="[275,338,1626,1652]" pageId="8" pageNumber="9">n.sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="4E83EEBC3C28FF93FE8FF9E0FE2D9058" box="[356,469,1626,1652]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="8" pageNumber="9">
Figs 3
<emphasis id="E4CC2E2B3C28FF93FE5FF9E0FE3B9058" bold="true" box="[436,451,1626,1652]" pageId="8" pageNumber="9"></emphasis>
4
</figureCitation>
) by its more robust main piece, ventrally less narrowed behind insertions of parameres, and with a broadly truncate apex. The apical lobe of
<taxonomicName id="11B889BA3C28FF93FC67F93BFBE390B6" box="[908,1051,1665,1690]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FC67F93BFBE390B6" box="[908,1051,1665,1690]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FC67F93BFC5C90B6" box="[908,932,1665,1690]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. aguilerai
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FBC8F93AFBA690B6" box="[1059,1118,1664,1690]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FBC8F93AFBA690B6" bold="true" box="[1059,1118,1664,1690]" pageId="8" pageNumber="9">n.sp.</emphasis>
</taxonomicNameLabel>
is more slender and with a less sharp bend that that of
<taxonomicName id="11B889BA3C28FF93FE3CF912FD8790ED" box="[471,639,1704,1729]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C28FF93FE3CF912FD8790ED" box="[471,639,1704,1729]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FE3CF912FE1790ED" box="[471,495,1704,1729]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
(
<figureCitation id="4E83EEBC3C28FF93FD64F912FD0090EE" box="[655,760,1703,1730]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="8" pageNumber="9">
Figs 3
<emphasis id="E4CC2E2B3C28FF93FD32F91DFD1F90ED" bold="true" box="[729,743,1703,1729]" pageId="8" pageNumber="9"></emphasis>
6
</figureCitation>
).
</paragraph>
<paragraph id="D607F2393C28FF93FF2DF975FC219171" blockId="8.[151,1437,151,2004]" pageId="8" pageNumber="9">
<taxonomicName id="11B889BA3C28FF93FF2DF975FE4690C4" box="[198,446,1743,1768]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FF2DF975FE4690C4" box="[198,446,1743,1768]" italics="true" pageId="8" pageNumber="9">Ochthebius aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FE2FF974FE0490C4" box="[452,508,1742,1768]" pageId="8" pageNumber="9" rank="species">n.sp.</taxonomicNameLabel>
and
<taxonomicName id="11B889BA3C28FF93FDDFF975FD2390C4" box="[564,731,1743,1768]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="legionensis">
<emphasis id="E4CC2E2B3C28FF93FDDFF975FD2390C4" box="[564,731,1743,1768]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FDDFF975FDB490C4" box="[564,588,1743,1768]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. legionensis
</emphasis>
</taxonomicName>
can be easily distinguished from the other Iberian species of the
<taxonomicName id="11B889BA3C28FF93FF29F94CFE989123" box="[194,352,1782,1807]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FF29F94CFE989123" box="[194,352,1782,1807]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FF29F94CFF229123" box="[194,218,1782,1807]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
group by the presence of an elytral depression. The apex of the median lobe of the aedeagus of
<taxonomicName id="11B889BA3C28FF93FF53F8A6FEB49119" box="[184,332,1820,1845]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FF53F8A6FEB49119" box="[184,332,1820,1845]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FF53F8A6FF299119" box="[184,209,1820,1845]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. aguilerai
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FEBFF8A6FE68911A" box="[340,400,1820,1846]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FEBFF8A6FE68911A" bold="true" box="[340,400,1820,1846]" pageId="8" pageNumber="9">n.sp.</emphasis>
</taxonomicNameLabel>
is less truncated in ventral view than that of
<taxonomicName id="11B889BA3C28FF93FC58F8A6FBAE9119" box="[947,1110,1820,1845]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FC58F8A6FBAE9119" box="[947,1110,1820,1845]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FC58F8A6FC349119" box="[947,972,1820,1845]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
, and less pointed in lateral view (
<figureCitation id="4E83EEBC3C28FF93FF34F8F9FE6C9171" box="[223,404,1858,1885]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="8" pageNumber="9">
Figs 3
<emphasis id="E4CC2E2B3C28FF93FEC2F8F8FEC09170" bold="true" box="[297,312,1858,1884]" pageId="8" pageNumber="9"></emphasis>
4, 7
<emphasis id="E4CC2E2B3C28FF93FE8FF8F8FE8B9170" bold="true" box="[356,371,1858,1884]" pageId="8" pageNumber="9"></emphasis>
10
</figureCitation>
). The apical lobe of both species are very similar.
</paragraph>
<paragraph id="D607F2393C28FF92FF2DF8D0FD6A96F4" blockId="8.[151,1437,151,2004]" lastBlockId="9.[151,1436,152,410]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The size of the species of the
<taxonomicName id="11B889BA3C28FF93FDC3F8D0FD3F91AF" box="[552,711,1898,1923]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FDC3F8D0FD3F91AF" box="[552,711,1898,1923]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FDC3F8D0FDB891AF" box="[552,576,1898,1923]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
group is rather variable, and there is a significant correlation between body and aedeagus size (for Iberian specimens of the three species r2 = 0.74, p &lt;0.01, n = 13). Specimens of
<taxonomicName id="11B889BA3C28FF93FEA0F800FE1E91FF" box="[331,486,1978,2003]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C28FF93FEA0F800FE1E91FF" box="[331,486,1978,2003]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FEA0F800FE9D91FF" box="[331,357,1978,2003]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. aguilerai
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C28FF93FE04F800FDD691F8" box="[495,558,1978,2004]" pageId="8" pageNumber="9" rank="species">
<emphasis id="E4CC2E2B3C28FF93FE04F800FDD691F8" bold="true" box="[495,558,1978,2004]" pageId="8" pageNumber="9">sp.n.</emphasis>
</taxonomicNameLabel>
and of the southern clade of
<taxonomicName id="11B889BA3C28FF93FC4DF800FBAA91FF" box="[934,1106,1978,2003]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="exsculptus">
<emphasis id="E4CC2E2B3C28FF93FC4DF800FBAA91FF" box="[934,1106,1978,2003]" italics="true" pageId="8" pageNumber="9">
<collectingCountry id="AEAFB2A93C28FF93FC4DF800FC3891FF" box="[934,960,1978,2003]" name="Iceland" pageId="8" pageNumber="9">O</collectingCountry>
. exsculptus
</emphasis>
</taxonomicName>
(see below) seem to have proportionally smaller aedeagus, but in any case, and due to the large overlap between species, these would be statistichal trends without diagnostic value.
</paragraph>
</subSubSection>
<subSubSection id="9EA2A1B23C29FF92FF2DFF5CFD5697B6" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="D607F2393C29FF92FF2DFF5CFD5697B6" blockId="9.[151,1436,152,410]" pageId="9" pageNumber="10">
<taxonomicName id="11B889BA3C29FF92FF2DFF5CFE3996D3" ID-CoL="6SFRK" box="[198,449,230,255]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C29FF92FF2DFF5CFE3996D3" box="[198,449,230,255]" italics="true" pageId="9" pageNumber="10">Ochthebius aguilerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C29FF92FE21FF5FFDFD96D3" box="[458,517,229,255]" pageId="9" pageNumber="10" rank="species">
<emphasis id="E4CC2E2B3C29FF92FE21FF5FFDFD96D3" bold="true" box="[458,517,229,255]" pageId="9" pageNumber="10">sp.n.</emphasis>
</taxonomicNameLabel>
seems to be in some characters closer to the Central European
<taxonomicName id="11B889BA3C29FF92FAEBFF5CFA6D96D3" ID-CoL="6SG5H" box="[1280,1429,230,255]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="halbherri">
<emphasis id="E4CC2E2B3C29FF92FAEBFF5CFA6D96D3" box="[1280,1429,230,255]" italics="true" pageId="9" pageNumber="10">
<collectingCountry id="AEAFB2A93C29FF92FAEBFF5CFAE096D3" box="[1280,1304,230,255]" name="Iceland" pageId="9" pageNumber="10">O</collectingCountry>
. halbherri
</emphasis>
</taxonomicName>
. Both species share an aedeagus with a dorsally constricted main piece, ending in a narrow apex, although both species can be easily distinguished by the shape of the apex of the median lobe (see
<bibRefCitation id="B2298FC83C29FF92FB6DFE88FAFB9760" author="Jach" box="[1158,1283,306,332]" pageId="9" pageNumber="12" refString="Jach, M. A. (1992) Revision of the Palearctic species of the genus Ochthebius Leach. 7. The subgenus Enicocerus Stephens (Coleoptera: Hydraenidae). Elytron, 5 (1991), 139 - 158." type="journal article" year="1992">Jäch 1992</bibRefCitation>
:
<figureCitation id="4E83EEBC3C29FF92FAE5FE88FAAF9760" box="[1294,1367,306,332]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
). The female tergite X is also similar in both species (see
<bibRefCitation id="B2298FC83C29FF92FCE2FEE3FC70975F" author="Jach" box="[777,904,345,371]" pageId="9" pageNumber="12" refString="Jach, M. A. (1992) Revision of the Palearctic species of the genus Ochthebius Leach. 7. The subgenus Enicocerus Stephens (Coleoptera: Hydraenidae). Elytron, 5 (1991), 139 - 158." type="journal article" year="1992">Jäch 1992</bibRefCitation>
:
<figureCitation id="4E83EEBC3C29FF92FC7FFEE3FC26975F" box="[916,990,345,371]" captionStart="FIGURES 3 10" captionStartId="5.[151,275,1897,1921]" captionTargetBox="[176,1428,186,1866]" captionTargetId="figure@5.[164,1449,174,1875]" captionTargetPageId="5" captionText="FIGURES 3 10. Aedeagus of O. aguilerai sp. n. 3) lateral and 4) ventral view (Paratype from type locality); O. legionensis 5) lateral and 6) ventral view (voucher specimen MNCN-AI 906); O. exsculptus (northern clade, voucher specimen MNCN-AI 1069) 7) lateral and 8) ventral view; and O. exsculptus (southern clade, Granada, Río Genil) 9) lateral and 10) ventral view. See Table 1 for details of the localities. Scale bar, 0.25 mm." httpUri="https://zenodo.org/record/193456/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
), although the apical protuberance is distinctly less developed in
<taxonomicName id="11B889BA3C29FF92FE37FE3AFD9497B5" ID-CoL="6SFRK" box="[476,620,384,409]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aguilerai" status="sp. nov.">
<emphasis id="E4CC2E2B3C29FF92FE37FE3AFD9497B5" box="[476,620,384,409]" italics="true" pageId="9" pageNumber="10">
<collectingCountry id="AEAFB2A93C29FF92FE37FE3AFE0C97B5" box="[476,500,384,409]" name="Iceland" pageId="9" pageNumber="10">O</collectingCountry>
. aguilerai
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFFF93503C29FF92FD98FE3AFD5697B6" box="[627,686,384,410]" pageId="9" pageNumber="10" rank="species">
<emphasis id="E4CC2E2B3C29FF92FD98FE3AFD5697B6" bold="true" box="[627,686,384,410]" pageId="9" pageNumber="10">sp.n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>