treatments-xml/data/18/32/87/183287D983637E35FF13669A2AD6FE92.xml

<document id="E2FCD6FA3347FDC1B07D6C00894D4460" ID-CLB-Dataset="22489" ID-DOI="10.11646/zootaxa.4767.4.4" ID-GBIF-Dataset="a6094fee-c9fe-423a-9990-3e9e8b939267" ID-ISSN="1175-5334" ID-Zenodo-Dep="3774917" ID-ZooBank="urn:lsid:zoobank.org:pub:80B195FA-02DE-4B4C-80AB-9DDC62E73875" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588142504801" checkinUser="plazi" docAuthor="Bahder, Brian W., Barrantes Barrantes, Edwin A., Zumbado Echavarria, Co. A., Mou, De-Fen, Helmick, Ericka E. &amp; Bartlett, Charles R." docDate="2020" docId="183287D983637E35FF13669A2AD6FE92" docLanguage="en" docName="zootaxa.4767.4.4.pdf" docOrigin="Zootaxa 4767 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Haplaxius dougwalshi Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett 2020, sp. n." docType="treatment" docVersion="6" lastPageNumber="551" masterDocId="E40BFFA183607E3DFF84626F2E79FFD1" masterDocTitle="A new species of planthopper in the genus Haplaxius (Hemiptera: Auchenorrhyncha: Fulgoroidea: Cixiidae) on palms in Costa Rica and a new country record for Haplaxius skarphion" masterLastPageNumber="552" masterPageNumber="543" pageNumber="546" updateTime="1698838000729" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AC804DCDAA4483E33D41D4ACC4027183">A new species of planthopper in the genus Haplaxius (Hemiptera: Auchenorrhyncha: Fulgoroidea: Cixiidae) on palms in Costa Rica and a new country record for Haplaxius skarphion</mods:title>
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<mods:namePart id="82C95602DB3C6F7CAB81484F2F7A9776">Bahder, Brian W.</mods:namePart>
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<mods:namePart id="C1F49B0316D420FE7F01CB4E9296AD1E">Barrantes Barrantes, Edwin A.</mods:namePart>
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<mods:namePart id="D5958AD123DA78CCA112E5E609BDECA8">Mou, De-Fen</mods:namePart>
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<mods:namePart id="47D51EFF391CAB2C60C6C4AB6A5667F2">Helmick, Ericka E.</mods:namePart>
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<treatment id="183287D983637E35FF13669A2AD6FE92" ID-DOI="http://doi.org/10.5281/zenodo.3796689" ID-GBIF-Taxon="163875146" ID-Zenodo-Dep="3796689" LSID="urn:lsid:plazi:treatment:183287D983637E35FF13669A2AD6FE92" httpUri="http://treatment.plazi.org/id/183287D983637E35FF13669A2AD6FE92" lastPageId="8" lastPageNumber="551" pageId="3" pageNumber="546">
<subSubSection id="D881654483637E3EFF13669A2F89FAC1" box="[151,496,1269,1296]" pageId="3" pageNumber="546" type="nomenclature">
<paragraph id="902436CF83637E3EFF13669A2F89FAC1" blockId="3.[151,496,1269,1331]" box="[151,496,1269,1296]" pageId="3" pageNumber="546">
<heading id="CB6C81A383637E3EFF13669A2F89FAC1" bold="true" box="[151,496,1269,1296]" fontSize="10" level="2" pageId="3" pageNumber="546" reason="2">
<emphasis id="A2EFEADD83637E3EFF13669A2F89FAC1" bold="true" box="[151,496,1269,1296]" pageId="3" pageNumber="546">
<taxonomicName id="579B4D4C83637E3EFF13669A2FDEFAC1" ID-CoL="3JL28" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[151,423,1269,1296]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83637E3EFF13669A2FDEFAC1" bold="true" box="[151,423,1269,1296]" italics="true" pageId="3" pageNumber="546">Haplaxius dougwalshi</emphasis>
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<taxonomicNameLabel id="B9DC57A683637E3EFE2A66992F89FAC1" box="[430,496,1270,1296]" pageId="3" pageNumber="546" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="D881654483637E3EFF1367762F56FAE2" box="[151,303,1305,1331]" pageId="3" pageNumber="546" type="description">
<paragraph id="902436CF83637E3EFF1367762F56FAE2" blockId="3.[151,496,1269,1331]" box="[151,303,1305,1331]" pageId="3" pageNumber="546">
(
<figureCitation id="08A02A4A83637E3EFF1B67762F5EFAE2" box="[159,295,1305,1331]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE7" captionStartId-0="4.[151,250,911,936]" captionStartId-1="4.[151,250,1802,1827]" captionStartId-2="5.[151,250,1250,1275]" captionStartId-3="5.[151,250,2014,2039]" captionStartId-4="6.[151,250,894,919]" captionStartId-5="6.[151,250,1587,1612]" captionTargetBox-0="[387,1199,181,885]" captionTargetBox-1="[201,1386,1324,1771]" captionTargetBox-2="[280,1307,561,1219]" captionTargetBox-3="[269,1318,1297,1987]" captionTargetBox-5="[295,1287,942,1552]" captionTargetId-0="figure@4.[387,1199,181,885]" captionTargetId-1="figure@4.[195,1392,1318,1777]" captionTargetId-2="figure@5.[276,1312,561,1225]" captionTargetId-3="figure@5.[269,1318,1296,1987]" captionTargetId-5="figure@6.[294,1293,941,1562]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-5="6" captionText-0="FIGURE 2. Adult habitus Haplaxius dougwalshi sp. n.; (A) body dorsal view male (B) body lateral male, scale = 1 mm." captionText-1="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." captionText-2="FIGURE 4. Forewing venation of Haplaxius dougwalshi sp. n." captionText-3="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." captionText-4="FIGURE 6. Caudal view of pygofer of adult male Haplaxius dougwalshi sp. n." captionText-5="FIGURE7. Aedeagusof adult male Haplaxiusdougwalshi sp. n.; (A) leftlateral view,(B) rightlateral view, and(C) dorsalview." figureDoi-0="http://doi.org/10.5281/zenodo.3774919" figureDoi-1="http://doi.org/10.5281/zenodo.3774921" figureDoi-2="http://doi.org/10.5281/zenodo.3774923" figureDoi-3="http://doi.org/10.5281/zenodo.3774925" figureDoi-5="http://doi.org/10.5281/zenodo.3774927" httpUri-0="https://zenodo.org/record/3774919/files/figure.png" httpUri-1="https://zenodo.org/record/3774921/files/figure.png" httpUri-2="https://zenodo.org/record/3774923/files/figure.png" httpUri-3="https://zenodo.org/record/3774925/files/figure.png" httpUri-5="https://zenodo.org/record/3774927/files/figure.png" pageId="3" pageNumber="546">Figures 2–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D881654483637E3EFF13670D2D19FAAA" box="[151,864,1377,1404]" pageId="3" pageNumber="546" type="materials_examined">
<paragraph id="902436CF83637E3EFF13670D2D19FAAA" blockId="3.[151,1437,1377,2016]" box="[151,864,1377,1404]" pageId="3" pageNumber="546">
<emphasis id="A2EFEADD83637E3EFF13670D2F4FFAAD" bold="true" box="[151,310,1377,1404]" pageId="3" pageNumber="546">
<typeStatus id="4F20886D83637E3EFF13670D2EABFAAD" box="[151,210,1378,1404]" pageId="3" pageNumber="546">Type</typeStatus>
locality.
</emphasis>
La Selva Biological Station, 
<collectingRegion id="525FF82D83637E3EFDFB670E2CA2FAAA" box="[639,731,1377,1403]" country="Costa Rica" name="Heredia" pageId="3" pageNumber="546">Heredia</collectingRegion>
, 
<collectingCountry id="E88C765F83637E3EFD62670E2D19FAAA" box="[742,864,1377,1403]" name="Costa Rica" pageId="3" pageNumber="546">Costa Rica</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="D881654483637E3EFF4367EA2E8DF9DA" pageId="3" pageNumber="546" type="diagnosis">
<paragraph id="902436CF83637E3EFF4367EA2E8DF9DA" blockId="3.[151,1437,1377,2016]" pageId="3" pageNumber="546">
<emphasis id="A2EFEADD83637E3EFF4367EA2F3BFA71" bold="true" box="[199,322,1413,1440]" pageId="3" pageNumber="546">Diagnosis.</emphasis>
This species is distinguished by a combination of bright orange coloration, facial color pattern of an orange frons with white arch traversing width of frons resulting in a trapezoidal shape dorsad of frontoclypeal suture, large, incurved processes on lateral margins of the pygofer and aedeagus with curved spine near midlength of shaft.
</paragraph>
</subSubSection>
<subSubSection id="D881654483637E3BFF43647A2A7FF945" lastPageId="6" lastPageNumber="549" pageId="3" pageNumber="546" type="description">
<paragraph id="902436CF83637E3EFF43647A2D65F96E" blockId="3.[151,1437,1377,2016]" pageId="3" pageNumber="546">
<emphasis id="A2EFEADD83637E3EFF43647A2F20F9E1" bold="true" box="[199,345,1557,1584]" pageId="3" pageNumber="546">Description.</emphasis>
<emphasis id="A2EFEADD83637E3EFEE6647A2FDAF9FE" box="[354,419,1557,1583]" italics="true" pageId="3" pageNumber="546">Color</emphasis>
. General body color is uniformly orange with abdominal sternites, middle and hind legs paler orange (
<figureCitation id="08A02A4A83637E3EFEB264562F07F982" box="[310,382,1593,1619]" captionStart="FIGURE 2" captionStartId="4.[151,250,911,936]" captionTargetBox="[387,1199,181,885]" captionTargetId="figure@4.[387,1199,181,885]" captionTargetPageId="4" captionText="FIGURE 2. Adult habitus Haplaxius dougwalshi sp. n.; (A) body dorsal view male (B) body lateral male, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774919" httpUri="https://zenodo.org/record/3774919/files/figure.png" pageId="3" pageNumber="546">Fig. 2</figureCitation>
). Frons with median white patch merging with lateral white patches, reaching frontoclypeal suture, orange patch dorsad of frontoclypeal suture trapezoidal in shape (
<figureCitation id="08A02A4A83637E3EFC4464322A60F9A6" box="[960,1049,1629,1655]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="3" pageNumber="546">Fig. 3A</figureCitation>
). Forewing veins pale orange (
<figureCitation id="08A02A4A83637E3EFAEB64322ED1F94A" captionStart="FIGURE 2" captionStartId="4.[151,250,911,936]" captionTargetBox="[387,1199,181,885]" captionTargetId="figure@4.[387,1199,181,885]" captionTargetPageId="4" captionText="FIGURE 2. Adult habitus Haplaxius dougwalshi sp. n.; (A) body dorsal view male (B) body lateral male, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774919" httpUri="https://zenodo.org/record/3774919/files/figure.png" pageId="3" pageNumber="546">Fig. 2</figureCitation>
). 
<emphasis id="A2EFEADD83637E3EFF3964ED2F5DF94D" box="[189,292,1666,1692]" italics="true" pageId="3" pageNumber="546">Structure</emphasis>
. Body length males (
<emphasis id="A2EFEADD83637E3EFD9164ED2C4FF94A" box="[533,566,1666,1691]" italics="true" pageId="3" pageNumber="546">n=</emphasis>
14): 
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with wings; 
<quantity id="57639B2A83637E3EFC1B64EE2A3BF94D" box="[927,1090,1665,1692]" metricMagnitude="-3" metricUnit="m" metricValue="2.545" metricValueMax="2.56" metricValueMin="2.53" pageId="3" pageNumber="546" unit="mm" value="2.545" valueMax="2.56" valueMin="2.53">2.53–2.56 mm</quantity>
without wings; females (
<emphasis id="A2EFEADD83637E3EFADB64ED2B14F94A" box="[1375,1389,1666,1691]" italics="true" pageId="3" pageNumber="546">n</emphasis>
=9): 
<quantity id="57639B2A83637E3EFF1364CA2F43F911" box="[151,314,1701,1728]" metricMagnitude="-3" metricUnit="m" metricValue="3.195" metricValueMax="3.2" metricValueMin="3.19" pageId="3" pageNumber="546" unit="mm" value="3.195" valueMax="3.2" valueMin="3.19">3.19–3.20 mm</quantity>
with wings; 
<quantity id="57639B2A83637E3EFE4F64CA2C17F911" box="[459,622,1701,1728]" metricMagnitude="-3" metricUnit="m" metricValue="2.57" metricValueMax="2.58" metricValueMin="2.56" pageId="3" pageNumber="546" unit="mm" value="2.57" valueMax="2.58" valueMin="2.56">2.56–2.58 mm</quantity>
without wings.
</paragraph>
<paragraph id="902436CF83637E3EFF4364A62C8FF831" blockId="3.[151,1437,1377,2016]" pageId="3" pageNumber="546">
Head. In frontal view, frons narrowed between eyes, expanding at ventral margin, transverse carina present separating frons and vertex (
<figureCitation id="08A02A4A83637E3EFE5664822C6CF8D6" box="[466,533,1773,1799]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="3" pageNumber="546">Fig. 3</figureCitation>
). Fastigium rounded in lateral view (
<figureCitation id="08A02A4A83637E3EFC2964822D88F8D6" box="[941,1009,1773,1799]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="3" pageNumber="546">Fig. 3</figureCitation>
). In dorsal view, frons concave, vertex widest posteriorly, narrowing to almost half width at base (
<figureCitation id="08A02A4A83637E3EFCBA657E2DFCF8FA" box="[830,901,1809,1835]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="3" pageNumber="546">Fig. 3</figureCitation>
). Vertex length males: 
<quantity id="57639B2A83637E3EFB0B657E2B41F8FD" box="[1167,1336,1809,1836]" metricMagnitude="-4" metricUnit="m" metricValue="3.35" metricValueMax="3.4" metricValueMin="3.3" pageId="3" pageNumber="546" unit="mm" value="0.335" valueMax="0.34" valueMin="0.33">0.33–0.34 mm</quantity>
; females 
<quantity id="57639B2A83637E3EFF13655A2E87F881" box="[151,254,1845,1872]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="546" unit="mm" value="0.35">0.35 mm</quantity>
. Vertex width at hind margin males: 
<quantity id="57639B2A83637E3EFD11655A2D43F881" box="[661,826,1845,1872]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="3" pageNumber="546" unit="mm" value="0.215" valueMax="0.22" valueMin="0.21">0.21–0.22 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFC2F655A2A29F881" box="[939,1104,1845,1872]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="3" pageNumber="546" unit="mm" value="0.225" valueMax="0.23" valueMin="0.22">0.22–0.23 mm</quantity>
. Vertex width at distal margin males: 
<quantity id="57639B2A83637E3EFF6365362FF4F8A5" box="[231,397,1881,1908]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="546" unit="mm" value="0.11499999999999999" valueMax="0.12" valueMin="0.11">0.11–0.12 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFE7A65362CDDF8A5" box="[510,676,1881,1908]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="3" pageNumber="546" unit="mm" value="0.125" valueMax="0.13" valueMin="0.12">0.12–0.13 mm</quantity>
. Frons length males: 
<quantity id="57639B2A83637E3EFC1065362A43F8A5" box="[916,1082,1881,1908]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="3" pageNumber="546" unit="mm" value="0.505" valueMax="0.51" valueMin="0.5">0.50–0.51 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFB2F65362B2BF8A5" box="[1195,1362,1881,1908]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="3" pageNumber="546" unit="mm" value="0.515" valueMax="0.52" valueMin="0.51">0.51–0.52 mm</quantity>
. Frons dorsal width males: 
<quantity id="57639B2A83637E3EFEF965122C5AF849" box="[381,547,1917,1944]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="546" unit="mm" value="0.105" valueMax="0.11" valueMin="0.1">0.10–0.11 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFD1365122D44F849" box="[663,829,1917,1944]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="546" unit="mm" value="0.11499999999999999" valueMax="0.12" valueMin="0.11">0.11–0.12 mm</quantity>
. Frons width at widest portion males: 
<quantity id="57639B2A83637E3EFB7565122BE1F849" box="[1265,1432,1917,1944]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.5999999999999996" pageId="3" pageNumber="546" unit="mm" value="0.365" valueMax="0.37" valueMin="0.36">0.36–0.37 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFF7965CE2FDBF86D" box="[253,418,1953,1980]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="3" pageNumber="546" unit="mm" value="0.375" valueMax="0.38" valueMin="0.37">0.37–0.38 mm</quantity>
. Frons frontoclypeal margin width males: 
<quantity id="57639B2A83637E3EFCFE65CE2A59F86D" box="[890,1056,1953,1980]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="3.0" metricValueMin="2.9" pageId="3" pageNumber="546" unit="mm" value="0.295" valueMax="0.3" valueMin="0.29">0.29–0.30 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFB1565CE2B4FF86D" box="[1169,1334,1953,1980]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="3" pageNumber="546" unit="mm" value="0.315" valueMax="0.32" valueMin="0.31">0.31–0.32 mm</quantity>
. Clypeus length males: 
<quantity id="57639B2A83637E3EFEB065AA2FA3F831" box="[308,474,1989,2016]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.0" metricValueMin="3.9" pageId="3" pageNumber="546" unit="mm" value="0.395" valueMax="0.4" valueMin="0.39">0.39–0.40 mm</quantity>
; females: 
<quantity id="57639B2A83637E3EFDC865AA2C8BF831" box="[588,754,1989,2016]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="3" pageNumber="546" unit="mm" value="0.405" valueMax="0.41" valueMin="0.4">0.40–0.41 mm</quantity>
.
</paragraph>
<caption id="C4E4664783647E39FF1361E02B1DFC76" ID-DOI="http://doi.org/10.5281/zenodo.3774919" ID-Zenodo-Dep="3774919" box="[151,1380,911,936]" httpUri="https://zenodo.org/record/3774919/files/figure.png" pageId="4" pageNumber="547" startId="4.[151,250,911,936]" targetBox="[387,1199,181,885]" targetPageId="4">
<paragraph id="902436CF83647E39FF1361E02B1DFC76" blockId="4.[151,1380,911,936]" box="[151,1380,911,936]" pageId="4" pageNumber="547">
<emphasis id="A2EFEADD83647E39FF1361E02F6AFC79" bold="true" box="[151,275,911,936]" pageId="4" pageNumber="547">FIGURE 2.</emphasis>
Adult 
<emphasis id="A2EFEADD83647E39FEDD61FF2CF7FC79" box="[345,654,912,936]" italics="true" pageId="4" pageNumber="547">
habitus 
<taxonomicName id="579B4D4C83647E39FE2E61FF2CF7FC79" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[426,654,912,936]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="547" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">Haplaxius dougwalshi</taxonomicName>
</emphasis>
<emphasis id="A2EFEADD83647E39FD1061FF2CB4FC79" bold="true" box="[660,717,912,936]" pageId="4" pageNumber="547">
<taxonomicNameLabel id="B9DC57A683647E39FD1061FF2CB4FC79" box="[660,717,912,936]" pageId="4" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; (A) body dorsal view male (B) body lateral male, scale = 1 mm.
</paragraph>
</caption>
<paragraph id="902436CF83647E39FF4361B92F38FAC0" blockId="4.[151,1437,982,1297]" pageId="4" pageNumber="547">
Thorax. Carinae extending to lateral margin in frontal view (
<figureCitation id="08A02A4A83647E39FC0761B92DB3FC21" box="[899,970,982,1008]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="4" pageNumber="547">Fig. 3</figureCitation>
). Mesonotum tricarinate, lateral margins subparallel (
<figureCitation id="08A02A4A83647E39FEA561952F1CFBC5" box="[289,357,1018,1044]" captionStart="FIGURE 3" captionStartId="4.[151,250,1802,1827]" captionTargetBox="[201,1386,1324,1771]" captionTargetId="figure@4.[195,1392,1318,1777]" captionTargetPageId="4" captionText="FIGURE 3. Adult Haplaxius dougwalshi sp. n.; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="4" pageNumber="547">Fig. 3</figureCitation>
). Pronotum length at midline males: 
<quantity id="57639B2A83647E39FC8761962DD0FBC4" box="[771,937,1017,1045]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="547" unit="mm" value="0.055" valueMax="0.06" valueMin="0.05">0.05–0.06 mm</quantity>
; females: 
<quantity id="57639B2A83647E39FB9E61952AC6FBC4" box="[1050,1215,1018,1045]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" metricValueMax="7" metricValueMin="6.0" pageId="4" pageNumber="547" unit="mm" value="0.065" valueMax="0.07" valueMin="0.06">0.06–0.07 mm</quantity>
. Mesonotum length at midline males: 
<quantity id="57639B2A83647E39FEDA66712C7DFBE8" box="[350,516,1054,1081]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="4" pageNumber="547" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47–0.48 mm</quantity>
; females: 
<quantity id="57639B2A83647E39FDF066712D63FBE8" box="[628,794,1054,1081]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="4.9" metricValueMin="4.8" pageId="4" pageNumber="547" unit="mm" value="0.485" valueMax="0.49" valueMin="0.48">0.48–0.49 mm</quantity>
. Mesonotum width males: 
<quantity id="57639B2A83647E39FBC366722A94FBE8" box="[1095,1261,1053,1081]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="4" pageNumber="547" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.65–0.66 mm</quantity>
; females: 
<quantity id="57639B2A83647E39FAD966712F78FB8C" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="4" pageNumber="547" unit="mm" value="0.665" valueMax="0.67" valueMin="0.66">0.66– 0.67 mm</quantity>
. Fore wings transparent, clouded posterior to pseudostigma (
<figureCitation id="08A02A4A83647E39FC44662D2A7EFB8D" box="[960,1031,1090,1116]" captionStart="FIGURE 4" captionStartId="5.[151,250,1250,1275]" captionTargetBox="[280,1307,561,1219]" captionTargetId="figure@5.[276,1312,561,1225]" captionTargetPageId="5" captionText="FIGURE 4. Forewing venation of Haplaxius dougwalshi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3774923" httpUri="https://zenodo.org/record/3774923/files/figure.png" pageId="4" pageNumber="547">Fig. 4</figureCitation>
), with conspicuous tubercles along main veins (
<figureCitation id="08A02A4A83647E39FEA766092F13FB51" box="[291,362,1126,1152]" captionStart="FIGURE 4" captionStartId="5.[151,250,1250,1275]" captionTargetBox="[280,1307,561,1219]" captionTargetId="figure@5.[276,1312,561,1225]" captionTargetPageId="5" captionText="FIGURE 4. Forewing venation of Haplaxius dougwalshi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3774923" httpUri="https://zenodo.org/record/3774923/files/figure.png" pageId="4" pageNumber="547">Fig. 4</figureCitation>
). Apex of clavus just beyond midlength of wing, PCu+A1 reaching wing margin before claval apex. Fork of CuA just basad of RA(+ScP)+RP fork, both just basad of wing midlength; MP forked from ScP+R in basal forth of wing; forewing branching pattern: RA 1-branched, RP 3, MP 5, CuA 4 (or 3 with icu crossvein); icu joining apex of clavus. PCu joining with A1 at midpoint of clavus. Forewing length males: 
<quantity id="57639B2A83647E39FB0966BE2B4AFB3C" box="[1165,1331,1233,1261]" metricMagnitude="-3" metricUnit="m" metricValue="2.59" metricValueMax="2.6" metricValueMin="2.58" pageId="4" pageNumber="547" unit="mm" value="2.59" valueMax="2.6" valueMin="2.58">2.58–2.60 mm</quantity>
; females: 
<quantity id="57639B2A83647E39FF1366992F44FAC0" box="[151,317,1270,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" metricValueMax="2.62" metricValueMin="2.6" pageId="4" pageNumber="547" unit="mm" value="2.61" valueMax="2.62" valueMin="2.6">2.60–2.62 mm</quantity>
.
</paragraph>
<caption id="C4E4664783647E39FF1365652DEAF897" ID-DOI="http://doi.org/10.5281/zenodo.3774921" ID-Zenodo-Dep="3774921" httpUri="https://zenodo.org/record/3774921/files/figure.png" pageId="4" pageNumber="547" startId="4.[151,250,1802,1827]" targetBox="[201,1386,1324,1771]" targetPageId="4">
<paragraph id="902436CF83647E39FF1365652DEAF897" blockId="4.[151,1436,1802,1863]" pageId="4" pageNumber="547">
<emphasis id="A2EFEADD83647E39FF1365652F6AF8F2" bold="true" box="[151,275,1802,1827]" pageId="4" pageNumber="547">FIGURE 3.</emphasis>
Adult 
<taxonomicName id="579B4D4C83647E39FEDE65652C47F8F3" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[346,574,1802,1826]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="547" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83647E39FEDE65652C47F8F3" box="[346,574,1802,1826]" italics="true" pageId="4" pageNumber="547">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83647E39FDC065652C07F8F3" bold="true" box="[580,638,1802,1826]" pageId="4" pageNumber="547">
<taxonomicNameLabel id="B9DC57A683647E39FDC065652C07F8F3" box="[580,638,1802,1826]" pageId="4" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; (A) male head frontal view, (B) male head, pronotum, and mesonotum dorsal view (C) male head, pronotum, and mesonotum lateral view, scale = 1 mm.
</paragraph>
</caption>
<paragraph id="902436CF83647E38FF43651A2A7BFDCA" blockId="4.[151,1436,1909,2043]" lastBlockId="5.[151,1437,152,539]" lastPageId="5" lastPageNumber="548" pageId="4" pageNumber="547">
Terminalia. Pygofer in lateral view widest ventrad, narrowing dorsally, posterior margin with large process on median posterior margin (
<figureCitation id="08A02A4A83647E39FE3B65F62C65F862" box="[447,540,1944,1971]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="4" pageNumber="547">Fig. 5A</figureCitation>
). In ventral view, pygofer opening bearing an elongate rounded lobe, approximately 2x long as wide (
<figureCitation id="08A02A4A83647E39FE2165D22F85F806" box="[421,508,1980,2007]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="4" pageNumber="547">Fig. 5B</figureCitation>
); in caudal view, pygofer with lateral processes, medially angled (
<figureCitation id="08A02A4A83647E39FB5265D22B63F809" box="[1238,1306,1981,2008]" captionStart="FIGURE 6" captionStartId="6.[151,250,894,919]" captionText="FIGURE 6. Caudal view of pygofer of adult male Haplaxius dougwalshi sp. n." pageId="4" pageNumber="547">Fig. 6</figureCitation>
). Gonostyli in lateral view angled upwards with acute apex, strongly sinuate on dorsal margin with basal curve strong, resulting in constriction of gonostyli, appearing spoon-like (
<figureCitation id="08A02A4A83657E38FD4E62F62D5AFF62" box="[714,803,152,179]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548">Fig. 5A</figureCitation>
). In ventral view, appearing clubbed with inner and outer margins of clubbed apex irregularly sinuate (
<figureCitation id="08A02A4A83657E38FD1762D22C95FF06" box="[659,748,188,215]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548">Fig. 5B</figureCitation>
), inner margin strongly sinuate, outer margin weakly sinuate (
<figureCitation id="08A02A4A83657E38FF24628E2E81FF2A" box="[160,248,224,251]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548">Fig. 5B</figureCitation>
). Aedeagus asymmetrical and complex (
<figureCitation id="08A02A4A83657E38FD39628E2C86FF2A" box="[701,767,225,251]" captionStart="FIGURE7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[295,1287,942,1552]" captionTargetId="figure@6.[294,1293,941,1562]" captionTargetPageId="6" captionText="FIGURE7. Aedeagusof adult male Haplaxiusdougwalshi sp. n.; (A) leftlateral view,(B) rightlateral view, and(C) dorsalview." figureDoi="http://doi.org/10.5281/zenodo.3774927" httpUri="https://zenodo.org/record/3774927/files/figure.png" pageId="5" pageNumber="548">Fig. 7</figureCitation>
A–C); aedeagal shaft with small spine at midlength on right lateral margin, strongly curved (almost 90° angle) (
<figureCitation id="08A02A4A83657E38FD60636A2D45FECE" box="[740,828,261,287]" captionStart="FIGURE7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[295,1287,942,1552]" captionTargetId="figure@6.[294,1293,941,1562]" captionTargetPageId="6" captionText="FIGURE7. Aedeagusof adult male Haplaxiusdougwalshi sp. n.; (A) leftlateral view,(B) rightlateral view, and(C) dorsalview." figureDoi="http://doi.org/10.5281/zenodo.3774927" httpUri="https://zenodo.org/record/3774927/files/figure.png" pageId="5" pageNumber="548">Fig. 7B</figureCitation>
) and larger spine at apex, angled dorsad and slightly distad on left lateral margin, nearly extending to base of aedeagus, approximately 2x length and thickness of small spine. Flagellum (endosoma) with one dorsal spine angled distad toward right lateral margin, terminating approximately at base of small aedeagal spine (
<figureCitation id="08A02A4A83657E38FDD4631E2CDFFE5A" box="[592,678,369,395]" captionStart="FIGURE7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[295,1287,942,1552]" captionTargetId="figure@6.[294,1293,941,1562]" captionTargetPageId="6" captionText="FIGURE7. Aedeagusof adult male Haplaxiusdougwalshi sp. n.; (A) leftlateral view,(B) rightlateral view, and(C) dorsalview." figureDoi="http://doi.org/10.5281/zenodo.3774927" httpUri="https://zenodo.org/record/3774927/files/figure.png" pageId="5" pageNumber="548">Fig. 7C</figureCitation>
) and one larger, ventral spine, arching ventrad, approximately twice length of dorsal flagellar spine, terminating approximately at same point as smaller aedeagal spine (
<figureCitation id="08A02A4A83657E38FB7E63FA2B2AFE7E" box="[1274,1363,405,431]" captionStart="FIGURE7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[295,1287,942,1552]" captionTargetId="figure@6.[294,1293,941,1562]" captionTargetPageId="6" captionText="FIGURE7. Aedeagusof adult male Haplaxiusdougwalshi sp. n.; (A) leftlateral view,(B) rightlateral view, and(C) dorsalview." figureDoi="http://doi.org/10.5281/zenodo.3774927" httpUri="https://zenodo.org/record/3774927/files/figure.png" pageId="5" pageNumber="548">Fig. 7B</figureCitation>
). Anal segment in lateral view with parallel dorsal and ventral margins, approximately 5X long at apex as wide (
<figureCitation id="08A02A4A83657E38FAB363D62BF6FE02" box="[1335,1423,440,467]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548">Fig. 5B</figureCitation>
), apex angled slightly downward, median process angled ventrad approximately 2/3 length distad of pygofer, median process of anal segment about equal size to lateral process of pygofer (
<figureCitation id="08A02A4A83657E38FC2B606E2D8DFDCA" box="[943,1012,512,539]" captionStart="FIGURE 5" captionStartId="5.[151,250,2014,2039]" captionTargetBox="[269,1318,1297,1987]" captionTargetId="figure@5.[269,1318,1296,1987]" captionTargetPageId="5" captionText="FIGURE 5. Male terminalia of Haplaxius dougwalshi sp. n.; (A) lateral view, (B) ventral view, and (C) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3774925" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548">Fig. 5</figureCitation>
).
</paragraph>
<caption id="C4E4664783657E38FF13668D2D65FB2A" ID-DOI="http://doi.org/10.5281/zenodo.3774923" ID-Zenodo-Dep="3774923" box="[151,796,1250,1275]" httpUri="https://zenodo.org/record/3774923/files/figure.png" pageId="5" pageNumber="548" startId="5.[151,250,1250,1275]" targetBox="[280,1307,561,1219]" targetPageId="5">
<paragraph id="902436CF83657E38FF13668D2D65FB2A" blockId="5.[151,796,1250,1275]" box="[151,796,1250,1275]" pageId="5" pageNumber="548">
<emphasis id="A2EFEADD83657E38FF13668D2F6AFB2A" bold="true" box="[151,275,1250,1275]" pageId="5" pageNumber="548">FIGURE 4.</emphasis>
Forewing venation of 
<taxonomicName id="579B4D4C83657E38FE7E668D2CA4FB2B" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[506,733,1250,1274]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="548" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83657E38FE7E668D2CA4FB2B" box="[506,733,1250,1274]" italics="true" pageId="5" pageNumber="548">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9DC57A683657E38FD67668C2D65FB2A" box="[739,796,1251,1275]" pageId="5" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="C4E4664783657E38FF1365B12B4EF826" ID-DOI="http://doi.org/10.5281/zenodo.3774925" ID-Zenodo-Dep="3774925" box="[151,1335,2014,2039]" httpUri="https://zenodo.org/record/3774925/files/figure.png" pageId="5" pageNumber="548" startId="5.[151,250,2014,2039]" targetBox="[269,1318,1297,1987]" targetPageId="5">
<paragraph id="902436CF83657E38FF1365B12B4EF826" blockId="5.[151,1335,2014,2039]" box="[151,1335,2014,2039]" pageId="5" pageNumber="548">
<emphasis id="A2EFEADD83657E38FF1365B12F6AF826" bold="true" box="[151,275,2014,2039]" pageId="5" pageNumber="548">FIGURE 5.</emphasis>
Male terminalia of 
<taxonomicName id="579B4D4C83657E38FE5865B12CB9F827" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[476,704,2014,2038]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="548" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83657E38FE5865B12CB9F827" box="[476,704,2014,2038]" italics="true" pageId="5" pageNumber="548">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83657E38FD4265B02C86F826" bold="true" box="[710,767,2015,2039]" pageId="5" pageNumber="548">
<taxonomicNameLabel id="B9DC57A683657E38FD4265B02C86F826" box="[710,767,2015,2039]" pageId="5" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; (A) lateral view, (B) ventral view, and (C) dorsal view.
</paragraph>
</caption>
<caption id="C4E4664783667E3BFF1361112DC7FC46" box="[151,958,894,919]" pageId="6" pageNumber="549">
<paragraph id="902436CF83667E3BFF1361112DC7FC46" blockId="6.[151,958,894,919]" box="[151,958,894,919]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF1361112F6AFC46" bold="true" box="[151,275,894,919]" pageId="6" pageNumber="549">FIGURE 6.</emphasis>
Caudal view of pygofer of adult male 
<taxonomicName id="579B4D4C83667E3BFD1861112D06FC47" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[668,895,894,918]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83667E3BFD1861112D06FC47" box="[668,895,894,918]" italics="true" pageId="6" pageNumber="549">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9DC57A683667E3BFC0161102DC7FC46" box="[901,958,895,919]" pageId="6" pageNumber="549" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="C4E4664783667E3BFF13645C2BE4F99D" ID-DOI="http://doi.org/10.5281/zenodo.3774927" ID-Zenodo-Dep="3774927" box="[151,1437,1587,1612]" httpUri="https://zenodo.org/record/3774927/files/figure.png" pageId="6" pageNumber="549" startId="6.[151,250,1587,1612]" targetBox="[295,1287,942,1552]" targetPageId="6">
<paragraph id="902436CF83667E3BFF13645C2BE4F99D" blockId="6.[151,1437,1587,1612]" box="[151,1437,1587,1612]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF13645C2F69F99D" bold="true" box="[151,272,1587,1612]" pageId="6" pageNumber="549">FIGURE 7.</emphasis>
Aedeagus of adult male 
<taxonomicName id="579B4D4C83667E3BFD97645B2C8AF99D" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[531,755,1588,1612]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83667E3BFD97645B2C8AF99D" box="[531,755,1588,1612]" italics="true" pageId="6" pageNumber="549">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83667E3BFD73645B2D54F99D" bold="true" box="[759,813,1588,1612]" pageId="6" pageNumber="549">
<taxonomicNameLabel id="B9DC57A683667E3BFD73645B2D54F99D" box="[759,813,1588,1612]" pageId="6" pageNumber="549" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; (A) left lateral view, (B) right lateral view, and (C) dorsal view.
</paragraph>
</caption>
<paragraph id="902436CF83667E3BFF4364152A7FF945" blockId="6.[151,1436,1657,2045]" box="[199,1030,1657,1685]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF4364152FDAF945" bold="true" box="[199,419,1657,1684]" pageId="6" pageNumber="549">Plant associations.</emphasis>
Pata de gallo palm (
<taxonomicName id="579B4D4C83667E3BFD0D64142D0DF945" box="[649,884,1659,1685]" class="Liliopsida" family="Arecaceae" genus="Asterogyne" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="549" phylum="Tracheophyta" rank="species" species="martiana">
<emphasis id="A2EFEADD83667E3BFD0D64142D0DF945" box="[649,884,1659,1685]" italics="true" pageId="6" pageNumber="549">Asterogyne martiana</emphasis>
</taxonomicName>
), 
<taxonomicName id="579B4D4C83667E3BFC0E64152A7BF945" box="[906,1026,1658,1684]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="549" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D881654483667E3BFF4364F22DF8F968" box="[199,897,1693,1721]" pageId="6" pageNumber="549" type="distribution">
<paragraph id="902436CF83667E3BFF4364F22DF8F968" blockId="6.[151,1436,1657,2045]" box="[199,897,1693,1721]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF4364F22F18F969" bold="true" box="[199,353,1693,1720]" pageId="6" pageNumber="549">Distribution.</emphasis>
<collectingCountry id="E88C765F83667E3BFEE364F12F98F969" box="[359,481,1694,1720]" name="Costa Rica" pageId="6" pageNumber="549">Costa Rica</collectingCountry>
(
<collectingRegion id="525FF82D83667E3BFE7464F12C34F969" box="[496,589,1694,1720]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="549">Heredia</collectingRegion>
, Puerto Viejo de Sarapiqui)
</paragraph>
</subSubSection>
<subSubSection id="D881654483667E3BFF4364AD2D58F8F4" pageId="6" pageNumber="549" type="etymology">
<paragraph id="902436CF83667E3BFF4364AD2D58F8F4" blockId="6.[151,1436,1657,2045]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF4364AD2F37F90D" bold="true" box="[199,334,1730,1756]" pageId="6" pageNumber="549">Etymology.</emphasis>
The specific name given is an honorarium in reference to the lead authors professor and chair during graduate school (Ph.D.), Dr. Doug Walsh at 
<collectingRegion id="525FF82D83667E3BFD3064892D01F8D1" box="[692,888,1766,1792]" country="United States of America" name="Washington" pageId="6" pageNumber="549">Washington State</collectingRegion>
University, who’s signature mustache resembles the color pattern observed on the frons of the novel taxon.
</paragraph>
</subSubSection>
<subSubSection id="D881654483667E3AFF4365422CDDFB17" lastPageId="7" lastPageNumber="550" pageId="6" pageNumber="549" type="materials_examined">
<paragraph id="902436CF83667E3BFF4365422A44F841" blockId="6.[151,1436,1657,2045]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF4365422FCBF899" bold="true" box="[199,434,1837,1864]" pageId="6" pageNumber="549">Material examined.</emphasis>
<materialsCitation id="20F33C9283667E3BFE3D65412AD6F8BD" ID-GBIF-Occurrence="2609409778" collectingDate="2019-05-26" collectionCode="FLREC" collectorName="Coll. &amp; B. W. Bahder" country="Costa Rica" location="La Selva Biological Station" pageId="6" pageNumber="549" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="4F20886D83667E3BFE3D65412C58F899" box="[441,545,1838,1864]" pageId="6" pageNumber="549" type="holotype">Holotype</typeStatus>
male “ 
<collectingCountry id="E88C765F83667E3BFDF665412C8BF899" box="[626,754,1838,1864]" name="Costa Rica" pageId="6" pageNumber="549">Costa Rica</collectingCountry>
, 
<collectingRegion id="525FF82D83667E3BFD7A65412D2EF899" box="[766,855,1838,1864]" country="Costa Rica" name="Heredia" pageId="6" pageNumber="549">Heredia</collectingRegion>
/ 
<location id="9544601483667E3BFCEB65402ADFF899" LSID="urn:lsid:plazi:treatment:183287D983637E35FF13669A2AD6FE92:9544601483667E3BFCEB65402ADFF899" box="[879,1190,1838,1865]" country="Costa Rica" name="La Selva Biological Station" pageId="6" pageNumber="549" stateProvince="Heredia">La Selva Biological Station</location>
/ 
<date id="E425100F83667E3BFB3A65412B48F899" box="[1214,1329,1838,1864]" pageId="6" pageNumber="549" value="2019-05-26">
<collectingDate id="F461E9E783667E3BFB3A65412B48F899" box="[1214,1329,1838,1864]" pageId="6" pageNumber="549" value="2019-05-26">26.V.2019</collectingDate>
</date>
/ Sweeping 
<taxonomicName id="579B4D4C83667E3BFF40653C2FDDF8BD" box="[196,420,1875,1901]" class="Magnoliopsida" family="Celastraceae" genus="Astrogyne" higherTaxonomySource="CoL" kingdom="Plantae" order="Celastrales" pageId="6" pageNumber="549" phylum="Tracheophyta" rank="species" species="martiana">
<emphasis id="A2EFEADD83667E3BFF40653C2FDDF8BD" box="[196,420,1875,1901]" italics="true" pageId="6" pageNumber="549">Astrogyne martiana</emphasis>
</taxonomicName>
/ 
<collectorName id="3D6E531983667E3BFE38653D2F81F8BD" box="[444,504,1874,1900]" pageId="6" pageNumber="549">Coll.</collectorName>
: 
<collectorName id="3D6E531983667E3BFD80653D2CF4F8BD" box="[516,653,1874,1900]" pageId="6" pageNumber="549">B.W.Bahder</collectorName>
// 
<typeStatus id="4F20886D83667E3BFD2A653D2D6FF8BD" box="[686,790,1874,1900]" pageId="6" pageNumber="549" type="holotype">Holotype</typeStatus>
/ 
<taxonomicName id="579B4D4C83667E3BFCAB653D2A55F8BD" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[815,1068,1874,1900]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="dougwalshi">
<emphasis id="A2EFEADD83667E3BFCAB653D2A55F8BD" box="[815,1068,1874,1900]" italics="true" pageId="6" pageNumber="549">Haplaxius dougwalshi</emphasis>
</taxonomicName>
” (
<collectionCode id="F68AAE0A83667E3BFBCD653D2ADEF8BD" box="[1097,1191,1874,1900]" pageId="6" pageNumber="549">FLREC</collectionCode>
)
</materialsCitation>
; 
<typeStatus id="4F20886D83667E3BFB3F653C2B52F8BC" box="[1211,1323,1875,1901]" pageId="6" pageNumber="549" type="paratype">Paratypes</typeStatus>
, La Selva Biological Research Station [
<date id="E425100F83667E3BFE5A651A2C2CF841" box="[478,597,1909,1936]" pageId="6" pageNumber="549" value="2019-05-26">
26. 
<collectionCode id="F68AAE0A83667E3BFE7B651A2C6CF841" box="[511,533,1909,1936]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="549" type="Museum">V</collectionCode>
.2019
</date>
] (
<specimenCount id="869DFD4683667E3BFDE865192CAAF841" box="[620,723,1910,1936]" pageId="6" pageNumber="549" type="male">13 males</specimenCount>
, 
<specimenCount id="869DFD4683667E3BFD5B65192D37F841" box="[735,846,1910,1936]" pageId="6" pageNumber="549" type="female">9 females</specimenCount>
, 
<collectionCode id="F68AAE0A83667E3BFCDD65192DC9F841" box="[857,944,1910,1936]" pageId="6" pageNumber="549">FLREC</collectionCode>
and 
<collectionCode id="F68AAE0A83667E3BFC6265192A48F841" box="[998,1073,1910,1936]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="6" pageNumber="549" type="Museum">FSCA</collectionCode>
).
</paragraph>
<paragraph id="902436CF83667E3BFF4365F62C24F865" blockId="6.[151,1436,1657,2045]" box="[199,605,1945,1972]" pageId="6" pageNumber="549">
<emphasis id="A2EFEADD83667E3BFF4365F62C24F865" bold="true" box="[199,605,1945,1972]" pageId="6" pageNumber="549">
Other 
<taxonomicName id="579B4D4C83667E3BFE9165F62FF4F865" authorityName="Fowler" authorityYear="1904" box="[277,397,1945,1972]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="A2EFEADD83667E3BFE9165F62FF4F865" bold="true" box="[277,397,1945,1972]" italics="true" pageId="6" pageNumber="549">Haplaxius</emphasis>
</taxonomicName>
species collected.
</emphasis>
</paragraph>
<paragraph id="902436CF83667E3BFF4365D12C7FF82D" blockId="6.[151,1436,1657,2045]" pageId="6" pageNumber="549">
<taxonomicName id="579B4D4C83667E3BFF4365D12FCAF809" box="[199,435,1982,2008]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="skarphion">
<emphasis id="A2EFEADD83667E3BFF4365D12FCAF809" box="[199,435,1982,2008]" italics="true" pageId="6" pageNumber="549">Haplaxius skarphion</emphasis>
</taxonomicName>
– “ 
<collectingCountry id="E88C765F83667E3BFE5F65D12C25F809" box="[475,604,1982,2008]" name="Costa Rica" pageId="6" pageNumber="549">Costa Rica</collectingCountry>
, 
<collectingRegion id="525FF82D83667E3BFDEC65D12CCAF809" box="[616,691,1982,2008]" country="Costa Rica" name="Limon" pageId="6" pageNumber="549">Limón</collectingRegion>
/ Gandoca, nr. 
<collectingCountry id="E88C765F83667E3BFCE765D02DC5F808" box="[867,956,1983,2009]" name="Panama" pageId="6" pageNumber="549">Panama</collectingCountry>
/ B.W.Bahder; 
<date id="E425100F83667E3BFBEA65D12A99F809" box="[1134,1248,1982,2008]" pageId="6" pageNumber="549" value="2018-05-14">
14. 
<collectionCode id="F68AAE0A83667E3BFB0865D12AD8F809" box="[1164,1185,1982,2008]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="549" type="Museum">V</collectionCode>
.2018
</date>
/ aspirated from coconut palm” (
<specimenCount id="869DFD4683667E3BFECF658D2FDDF82C" box="[331,420,2018,2045]" pageId="6" pageNumber="549" type="male">2 males</specimenCount>
) (
<figureCitation id="08A02A4A83667E3BFE3F658D2F84F82D" box="[443,509,2018,2044]" captionStart="FIGURE 8" captionStartId="7.[151,250,693,718]" captionTargetBox="[205,1368,182,667]" captionTargetId="figure@7.[201,1386,181,667]" captionTargetPageId="7" captionText="FIGURE 8. Adult male of Haplaxius skarphion; (A) body lateral view, (B) frontal view of head and prothorax, (C) ventral view of terminalia, (D) lateral view of terminalia, (E) left lateral view of aedeagus, and (F) right lateral view of aedeagus, scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774929" httpUri="https://zenodo.org/record/3774929/files/figure.png" pageId="6" pageNumber="549">Fig. 8</figureCitation>
)
</paragraph>
<caption id="C4E4664783677E3AFF1360DA2E94FCC4" ID-DOI="http://doi.org/10.5281/zenodo.3774929" ID-Zenodo-Dep="3774929" httpUri="https://zenodo.org/record/3774929/files/figure.png" pageId="7" pageNumber="550" startId="7.[151,250,693,718]" targetBox="[205,1368,182,667]" targetPageId="7">
<paragraph id="902436CF83677E3AFF1360DA2E94FCC4" blockId="7.[151,1436,693,790]" pageId="7" pageNumber="550">
<emphasis id="A2EFEADD83677E3AFF1360DA2F6BFD1F" bold="true" box="[151,274,693,718]" pageId="7" pageNumber="550">FIGURE 8.</emphasis>
Adult male of 
<taxonomicName id="579B4D4C83677E3AFE2C60D92C02FD1F" box="[424,635,694,718]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="skarphion">
<emphasis id="A2EFEADD83677E3AFE2C60D92C02FD1F" box="[424,635,694,718]" italics="true" pageId="7" pageNumber="550">Haplaxius skarphion</emphasis>
</taxonomicName>
; (A) body lateral view, (B) frontal view of head and prothorax, (C) ventral view of terminalia, (D) lateral view of terminalia, (E) left lateral view of aedeagus, and (F) right lateral view of aedeagus, scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="902436CF83677E3AFF43612C2CDDFB17" blockId="7.[151,1436,835,1222]" pageId="7" pageNumber="550">
<emphasis id="A2EFEADD83677E3AFF43612C2F00FC8F" bold="true" box="[199,377,835,862]" pageId="7" pageNumber="550">Sequence data.</emphasis>
For the COI locus, 683 bp of the 5-prime region of the gene were obtained for 
<taxonomicName id="579B4D4C83677E3AFB66612B2E99FC53" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFB66612B2E99FC53" italics="true" pageId="7" pageNumber="550">Haplaxius dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFF6C61072F50FC53" bold="true" box="[232,297,872,898]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFF6C61072F50FC53" box="[232,297,872,898]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(GenBank Accession No. 
<accessionNumber id="8FC8AB2C83677E3AFDD361072CAEFC53" box="[599,727,872,898]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT080284" pageId="7" pageNumber="550">
<collectionCode id="F68AAE0A83677E3AFDD361072CF8FC53" box="[599,641,872,898]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="7" pageNumber="550">MT</collectionCode>
080284
</accessionNumber>
) and was compared to the corresponding region for 
<taxonomicName id="579B4D4C83677E3AFAAD61072E9AFC77" baseAuthorityName="Van Duzee" baseAuthorityYear="1907" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="crudus">
<emphasis id="A2EFEADD83677E3AFAAD61072E9AFC77" italics="true" pageId="7" pageNumber="550">Haplaxius crudus</emphasis>
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFF7561E32C72FC77" box="[241,523,908,934]" class="Insecta" family="Cixiidae" genus="Nymphomyndus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="carribea">
<emphasis id="A2EFEADD83677E3AFF7561E32C72FC77" box="[241,523,908,934]" italics="true" pageId="7" pageNumber="550">Nymphomyndus carribea</emphasis>
</taxonomicName>
and 
<taxonomicName id="579B4D4C83677E3AFDC661E32D42FC77" box="[578,827,908,934]" class="Insecta" family="Cixiidae" genus="Oecleus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="mackaspringi">
<emphasis id="A2EFEADD83677E3AFDC661E32D42FC77" box="[578,827,908,934]" italics="true" pageId="7" pageNumber="550">Oecleus mackaspringi</emphasis>
</taxonomicName>
. There was strong bootstrap support (88) for 
<taxonomicName id="579B4D4C83677E3AFABD61E22E99FC1B" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFABD61E22E99FC1B" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFF6361DF2F5EFC1B" bold="true" box="[231,295,944,970]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFF6361DF2F5EFC1B" box="[231,295,944,970]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resolving next to 
<taxonomicName id="579B4D4C83677E3AFE7461DE2C27FC1B" baseAuthorityName="Van Duzee" baseAuthorityYear="1907" box="[496,606,944,970]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="crudus">
<emphasis id="A2EFEADD83677E3AFE7461DE2C27FC1B" box="[496,606,944,970]" italics="true" pageId="7" pageNumber="550">H. crudus</emphasis>
</taxonomicName>
relative to the other two genera of oecleines included (
<figureCitation id="08A02A4A83677E3AFB4261DF2B67FC1B" box="[1222,1310,944,970]" captionStart="FIGURE 9" captionStartId="7.[151,250,1657,1682]" captionTargetBox="[227,1358,1249,1629]" captionTargetId="figure@7.[227,1361,1244,1633]" captionTargetPageId="7" captionText="FIGURE 9. Maximum Likelihood phylogenetic tree based on COI sequence data (A) and 18S sequence data (B). Branch support was assessed using 1,000 bootstrap replicates." figureDoi="http://doi.org/10.5281/zenodo.3774931" httpUri="https://zenodo.org/record/3774931/files/figure.png" pageId="7" pageNumber="550">Fig. 9A</figureCitation>
) and based on the pairwise comparison, was 12% different than 
<taxonomicName id="579B4D4C83677E3AFD5A61BA2D32FC3F" baseAuthorityName="Van Duzee" baseAuthorityYear="1907" box="[734,843,980,1006]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="crudus">
<emphasis id="A2EFEADD83677E3AFD5A61BA2D32FC3F" box="[734,843,980,1006]" italics="true" pageId="7" pageNumber="550">H. crudus</emphasis>
</taxonomicName>
, 16.4% different than 
<taxonomicName id="579B4D4C83677E3AFBC061BA2AB1FC3F" baseAuthorityName="Fennah" baseAuthorityYear="1971" box="[1092,1224,980,1006]" class="Insecta" family="Cixiidae" genus="Nymphomyndus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="caribbea">
<emphasis id="A2EFEADD83677E3AFBC061BA2AB1FC3F" box="[1092,1224,980,1006]" italics="true" pageId="7" pageNumber="550">N. caribbea</emphasis>
</taxonomicName>
, and 17% different from 
<taxonomicName id="579B4D4C83677E3AFF5061972FF7FBC3" box="[212,398,1016,1042]" class="Insecta" family="Cixiidae" genus="Oecleus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="mackaspringi">
<emphasis id="A2EFEADD83677E3AFF5061972FF7FBC3" box="[212,398,1016,1042]" italics="true" pageId="7" pageNumber="550">O. mackaspringi</emphasis>
</taxonomicName>
(
<tableCitation id="DD19037483677E3AFE1861972F89FBC3" box="[412,496,1016,1042]" captionStart="TABLE 1" captionStartId="8.[151,239,370,394]" captionText="TABLE 1. Pairwise comparison for the COI gene based on 1,000 bootstrap replications using the p-distance method; numbers on bottom left=percent difference, numbers in upper right=standard error." pageId="7" pageNumber="550">Table 1</tableCitation>
). For the 18S locus, 1,358 bp were obtained for 
<taxonomicName id="579B4D4C83677E3AFB8861962AD7FBC3" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[1036,1198,1016,1042]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFB8861962AD7FBC3" box="[1036,1198,1016,1042]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFB3161972A8DFBC3" bold="true" box="[1205,1268,1016,1042]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFB3161972A8DFBC3" box="[1205,1268,1016,1042]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(GenBank Accession No. 
<accessionNumber id="8FC8AB2C83677E3AFEA466742FD9FBE7" box="[288,416,1051,1078]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT002395" pageId="7" pageNumber="550">
<collectionCode id="F68AAE0A83677E3AFEA466742F32FBE7" box="[288,331,1051,1078]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="7" pageNumber="550">MT</collectionCode>
002395
</accessionNumber>
) and was compared to the corresponding region for 
<taxonomicName id="579B4D4C83677E3AFC6366722A2CFBE7" baseAuthorityName="Van Duzee" baseAuthorityYear="1907" box="[999,1109,1052,1078]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="crudus">
<emphasis id="A2EFEADD83677E3AFC6366722A2CFBE7" box="[999,1109,1052,1078]" italics="true" pageId="7" pageNumber="550">H. crudus</emphasis>
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFBE766722A9EFBE7" baseAuthorityName="Fennah" baseAuthorityYear="1971" box="[1123,1255,1052,1078]" class="Insecta" family="Cixiidae" genus="Nymphomyndus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="caribbea">
<emphasis id="A2EFEADD83677E3AFBE766722A9EFBE7" box="[1123,1255,1052,1078]" italics="true" pageId="7" pageNumber="550">N. caribbea</emphasis>
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFB7166722BECFBE7" box="[1269,1429,1052,1078]" class="Bivalvia" family="Veneridae" genus="Myxia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="7" pageNumber="550" phylum="Mollusca" rank="species" species="belinda">
<emphasis id="A2EFEADD83677E3AFB7166722BECFBE7" box="[1269,1429,1052,1078]" italics="true" pageId="7" pageNumber="550">Myxia belinda</emphasis>
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFF13662F2F43FB8B" box="[151,314,1088,1114]" class="Insecta" family="Cixiidae" genus="Myndus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="taffini">
<emphasis id="A2EFEADD83677E3AFF13662F2F43FB8B" box="[151,314,1088,1114]" italics="true" pageId="7" pageNumber="550">Myndus taffini</emphasis>
</taxonomicName>
, and 
<taxonomicName id="579B4D4C83677E3AFEFC662F2C4BFB8B" box="[376,562,1088,1114]" class="Insecta" family="Cixiidae" genus="Oecleus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="mackaspringi">
<emphasis id="A2EFEADD83677E3AFEFC662F2C4BFB8B" box="[376,562,1088,1114]" italics="true" pageId="7" pageNumber="550">O. mackaspringi</emphasis>
</taxonomicName>
. There was strong bootstrap support for 
<taxonomicName id="579B4D4C83677E3AFC7F662E2AE7FB8B" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[1019,1182,1088,1114]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi">
<emphasis id="A2EFEADD83677E3AFC7F662E2AE7FB8B" box="[1019,1182,1088,1114]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
resolving in the genus 
<taxonomicName id="579B4D4C83677E3AFF13660B2F73FBAF" authorityName="Fowler" authorityYear="1904" box="[151,266,1124,1150]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="A2EFEADD83677E3AFF13660B2F73FBAF" box="[151,266,1124,1150]" italics="true" pageId="7" pageNumber="550">Haplaxius</emphasis>
</taxonomicName>
(98) relative to other genera of oecleines (
<figureCitation id="08A02A4A83677E3AFD5A660B2D4CFBAF" box="[734,821,1124,1150]" captionStart="FIGURE 9" captionStartId="7.[151,250,1657,1682]" captionTargetBox="[227,1358,1249,1629]" captionTargetId="figure@7.[227,1361,1244,1633]" captionTargetPageId="7" captionText="FIGURE 9. Maximum Likelihood phylogenetic tree based on COI sequence data (A) and 18S sequence data (B). Branch support was assessed using 1,000 bootstrap replicates." figureDoi="http://doi.org/10.5281/zenodo.3774931" httpUri="https://zenodo.org/record/3774931/files/figure.png" pageId="7" pageNumber="550">Fig. 9B</figureCitation>
). Additionally, 
<taxonomicName id="579B4D4C83677E3AFC5B660A2AF8FBAF" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[991,1153,1124,1150]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFC5B660A2AF8FBAF" box="[991,1153,1124,1150]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFB03660B2ABCFBAF" bold="true" box="[1159,1221,1124,1150]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFB03660B2ABCFBAF" box="[1159,1221,1124,1150]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
was 0.6% different from 
<taxonomicName id="579B4D4C83677E3AFF5066E62F38FB73" baseAuthorityName="Van Duzee" baseAuthorityYear="1907" box="[212,321,1160,1186]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="crudus">
<emphasis id="A2EFEADD83677E3AFF5066E62F38FB73" box="[212,321,1160,1186]" italics="true" pageId="7" pageNumber="550">H. crudus</emphasis>
</taxonomicName>
for the 18S gene whereas it was on average, 3.0% different from the other oecleine genera included, ranging from 1.8% to 5.1% different (
<tableCitation id="DD19037483677E3AFDBB66C32CEEFB17" box="[575,663,1196,1222]" captionStart="TABLE 2" captionStartId="8.[151,239,681,705]" captionText="TABLE 2. Pairwise comparison for the 18S gene based on 1,000 bootstrap replications using the p-distance method; numbers on bottom left=percent difference, numbers in upper right=standard error." pageId="7" pageNumber="550">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="C4E4664783677E3AFF1364162CE3F967" ID-DOI="http://doi.org/10.5281/zenodo.3774931" ID-Zenodo-Dep="3774931" httpUri="https://zenodo.org/record/3774931/files/figure.png" pageId="7" pageNumber="550" startId="7.[151,250,1657,1682]" targetBox="[227,1358,1249,1629]" targetPageId="7">
<paragraph id="902436CF83677E3AFF1364162CE3F967" blockId="7.[151,1437,1657,1718]" pageId="7" pageNumber="550">
<emphasis id="A2EFEADD83677E3AFF1364162F6AF943" bold="true" box="[151,275,1657,1682]" pageId="7" pageNumber="550">FIGURE 9.</emphasis>
Maximum Likelihood phylogenetic tree based on COI sequence data (A) and 18S sequence data (B). Branch support was assessed using 1,000 bootstrap replicates.
</paragraph>
</caption>
<subSubSection id="D881654483677E35FF43648A2AD6FE92" lastPageId="8" lastPageNumber="551" pageId="7" pageNumber="550" type="discussion">
<paragraph id="902436CF83677E35FF43648A2CD9FF06" blockId="7.[151,1437,1764,2043]" lastBlockId="8.[151,1437,153,323]" lastPageId="8" lastPageNumber="551" pageId="7" pageNumber="550">
<emphasis id="A2EFEADD83677E3AFF43648A2F42F92E" bold="true" box="[199,315,1765,1791]" pageId="7" pageNumber="550">Remarks.</emphasis>
The general form of the aedeagus of 
<taxonomicName id="579B4D4C83677E3AFD63648A2DF2F92F" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[743,907,1764,1790]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFD63648A2DF2F92F" box="[743,907,1764,1790]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFC17648A2DAAF92E" bold="true" box="[915,979,1765,1791]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFC17648A2DAAF92E" box="[915,979,1765,1791]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to many species illustrated by 
<bibRefCitation id="F40A4B3E83677E3AFF1365662F38F8F2" author="Kramer, J. P." box="[151,321,1801,1827]" pageId="7" pageNumber="550" pagination="301 - 389" refId="ref5846" refString="Kramer, J. P. (1979) Taxonomic study of the planthopper genus Myndus in the Americas (Homoptera: Fulgoroidea: Cixiidae). Transactions of the American Entomological Society, 105 (3), 301 - 389." type="journal article" year="1979">Kramer (1979)</bibRefCitation>
. While the presence of a spine on the aedeagus before the apex is not unique to 
<taxonomicName id="579B4D4C83677E3AFB3165662B2EF8F3" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[1205,1367,1800,1826]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFB3165662B2EF8F3" box="[1205,1367,1800,1826]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFADA65662BE4F8F2" bold="true" box="[1374,1437,1801,1827]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFADA65662BE4F8F2" box="[1374,1437,1801,1827]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(shared with 
<taxonomicName id="579B4D4C83677E3AFEAF65422C7BF896" authority="(Osborn)" baseAuthorityName="Osborn" baseAuthorityYear="1903" box="[299,514,1836,1863]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="fulvus">
<emphasis id="A2EFEADD83677E3AFEAF65422FEBF897" box="[299,402,1836,1862]" italics="true" pageId="7" pageNumber="550">H. fulvus</emphasis>
(Osborn)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFD8A65422D58F896" authority="(Van Duzee)" baseAuthorityName="Van Duzee" baseAuthorityYear="1909" box="[526,801,1836,1863]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="A2EFEADD83677E3AFD8A65422CF1F897" box="[526,648,1836,1862]" italics="true" pageId="7" pageNumber="550">H. pusillus</emphasis>
(Van Duzee)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFCA965422D98F896" authority="(Ball)" baseAuthorityName="Ball" baseAuthorityYear="1902" box="[813,993,1836,1863]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="A2EFEADD83677E3AFCA965422DE1F897" box="[813,920,1836,1862]" italics="true" pageId="7" pageNumber="550">H. viridis</emphasis>
(Ball)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFC7465422A92F896" authority="(Kramer)" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[1008,1259,1837,1863]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="jamaicae">
<emphasis id="A2EFEADD83677E3AFC7465422A02F897" box="[1008,1147,1837,1862]" italics="true" pageId="7" pageNumber="550">H. jamaicae</emphasis>
(Kramer)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFB7D65422E85F8BB" authority="Caldwell" authorityName="Caldwell" authorityYear="1946" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="simplicatus">
<emphasis id="A2EFEADD83677E3AFB7D65422BE4F897" box="[1273,1437,1836,1862]" italics="true" pageId="7" pageNumber="550">H. simplicatus</emphasis>
Caldwell
</taxonomicName>
, and 
<taxonomicName id="579B4D4C83677E3AFEB8653E2C42F8BA" authority="(Kramer))" authorityName="" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[316,571,1872,1899]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="vilbastei">
<emphasis id="A2EFEADD83677E3AFEB8653E2FC7F8BB" box="[316,446,1872,1898]" italics="true" pageId="7" pageNumber="550">H. vilbastei</emphasis>
(Kramer))
</taxonomicName>
, the spine on the shaft in 
<taxonomicName id="579B4D4C83677E3AFCE4653E2A7DF8BB" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[864,1028,1872,1898]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi">
<emphasis id="A2EFEADD83677E3AFCE4653E2A7DF8BB" box="[864,1028,1872,1898]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFB88653E2A3FF8BA" bold="true" box="[1036,1094,1873,1899]" pageId="7" pageNumber="550">sp n.</emphasis>
is distinct in being positioned further basad than the other species and strongly curved, whereas the other taxa possess a spine that is either straight or only slightly curved. The lateral processes of the pygofer of 
<taxonomicName id="579B4D4C83677E3AFCDD65F62D85F863" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[857,1020,1944,1970]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83677E3AFCDD65F62D85F863" box="[857,1020,1944,1970]" italics="true" pageId="7" pageNumber="550">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83677E3AFB8765F62A3AF862" bold="true" box="[1027,1091,1945,1971]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="B9DC57A683677E3AFB8765F62A3AF862" box="[1027,1091,1945,1971]" pageId="7" pageNumber="550" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is unique and easily diagnoses it, being much longer and narrower than any other species of 
<taxonomicName id="579B4D4C83677E3AFCDF65D32DB7F807" authorityName="Fowler" authorityYear="1904" box="[859,974,1980,2006]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="genus">
<emphasis id="A2EFEADD83677E3AFCDF65D32DB7F807" box="[859,974,1980,2006]" italics="true" pageId="7" pageNumber="550">Haplaxius</emphasis>
</taxonomicName>
. Other taxa that also a possess a similar process include 
<taxonomicName id="579B4D4C83677E3AFECF658E2C48F82A" authority="(Kramer)" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[331,561,2016,2043]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="glyphis">
<emphasis id="A2EFEADD83677E3AFECF658E2FC6F82B" box="[331,447,2016,2042]" italics="true" pageId="7" pageNumber="550">H. glyphis</emphasis>
(Kramer)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFDBF658E2D56F82A" authority="(Kramer)" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[571,815,2016,2043]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="gnophos">
<emphasis id="A2EFEADD83677E3AFDBF658E2CC4F82B" box="[571,701,2016,2042]" italics="true" pageId="7" pageNumber="550">H. gnophos</emphasis>
(Kramer)
</taxonomicName>
, 
<taxonomicName id="579B4D4C83677E3AFCBE658E2A34F82A" authority="(Kramer)" baseAuthorityName="Kramer" baseAuthorityYear="1979" box="[826,1101,2016,2043]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="neopusillus">
<emphasis id="A2EFEADD83677E3AFCBE658E2DA5F82B" box="[826,988,2016,2042]" italics="true" pageId="7" pageNumber="550">H. neopusillus</emphasis>
(Kramer)
</taxonomicName>
, and 
<taxonomicName id="579B4D4C83677E3AFB0C658E2BE1F82A" authority="(Van Duzee)" baseAuthorityName="Van Duzee" baseAuthorityYear="1909" box="[1160,1432,2016,2043]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="550" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="A2EFEADD83677E3AFB0C658E2B78F82B" box="[1160,1281,2016,2042]" italics="true" pageId="7" pageNumber="550">H. pusillus</emphasis>
(Van Duzee)
</taxonomicName>
; however, in these taxa the base of the process is broad whereas in 
<taxonomicName id="579B4D4C83687E35FC0A62F52A4AFF62" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[910,1075,153,179]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="551" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83687E35FC0A62F52A4AFF62" box="[910,1075,153,179]" italics="true" pageId="8" pageNumber="551">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83687E35FBB862F62A04FF62" bold="true" box="[1084,1149,153,179]" pageId="8" pageNumber="551">
<taxonomicNameLabel id="B9DC57A683687E35FBB862F62A04FF62" box="[1084,1149,153,179]" pageId="8" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the base is strongly narrowed, producing a very well-defined process.
</paragraph>
<paragraph id="902436CF83687E35FF43628E2AD6FE92" blockId="8.[151,1437,153,323]" pageId="8" pageNumber="551">
While the features of the genitalia are the most reliable to distinguish 
<taxonomicName id="579B4D4C83687E35FC5E628D2A07FF2A" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[986,1150,225,251]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="551" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83687E35FC5E628D2A07FF2A" box="[986,1150,225,251]" italics="true" pageId="8" pageNumber="551">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83687E35FB02628E2ABFFF2A" bold="true" box="[1158,1222,225,251]" pageId="8" pageNumber="551">
<taxonomicNameLabel id="B9DC57A683687E35FB02628E2ABFFF2A" box="[1158,1222,225,251]" pageId="8" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the white patterns observed on the frons also appear distinct, along with the uniform orange coloration. In addition, there is molecular support for 
<taxonomicName id="579B4D4C83687E35FE9E63452FC4FE92" authority="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett, 2020" authorityName="Bahder &amp; Barrantes Barrantes &amp; Zumbado Echavarria &amp; Mou &amp; Helmick &amp; Bartlett" authorityYear="2020" box="[282,445,297,323]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="551" phylum="Arthropoda" rank="species" species="dougwalshi" status="sp. nov.">
<emphasis id="A2EFEADD83687E35FE9E63452FC4FE92" box="[282,445,297,323]" italics="true" pageId="8" pageNumber="551">H. dougwalshi</emphasis>
</taxonomicName>
<emphasis id="A2EFEADD83687E35FE4063462C7DFE92" bold="true" box="[452,516,297,323]" pageId="8" pageNumber="551">
<taxonomicNameLabel id="B9DC57A683687E35FE4063462C7DFE92" box="[452,516,297,323]" pageId="8" pageNumber="551" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
belonging to 
<taxonomicName id="579B4D4C83687E35FD1A63462D68FE92" authorityName="Fowler" authorityYear="1904" box="[670,785,297,323]" class="Insecta" family="Cixiidae" genus="Haplaxius" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="551" phylum="Arthropoda" rank="genus">
<emphasis id="A2EFEADD83687E35FD1A63462D68FE92" box="[670,785,297,323]" italics="true" pageId="8" pageNumber="551">Haplaxius</emphasis>
</taxonomicName>
based on both the COI and 18S loci.
</paragraph>
</subSubSection>
</treatment>
</document>