351 lines
48 KiB
XML
351 lines
48 KiB
XML
<document id="BBBF20F355A10B084E3EF6A8B6A702FB" ID-DOI="10.11646/zootaxa.3602.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5259143" ID-ZooBank="10304FBF-3969-4EFA-83F1-BB8A5E2B37F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629836269209" checkinUser="felipe" docAuthor="TAYLOR, M. L., CAIRNS, S. D., AGNEW, D. J. & ROGERS, A. D." docDate="2013" docId="EE36E867FFC9FFBBFF0AADB5FCEF0FA9" docLanguage="en" docName="zt03602p105.pdf" docOrigin="Zootaxa 3602 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3602.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thouarella moseleyi Wright and Studer 1889" docType="treatment" docVersion="4" lastPageNumber="81" masterDocId="120F901FFF85FFEBFF9DAB72FFEA0E66" masterDocTitle="<strong> A revision of the genus <em> Thouarella </ em> Gray, 1870 (Octocorallia: Primnoidae), including an illustrated dichotomous key, a new species description, and comments on <em> Plumarella </ em> Gray, 1870 and <em> Dasystenella </ em>, Versluys, 1906 </ strong>" masterLastPageNumber="105" masterPageNumber="1" pageNumber="77" updateTime="1699119148921" updateUser="plazi">
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<mods:title id="60FC921ABD08BA0639A7A658DA4728F7"><strong> A revision of the genus <em> Thouarella </ em> Gray, 1870 (Octocorallia: Primnoidae), including an illustrated dichotomous key, a new species description, and comments on <em> Plumarella </ em> Gray, 1870 and <em> Dasystenella </ em>, Versluys, 1906 </ strong></mods:title>
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<mods:namePart id="4B8547CB46E96B3226C14F569F098C2F">TAYLOR, M. L.</mods:namePart>
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<mods:namePart id="A228A2A90C039AB8E8A240A9C0C8C71C">CAIRNS, S. D.</mods:namePart>
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<mods:namePart id="D175719B7BBDFB6F0B00C57EF32C714F">AGNEW, D. J.</mods:namePart>
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<mods:namePart id="0CE9233653CB13D0CECDDA3BE58A8E33">ROGERS, A. D.</mods:namePart>
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<mods:title id="17F65B01794E0A0D39D993E0539BC83B">Zootaxa</mods:title>
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<mods:date id="3210F2107136105E1A93D38A388C8AF8">2013</mods:date>
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<paragraph id="66205971FFC9FFA7FF0AADB5FD180887" blockId="76.[151,754,1735,1794]" box="[151,754,1735,1761]" pageId="76" pageNumber="77">
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<emphasis id="54EB8563FFC9FFA7FF0AADB5FD180887" bold="true" box="[151,754,1735,1761]" pageId="76" pageNumber="77">
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19.
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<taxonomicName id="A19F22F2FFC9FFA7FF5EADB5FD180887" ID-CoL="56L4W" authority="Wright and Studer, 1889" authorityName="Wright and Studer" authorityYear="1889" box="[195,754,1735,1761]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="76" pageNumber="77" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="54EB8563FFC9FFA7FF5EADB5FE530887" bold="true" box="[195,441,1735,1761]" italics="true" pageId="76" pageNumber="77">Thouarella moseleyi</emphasis>
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<bibRefCitation id="020E2480FFC9FFA7FE5CADB5FD180887" author="Wright, E. P. & Studer, T." box="[449,754,1735,1761]" pageId="76" pageNumber="77" pagination="1 - 314" refId="ref65269" refString="Wright, E. P. & Studer, T. (1889) Report on the Alcyonaria collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology, 31 (64), 1 - 314." type="journal article" year="1889">Wright and Studer, 1889</bibRefCitation>
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<paragraph id="66205971FFC9FFA7FF0AAD98FEFE0964" blockId="76.[151,754,1735,1794]" box="[151,276,1770,1794]" pageId="76" pageNumber="77">
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<figureCitation id="FEA445F4FFC9FFA7FF0AAD98FF060964" box="[151,236,1770,1794]" captionStart="FIGURE 30" captionStartId="78.[151,250,1980,2003]" captionTargetBox="[186,1400,177,1976]" captionTargetId="figure-47@78.[186,1401,167,1980]" captionTargetPageId="78" captionText="FIGURE 30. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) 6 cm colony; b) close up of the few remaining polyps; c) lateral view of polyp; d) stereo opercular view." figureDoi="http://doi.org/10.5281/zenodo.5259242" httpUri="https://zenodo.org/record/5259242/files/figure.png" pageId="76" pageNumber="77">Figs 30</figureCitation>
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,
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<figureCitation id="FEA445F4FFC9FFA7FF65AD98FEFE0964" box="[248,276,1770,1794]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="76" pageNumber="77">31</figureCitation>
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</paragraph>
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<taxonomicName id="A19F22F2FFC9FFA7FF0AAC40FD48092E" ID-CoL="56L4W" authority="Wright & Studer, 1889: 61 - 62" authorityName="Wright & Studer" authorityPageNumber="61 - 62" authorityYear="1889" box="[151,674,1842,1864]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="76" pageNumber="77" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="54EB8563FFC9FFA7FF0AAC40FE8F092E" box="[151,357,1842,1864]" italics="true" pageId="76" pageNumber="77">Thouarella moseleyi</emphasis>
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<bibRefCitation id="020E2480FFC9FFA7FEF1AC40FD48092E" author="Wright, E. P. & Studer, T." box="[364,674,1842,1864]" pageId="76" pageNumber="77" pagination="1 - 314" refId="ref65269" refString="Wright, E. P. & Studer, T. (1889) Report on the Alcyonaria collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology, 31 (64), 1 - 314." type="journal article" year="1889">Wright & Studer, 1889: 61–62</bibRefCitation>
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, pl. 14, fig. 1, 1a, pl. 21, fig. 2;
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<treatmentCitation id="E73E7F60FFC9FFA7FC7FAC40FB2B092E" author="Versluys, J." box="[994,1217,1842,1864]" page="29" pageId="76" pageNumber="77" year="1906">
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<bibRefCitation id="020E2480FFC9FFA7FC7FAC40FB2B092E" author="Versluys, J." box="[994,1217,1842,1864]" pageId="76" pageNumber="77" pagination="1 - 187" refId="ref65212" refString="Versluys, J. (1906) Die Gorgoniden der Siboga - Expedition. Siboga-Expeditie, 13 a, 1 - 187." type="book chapter" year="1906">Versluys 1906: 29–30</bibRefCitation>
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</treatmentCitation>
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||
, figs 26, 27 (in text)
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<treatmentCitation id="E73E7F60FFC9FFA7FF0AAC23FD0A0901" author="Kukenthal, W." box="[151,736,1873,1895]" page="150" pageId="76" pageNumber="77" year="1915">
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<taxonomicName id="A19F22F2FFC9FFA7FF0AAC23FD0A0901" ID-CoL="56L4W" authority="Kukenthal 1915: 150" authorityName="Kukenthal" authorityPageNumber="150" authorityYear="1915" box="[151,736,1873,1895]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="76" pageNumber="77" phylum="Cnidaria" rank="species" species="moseleyi" subGenus="Euthouarella">
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<emphasis id="54EB8563FFC9FFA7FF0AAC23FDEB0901" box="[151,513,1873,1895]" italics="true" pageId="76" pageNumber="77">Thouarella (Euthouarella) moseleyi</emphasis>
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<bibRefCitation id="020E2480FFC9FFA7FD9AAC23FD0A0901" author="Kukenthal, W." box="[519,736,1873,1895]" pageId="76" pageNumber="77" pagination="142 - 158" refId="ref63379" refString="Kukenthal, W. (1915) System und Stammesgeschichte der Primnoidae. Zoologischer Anzeiger, 46 (5), 142 - 158." type="journal article" year="1915">Kükenthal 1915: 150</bibRefCitation>
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;
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<treatmentCitation id="E73E7F60FFC9FFA7FD77AC23FC6C0901" author="Kukenthal, W." box="[746,902,1873,1895]" page="417" pageId="76" pageNumber="77" year="1919">
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<bibRefCitation id="020E2480FFC9FFA7FD77AC23FC6C0901" author="Kukenthal, W." box="[746,902,1873,1895]" pageId="76" pageNumber="77" pagination="1 - 946" refId="ref63404" refString="Kukenthal, W. (1919) Gorgonaria. Wissenschaftliche Ergebnisse der deutschen Tiefsee - Expedition auf dem Dampfer " Valdivia " 1898 - 1899, 13, 1 - 946 pp." type="book chapter" year="1919">1919: 417–418</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="E73E7F60FFC9FFA7FC0DAC23FC120901" author="Kukenthal, W." box="[912,1016,1873,1895]" page="292" pageId="76" pageNumber="77" year="1924">
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<bibRefCitation id="020E2480FFC9FFA7FC0DAC23FC120901" author="Kukenthal, W." box="[912,1016,1873,1895]" pageId="76" pageNumber="77" pagination="1 - 478" refId="ref63437" refString="Kukenthal, W. (1924) Coelenterata: Gorgonaria. Das Tierreich. Walter de Gruyter & Co., Berlin, 47, 1 - 478 pp." type="book chapter" year="1924">1924: 292</bibRefCitation>
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</treatmentCitation>
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, 294;
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<treatmentCitation id="E73E7F60FFC9FFA7FBABAC23FADE0901" author="Cairns, S. D. & Bayer, F. M." box="[1078,1332,1873,1895]" page="28" pageId="76" pageNumber="77" year="2009">
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<bibRefCitation id="020E2480FFC9FFA7FBABAC23FADE0901" author="Cairns, S. D. & Bayer, F. M." box="[1078,1332,1873,1895]" pageId="76" pageNumber="77" pagination="1 - 79" refId="ref62273" refString="Cairns, S. D. & Bayer, F. M. (2009) A generic revision and phylogenetic analysis of the Primnoidae (Cnidaria: Octocorallia). Smithsonian Contributions to Zoology, 629, 1 - 79. http: // dx. doi. org / 10.5479 / si. 00810282.629" type="journal article" year="2009">Cairns & Bayer 2009: 28</bibRefCitation>
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</treatmentCitation>
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(listed)
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<treatmentCitation id="E73E7F60FFC9FFA7FF0AAC02FCD709E0" author="Thomson, J. A. & Henderson, W. D." box="[151,829,1904,1926]" page="38" pageId="76" pageNumber="77" year="1906">
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<taxonomicName id="A19F22F2FFC9FFA7FF0AAC02FCD709E0" authority="Thomson & Henderson, 1906: 38" authorityName="Thomson & Henderson" authorityPageNumber="38" authorityYear="1906" box="[151,829,1904,1926]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="76" pageNumber="77" phylum="Cnidaria" rank="variety" species="moseleyi" variety="spicata">
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<emphasis id="54EB8563FFC9FFA7FF0AAC02FE8F09E0" box="[151,357,1904,1926]" italics="true" pageId="76" pageNumber="77">Thouarella moseleyi</emphasis>
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||
var.
|
||
<emphasis id="54EB8563FFC9FFA7FE0AAC03FE0A09E0" box="[407,480,1905,1926]" italics="true" pageId="76" pageNumber="77">spicata</emphasis>
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<bibRefCitation id="020E2480FFC9FFA7FE7AAC02FCD709E0" author="Thomson, J. A. & Henderson, W. D." box="[487,829,1904,1926]" pageId="76" pageNumber="77" pagination="1 - 132" refId="ref64798" refString="Thomson, J. A. & Henderson, W. D. (1906) An Account of the Alcyonarians collected by the Royal Indian Marine Survey Ship Investigator in the Indian Ocean. Part 1. The Alcyonarians of the Deep Sea. The Indian Museum, Calcutta, 1 - 132." type="book chapter" year="1906">Thomson & Henderson, 1906: 38</bibRefCitation>
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</taxonomicName>
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, 42–43, pl. 3, figs 2, 4 (sample not seen)
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</paragraph>
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<paragraph id="66205971FFC9FFA7FF0AACC1FC880988" blockId="76.[151,1436,1970,2030]" pageId="76" pageNumber="77">
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<emphasis id="54EB8563FFC9FFA7FF0AACC1FE8709AD" box="[151,365,1971,1995]" italics="true" pageId="76" pageNumber="77">Material examined</emphasis>
|
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:
|
||
<materialsCitation id="D6F7532CFFC9FFA7FEE3ACC1FC880988" collectingDate="1874-07-15" collectionCode="NHM" collectorName="H. M. S. Challenger & Kermadec Islands, NE" country="New Zealand" elevation="1097" location="Kermadec Islands" pageId="76" pageNumber="77" specimenCode="NHM 1889.5" specimenCount="1" typeStatus="holotype">
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<emphasis id="54EB8563FFC9FFA7FEE3ACC1FE0109AD" bold="true" box="[382,491,1971,1995]" pageId="76" pageNumber="77">
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<typeStatus id="B924E7D3FFC9FFA7FEE3ACC1FE0109AD" box="[382,491,1971,1995]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<specimenCode id="3639F10AFFC9FFA7FE61ACC1FD7F09AC" box="[508,661,1970,1994]" collectionCode="NHM" pageId="76" pageNumber="77">NHM 1889.5</specimenCode>
|
||
.27.39,
|
||
<collectorName id="CB6A3CA7FFC9FFA7FD74ACC0FC2809AD" box="[745,962,1970,1995]" pageId="76" pageNumber="77">
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||
H.M.S.
|
||
<emphasis id="54EB8563FFC9FFA7FCD8ACC1FC2809AD" box="[837,962,1971,1995]" italics="true" pageId="76" pageNumber="77">Challenger</emphasis>
|
||
</collectorName>
|
||
, sta. 171,
|
||
<collectorName id="CB6A3CA7FFC9FFA7FBA6ACC0FAD709AC" box="[1083,1341,1970,1994]" pageId="76" pageNumber="77">
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<location id="63400FAAFFC9FFA7FBA6ACC0FAE009AC" LSID="urn:lsid:plazi:treatment:EE36E867FFC9FFBBFF0AADB5FCEF0FA9:63400FAAFFC9FFA7FBA6ACC0FAE009AC" box="[1083,1290,1970,1994]" country="New Zealand" name="Kermadec Islands" pageId="76" pageNumber="77">Kermadec Islands</location>
|
||
, NE
|
||
</collectorName>
|
||
of
|
||
<collectingCountry id="1E8819E1FFC9FFA7FAFAACC1FF1E0988" name="New Zealand" pageId="76" pageNumber="77">New Zealand</collectingCountry>
|
||
, 28˚33’S, 177˚50’E,
|
||
<quantity id="A167F494FFC9FFA7FE42ACA4FDDC0988" box="[479,566,2006,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="76" pageNumber="77" unit="m" value="1097.0">
|
||
<elevation id="EDB2BE42FFC9FFA7FE42ACA4FDDC0988" box="[479,566,2006,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="76" pageNumber="77" unit="m" value="1097.0">1097 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="12217FB1FFC9FFA7FDDFACA4FD2C0988" box="[578,710,2006,2030]" pageId="76" pageNumber="77" value="1874-07-15">
|
||
<collectingDate id="02658659FFC9FFA7FDDFACA4FD2C0988" box="[578,710,2006,2030]" pageId="76" pageNumber="77" value="1874-07-15">15 Jul 1874</collectingDate>
|
||
</date>
|
||
, 6 cm colony.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AABEAFEF10EC9" blockId="77.[151,1437,152,1399]" box="[151,283,152,175]" pageId="77" pageNumber="78">
|
||
<emphasis id="54EB8563FFC8FFA6FF0AABEAFEF10EC9" box="[151,283,152,175]" italics="true" pageId="77" pageNumber="78">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AABCEFD4F0F02" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The
|
||
<typeStatus id="B924E7D3FFC8FFA6FF57ABCEFEC60EB2" box="[202,300,188,212]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
|
||
was originally approximately
|
||
<quantity id="A167F494FFC8FFA6FD1FABCEFD340EB5" box="[642,734,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="77" pageNumber="78" unit="mm" value="160.0">160 mm</quantity>
|
||
long (from
|
||
<bibRefCitation id="020E2480FFC8FFA6FCFBABCEFB8E0EB2" author="Wright, E. P. & Studer, T." box="[870,1124,188,212]" pageId="77" pageNumber="78" pagination="1 - 314" refId="ref65269" refString="Wright, E. P. & Studer, T. (1889) Report on the Alcyonaria collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology, 31 (64), 1 - 314." type="journal article" year="1889">Wright & Studer 1889</bibRefCitation>
|
||
: pl. 14, fig. 1) but only a
|
||
<quantity id="A167F494FFC8FFA6FA1DABCEFF280E91" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="77" pageNumber="78" unit="mm" value="60.0">60 mm</quantity>
|
||
fragment now remains. The branchlets are pinnate, possibly alternately pinnate (
|
||
<figureCitation id="FEA445F4FFC8FFA6FBE6ABADFB1E0E91" box="[1147,1268,223,247]" captionStart="FIGURE 30" captionStartId="78.[151,250,1980,2003]" captionTargetBox="[186,1400,177,1976]" captionTargetId="figure-47@78.[186,1401,167,1980]" captionTargetPageId="78" captionText="FIGURE 30. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) 6 cm colony; b) close up of the few remaining polyps; c) lateral view of polyp; d) stereo opercular view." figureDoi="http://doi.org/10.5281/zenodo.5259242" httpUri="https://zenodo.org/record/5259242/files/figure.png" pageId="77" pageNumber="78">Fig. 30a,b</figureCitation>
|
||
), although the colony can appear bottlebrush as branchlets are flexible and curve in all directions. The branchlets are mostly simple, some with secondary branching and they are
|
||
<quantity id="A167F494FFC8FFA6FD60AA55FC920F59" box="[765,888,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="77" pageNumber="78" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
|
||
long. The axis is thin, calcareous, flexible, and “somewhat flattened” (
|
||
<bibRefCitation id="020E2480FFC8FFA6FE04AA3EFD720F02" author="Wright, E. P. & Studer, T." box="[409,664,332,356]" pageId="77" pageNumber="78" pagination="1 - 314" refId="ref65269" refString="Wright, E. P. & Studer, T. (1889) Report on the Alcyonaria collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the Years 1873 - 76, Zoology, 31 (64), 1 - 314." type="journal article" year="1889">Wright & Studer 1889</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AAA1DFE990F95" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The polyps are clavate with a tall, conical operculum (
|
||
<figureCitation id="FEA445F4FFC8FFA6FCACAA1DFC780FE1" box="[817,914,367,391]" captionStart="FIGURE 30" captionStartId="78.[151,250,1980,2003]" captionTargetBox="[186,1400,177,1976]" captionTargetId="figure-47@78.[186,1401,167,1980]" captionTargetPageId="78" captionText="FIGURE 30. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) 6 cm colony; b) close up of the few remaining polyps; c) lateral view of polyp; d) stereo opercular view." figureDoi="http://doi.org/10.5281/zenodo.5259242" httpUri="https://zenodo.org/record/5259242/files/figure.png" pageId="77" pageNumber="78">Fig. 30c</figureCitation>
|
||
),
|
||
<quantity id="A167F494FFC8FFA6FC3AAA1DFC170FE1" box="[935,1021,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="77" pageNumber="78" unit="mm" value="1.5">1.5 mm</quantity>
|
||
high, paired, standing perpendicular to the branchlet, with some on the main stem. Each polyp has 6 longitudinal rows of body-wall scales and that number reduces rapidly adaxially from the marginal. There are 4–5 scales in the longitudinal abaxial row, and
|
||
<quantity id="A167F494FFC8FFA6FAEFAAC5FF470F92" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="77" pageNumber="78" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
|
||
the adaxial rows.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AAA8DFCA60C89" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The thin, lanceolate operculars (can be wider with rounded distal edge,
|
||
<figureCitation id="FEA445F4FFC8FFA6FC66AA8DFBB10C71" box="[1019,1115,511,535]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31b</figureCitation>
|
||
) measure 450–640 µm high (average 510 µm), 140–270 µm wide (average 200 µm), with a H:W of 2.2–3.1 (average 2.6). The inner surface is smooth with a simple keel running longitudinally from the tip for three-quarters of the scale length (
|
||
<figureCitation id="FEA445F4FFC8FFA6FAAFA935FA790C39" box="[1330,1427,583,607]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31c</figureCitation>
|
||
) whilst the proximal quarter is tuberculate. Wider operculars have a wider, ridged area on the inner surface rather than a distinct keel (
|
||
<figureCitation id="FEA445F4FFC8FFA6FE0DA9FDFE1D0CC1" box="[400,503,655,679]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31b</figureCitation>
|
||
). The outer scale surface is longitudinally concave and smooth, with granules arranged radially from the proximal centre and tubercles are visible at the proximal edge. The opercular edge appears smooth, but may be worn and thus not representative.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AA98EFDE30DA1" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The marginals are twice as wide as the operculars, 400–410 µm (average 406 µm), 455–510 µm high (average 480 µm), with a H:W of 1.1–1.3 (average 1.2), and are diamond-shaped (
|
||
<figureCitation id="FEA445F4FFC8FFA6FC4CA86DFBA60D51" box="[977,1100,799,823]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31d–f</figureCitation>
|
||
). The inner surface has 4 or 5 tall, thin keels (
|
||
<figureCitation id="FEA445F4FFC8FFA6FED0A836FE550D3A" box="[333,447,836,860]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31d,f</figureCitation>
|
||
) with smooth areas lateral to the keels whilst the remainder is tuberculate. The outer surface is longitudinally concave and smooth, with granules distributed radially from the proximal centre with some tubercles being visible at the proximal edge (
|
||
<figureCitation id="FEA445F4FFC8FFA6FD42A8FEFCAB0DC2" box="[735,833,908,932]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31e</figureCitation>
|
||
). The distal edge is irregular, possibly worn, and the proximal edge is coarsely lobate.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AA8A6FE830A31" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The submarginals are fan-shaped to elliptical (
|
||
<figureCitation id="FEA445F4FFC8FFA6FD4BA8A6FCA70D8A" box="[726,845,980,1004]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">
|
||
Fig.
|
||
<quantity id="A167F494FFC8FFA6FC91A8A6FCD20D8A" box="[780,824,980,1004]" metricMagnitude="-2" metricUnit="kg" metricValue="3.1" pageId="77" pageNumber="78" unit="g" value="31.0">31g</quantity>
|
||
,h
|
||
</figureCitation>
|
||
), 320–420 µm high (average 383 µm), 470–480 µm wide (average 475 µm), with a H:W of 0.7–0.9 (average 0.8). The inner surface is tuberculate, the distal edge having a wide area of perpendicular ridges rather than a keel. The outer surface and scale edges are as the bodywall scales, below.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AAF16FA630A81" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
The body-wall scales are circular to fan-shaped (
|
||
<figureCitation id="FEA445F4FFC8FFA6FD50AF16FCD20A1A" box="[717,824,1124,1148]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">Fig. 31i–l</figureCitation>
|
||
), 300–400 µm high (average 340 µm), 300–475 µm wide (average 370 µm), with a H:W of 0.7–1.2 (average 0.9). The inner surface is tuberculate with a narrow smooth band along the distal edge, and occasionally, small teeth perpendicular to the distal edge. The outer scale surface is smooth with sparse granules across the proximal area whilst the distal edge is smooth and the proximal edge is coarsely lobate.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AAF86FC190B11" blockId="77.[151,1437,152,1399]" pageId="77" pageNumber="78">
|
||
There are 2 layers of coenenchymal scales (
|
||
<figureCitation id="FEA445F4FFC8FFA6FD25AF86FCCE0B6A" box="[696,804,1268,1292]" captionStart="FIGURE 31" captionStartId="79.[151,250,1912,1935]" captionTargetBox="[221,1366,177,1907]" captionTargetId="figure-80@79.[221,1366,159,1907]" captionTargetPageId="79" captionText="FIGURE 31. Thouarella moseleyi, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR." figureDoi="http://doi.org/10.5281/zenodo.5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="77" pageNumber="78">
|
||
Fig.
|
||
<quantity id="A167F494FFC8FFA6FD70AF86FCCE0B6A" box="[749,804,1268,1292]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="77" pageNumber="78" unit="m" value="31.0">31m</quantity>
|
||
</figureCitation>
|
||
): sclerites from the inner layer are circular in shape and sclerites from the outside layer are elliptical. The inner surface of the coenenchymal scales is tuberculate with a smooth band running along the distal edge whilst the outer surface is smooth, sometimes with sparsely placed granules. The distal edge is smooth and the proximal edge is coarsely lobate.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AAEDAFECA0BA6" blockId="77.[151,1436,1448,1579]" box="[151,288,1448,1472]" pageId="77" pageNumber="78">
|
||
<emphasis id="54EB8563FFC8FFA6FF0AAEDAFECA0BA6" box="[151,288,1448,1472]" italics="true" pageId="77" pageNumber="78">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AAEBEFF5E084D" blockId="77.[151,1436,1448,1579]" pageId="77" pageNumber="78">
|
||
This species is known from 2 locations (excluding
|
||
<taxonomicName id="A19F22F2FFC8FFA6FD42AEBEFC880B82" authorityName="Thomson & Henderson" authorityYear="1906" box="[735,866,1484,1508]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="variety" species="moseleyi" variety="spicata">
|
||
var.
|
||
<emphasis id="54EB8563FFC8FFA6FC8CAEBFFC880B82" box="[785,866,1485,1508]" italics="true" pageId="77" pageNumber="78">spicata</emphasis>
|
||
</taxonomicName>
|
||
), northwest of the Kermadec Islands in the South Pacific and north of Sumbawa Island in
|
||
<collectingCountry id="1E8819E1FFC8FFA6FDFDAE9DFD3A0861" box="[608,720,1519,1543]" name="Indonesia" pageId="77" pageNumber="78">Indonesia</collectingCountry>
|
||
,
|
||
<quantity id="A167F494FFC8FFA6FD41AE9DFCAA0861" box="[732,832,1519,1543]" metricMagnitude="6" metricUnit="m" metricValue="6.6" pageId="77" pageNumber="78" unit="km" value="6600.0">6600 km</quantity>
|
||
away. It has been recorded from depths of
|
||
<quantity id="A167F494FFC8FFA6FAB1AE9DFF5A084D" metricMagnitude="2" metricUnit="m" metricValue="9.455" metricValueMax="10.97" metricValueMin="7.94" pageId="77" pageNumber="78" unit="m" value="945.5" valueMax="1097.0" valueMin="794.0">794–1097 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AAD2EFF130812" blockId="77.[151,1436,1628,1940]" box="[151,249,1628,1652]" pageId="77" pageNumber="78">
|
||
<emphasis id="54EB8563FFC8FFA6FF0AAD2EFF130812" box="[151,249,1628,1652]" italics="true" pageId="77" pageNumber="78">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF0AAD0DFD2808B9" blockId="77.[151,1436,1628,1940]" pageId="77" pageNumber="78">
|
||
The number of polyps per cm was impossible to tell from the small
|
||
<typeStatus id="B924E7D3FFC8FFA6FC12AD0DFC1B08F1" box="[911,1009,1663,1687]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
|
||
fragment that remains. More material of this species from the type locality is required to confirm the differences between it and
|
||
<taxonomicName id="A19F22F2FFC8FFA6FB18ADD7FB3B08DA" authorityName="Versluys" authorityYear="1906" box="[1157,1233,1700,1724]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="species" species="laxa">
|
||
<emphasis id="54EB8563FFC8FFA6FB18ADD7FB3B08DA" box="[1157,1233,1700,1724]" italics="true" pageId="77" pageNumber="78">T. laxa</emphasis>
|
||
</taxonomicName>
|
||
, which has a very similar polyp morphology and pinnate branching.
|
||
</paragraph>
|
||
<paragraph id="66205971FFC8FFA6FF5AAD9EFEC009F2" blockId="77.[151,1436,1628,1940]" pageId="77" pageNumber="78">
|
||
A variety of
|
||
<taxonomicName id="A19F22F2FFC8FFA6FEFFAD9FFE090962" authorityName="Kukenthal" authorityYear="1915" box="[354,483,1772,1796]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="54EB8563FFC8FFA6FEFFAD9FFE090962" box="[354,483,1772,1796]" italics="true" pageId="77" pageNumber="78">T. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="A19F22F2FFC8FFA6FE6BAD9EFD970962" authorityName="Thomson & Henderson" authorityYear="1906" box="[502,637,1772,1796]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="variety" species="moseleyi" variety="spicata">
|
||
var.
|
||
<emphasis id="54EB8563FFC8FFA6FDB1AD9FFD970962" box="[556,637,1773,1796]" italics="true" pageId="77" pageNumber="78">spicata</emphasis>
|
||
</taxonomicName>
|
||
) was described in
|
||
<bibRefCitation id="020E2480FFC8FFA6FCFDAD9EFB300962" author="Thomson, J. A. & Henderson, W. D." box="[864,1242,1772,1796]" pageId="77" pageNumber="78" pagination="1 - 132" refId="ref64798" refString="Thomson, J. A. & Henderson, W. D. (1906) An Account of the Alcyonarians collected by the Royal Indian Marine Survey Ship Investigator in the Indian Ocean. Part 1. The Alcyonarians of the Deep Sea. The Indian Museum, Calcutta, 1 - 132." type="book chapter" year="1906">Thomson and Henderson (1906)</bibRefCitation>
|
||
. The differences between the polyps of the
|
||
<typeStatus id="B924E7D3FFC8FFA6FE20AC7DFDF50941" box="[445,543,1807,1831]" pageId="77" pageNumber="78" type="holotype">holotype</typeStatus>
|
||
and those of the variant are mainly that the latter has a taller conical operculum and marginal scales with a more elongated, pointed spine (which may be bifid). The tall conical operculum is similar to that of the polyps of
|
||
<taxonomicName id="A19F22F2FFC8FFA6FE65AC2BFD6E0916" authorityName="Cairns" authorityYear="2006" box="[504,644,1880,1904]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="species" species="grasshoffi">
|
||
<emphasis id="54EB8563FFC8FFA6FE65AC2BFD6E0916" box="[504,644,1880,1904]" italics="true" pageId="77" pageNumber="78">T. grasshoffi</emphasis>
|
||
</taxonomicName>
|
||
. However, colonies of
|
||
<taxonomicName id="A19F22F2FFC8FFA6FC11AC2BFBF30916" authorityName="Cairns" authorityYear="2006" box="[908,1049,1880,1904]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="77" pageNumber="78" phylum="Cnidaria" rank="species" species="grasshoffi">
|
||
<emphasis id="54EB8563FFC8FFA6FC11AC2BFBF30916" box="[908,1049,1880,1904]" italics="true" pageId="77" pageNumber="78">T. grasshoffi</emphasis>
|
||
</taxonomicName>
|
||
have a true bottlebrush branching arrangement.
|
||
</paragraph>
|
||
<caption id="32E009F9FFCBFFA5FF0AACCEFD9B0997" ID-DOI="http://doi.org/10.5281/zenodo.5259242" ID-Zenodo-Dep="5259242" httpUri="https://zenodo.org/record/5259242/files/figure.png" pageId="78" pageNumber="79" startId="78.[151,250,1980,2003]" targetBox="[186,1400,177,1976]" targetPageId="78">
|
||
<paragraph id="66205971FFCBFFA5FF0AACCEFD9B0997" blockId="78.[151,1436,1980,2033]" pageId="78" pageNumber="79">
|
||
<emphasis id="54EB8563FFCBFFA5FF0AACCEFECA09B5" bold="true" box="[151,288,1980,2003]" pageId="78" pageNumber="79">FIGURE 30.</emphasis>
|
||
<emphasis id="54EB8563FFCBFFA5FEBBACCFFE1609B5" box="[294,508,1981,2003]" italics="true" pageId="78" pageNumber="79">
|
||
<taxonomicName id="A19F22F2FFCBFFA5FEBBACCFFE1209B5" authorityName="Kukenthal" authorityYear="1915" box="[294,504,1981,2003]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="78" pageNumber="79" phylum="Cnidaria" rank="species" species="moseleyi">Thouarella moseleyi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype, NHM 1889.5.27.39: a) 6 cm colony; b) close up of the few remaining polyps; c) lateral view of polyp; d) stereo opercular view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="32E009F9FFCAFFA4FF0AAC0AFE810992" ID-DOI="http://doi.org/10.5281/zenodo.5259244" ID-Zenodo-Dep="5259244" httpUri="https://zenodo.org/record/5259244/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,250,1912,1935]" targetBox="[221,1366,177,1907]" targetPageId="79">
|
||
<paragraph id="66205971FFCAFFA4FF0AAC0AFE810992" blockId="79.[151,1437,1912,2036]" pageId="79" pageNumber="80">
|
||
<emphasis id="54EB8563FFCAFFA4FF0AAC0AFEF509E9" bold="true" box="[151,287,1912,1935]" pageId="79" pageNumber="80">FIGURE 31.</emphasis>
|
||
<taxonomicName id="A19F22F2FFCAFFA4FEBBAC0AFE1F09E8" authorityName="Kukenthal" authorityYear="1915" box="[294,501,1912,1934]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="79" pageNumber="80" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="54EB8563FFCAFFA4FEBBAC0AFE1F09E8" box="[294,501,1912,1934]" italics="true" pageId="79" pageNumber="80">Thouarella moseleyi</emphasis>
|
||
</taxonomicName>
|
||
, holotype, NHM 1889.5.27.39: a) outer and b,c) inner surface of opercular scales; d,f) inner and e) outer surface of marginal scales; g,h) inner surface of submarginal scales; i,l) inner and j,k) outer surface of body-wall scales; m) coenenchymal scales—outer surface shown on top left and bottom right scales, remainder show the inner surface. Some SEM by ZGR.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="66205971FFD5FFBBFF0AABEAFEC60ED6" blockId="80.[151,1436,152,464]" box="[151,300,152,176]" pageId="80" pageNumber="81">
|
||
<emphasis id="54EB8563FFD5FFBBFF0AABEAFEC60ED6" box="[151,300,152,176]" italics="true" pageId="80" pageNumber="81">Comparisons</emphasis>
|
||
</paragraph>
|
||
<paragraph id="66205971FFD5FFBBFF0AABCEFC9B0F02" blockId="80.[151,1436,152,464]" pageId="80" pageNumber="81">
|
||
Within Group 2 only two other species,
|
||
<taxonomicName id="A19F22F2FFD5FFBBFDF3ABCFFD560EB2" authorityName="Versluys" authorityYear="1906" box="[622,700,188,212]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="laxa">
|
||
<emphasis id="54EB8563FFD5FFBBFDF3ABCFFD560EB2" box="[622,700,188,212]" italics="true" pageId="80" pageNumber="81">T. laxa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="54EB8563FFD5FFBBFD67ABCFFC6C0EB2" box="[762,902,188,212]" italics="true" pageId="80" pageNumber="81">
|
||
<taxonomicName id="A19F22F2FFD5FFBBFD67ABCFFC690EB2" authorityName="Versluys" authorityYear="1906" box="[762,899,188,212]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="tydemani">T. tydemani</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
have pinnate colonies or colonies that appear pinnate. The ridges running perpendicular to the distal edge of the body-wall scales on the polyps of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFB6BAB93FA990E9E" authorityName="Kukenthal" authorityYear="1915" box="[1270,1395,224,248]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="54EB8563FFD5FFBBFB6BAB93FA990E9E" box="[1270,1395,224,248]" italics="true" pageId="80" pageNumber="81">T. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
are similar to those reported on the body-wall scales in a recent description of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFB97AA77FE430F59" authority="(Zapata-Guardiola and Lopez-Gonzalez 2010 a)" baseAuthorityName="Zapata-Guardiola and Lopez-Gonzalez" baseAuthorityYear="2010" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="tydemani">
|
||
<emphasis id="54EB8563FFD5FFBBFB97AA77FB650F7A" box="[1034,1167,260,284]" italics="true" pageId="80" pageNumber="81">T. tydemani</emphasis>
|
||
(
|
||
<bibRefCitation id="020E2480FFD5FFBBFB3CAA76FE4B0F59" author="Zapata-Guardiola, R. & Lopez-Gonzalez, P. J." pageId="80" pageNumber="81" pagination="169 - 180" refId="ref65337" refString="Zapata-Guardiola, R. & Lopez-Gonzalez, P. J. (2010 a) Two new species of Antarctic gorgonians (Octocorallia: Primnoidae) with a redescription of Thouarella laxa Versluys, 1906. Helgoland Marine Research, 64 (3), 169 - 180. http: // dx. doi. org / 10.1007 / s 10152 - 009 - 0176 - 5" type="journal article" year="2010">Zapata-Guardiola and López-González 2010a</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The polyps of
|
||
<emphasis id="54EB8563FFD5FFBBFDC4AA5BFD080F26" box="[601,738,296,320]" italics="true" pageId="80" pageNumber="81">
|
||
<taxonomicName id="A19F22F2FFD5FFBBFDC4AA5BFD350F26" baseAuthorityName="Zapata-Guardiola and Lopez-Gonzalez" baseAuthorityYear="2010" box="[601,735,296,320]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="tydemani">T. tydemani</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
however, have long spinose marginals and are distally flared, whereas polyps of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFEF4AA3FFE0D0F02" authorityName="Kukenthal" authorityYear="1915" box="[361,487,332,356]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="54EB8563FFD5FFBBFEF4AA3FFE0D0F02" box="[361,487,332,356]" italics="true" pageId="80" pageNumber="81">T. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
are clavate with shorter marginals.
|
||
</paragraph>
|
||
<paragraph id="66205971FFD5FFBBFF5AAA1DFCEF0FA9" blockId="80.[151,1436,152,464]" pageId="80" pageNumber="81">
|
||
The polyps of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFEF1AA03FE010FEE" authorityName="Kukenthal" authorityYear="1915" box="[364,491,368,392]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="54EB8563FFD5FFBBFEF1AA03FE010FEE" box="[364,491,368,392]" italics="true" pageId="80" pageNumber="81">T. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
are more robust than those of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFCDAAA03FC790FEE" authorityName="Versluys" authorityYear="1906" box="[839,915,368,392]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="laxa">
|
||
<emphasis id="54EB8563FFD5FFBBFCDAAA03FC790FEE" box="[839,915,368,392]" italics="true" pageId="80" pageNumber="81">T. laxa</emphasis>
|
||
</taxonomicName>
|
||
, with wider marginals that are not as spinose, and a fitted opercular cone. There are also several ridges running perpendicular to the distal edge of the submarginals, which are absent in the polyps of
|
||
<taxonomicName id="A19F22F2FFD5FFBBFD2EAACBFD140FB6" authorityName="Versluys" authorityYear="1906" box="[691,766,440,464]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="80" pageNumber="81" phylum="Cnidaria" rank="species" species="laxa">
|
||
<emphasis id="54EB8563FFD5FFBBFD2EAACBFD140FB6" box="[691,766,440,464]" italics="true" pageId="80" pageNumber="81">T. laxa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |