168 lines
19 KiB
XML
168 lines
19 KiB
XML
<document id="91B1699C8AA1EAD8DC4F1A24DFDD0F49" ID-CLB-Dataset="2719" ID-DOI="10.11646/palaeoentomology.4.6.4" ID-GBIF-Dataset="09972b89-239e-49e6-af29-5bd9aa769f2d" ID-ISSN="2624-2834" ID-Zenodo-Dep="5778566" ID-ZooBank="FAB39DFF-4118-4AC4-870B-257458D56FA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639469354819" checkinUser="plazi" docAuthor="Radchenko, Alexander G." docDate="2021" docId="CE758787FFF0591AFC87FED7D22AFB00" docLanguage="en" docName="palaeoentomology.4.6.4.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Drymomyrmex rasnitsyni Radchenko 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="546" masterDocId="324CFFFFFFF15918FFADFFD8D76CFF84" masterDocTitle="New species of the fossil ant genus Drymomyrmex (Hymenoptera, Formicidae, Formicinae) from the late Eocene Rovno amber (Ukraine)" masterLastPageNumber="549" masterPageNumber="544" pageNumber="545" updateTime="1699472402092" updateUser="ExternalLinkService">
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<mods:mods id="95CE216DF8E648203433519D4FC14F96" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DD3189C629B7912DE93E1A646401B394">
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<mods:title id="A62580DBCFD4A30474843266CDA7523F">New species of the fossil ant genus Drymomyrmex (Hymenoptera, Formicidae, Formicinae) from the late Eocene Rovno amber (Ukraine)</mods:title>
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</mods:titleInfo>
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<mods:name id="CA44DC9B0B929264A77AD109DDDA3408" type="personal">
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<mods:role id="C93C0C46CB8D296AA9768B778F7B1948">
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<mods:roleTerm id="D15B2011F088EF7480A52CB7AEEBA2C5">Author</mods:roleTerm>
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<mods:namePart id="63E149B51772FA9A62A9A81A1DDF6840">Radchenko, Alexander G.</mods:namePart>
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<mods:typeOfResource id="3157AA08B47A8D86AB32E1B13D7C405C">text</mods:typeOfResource>
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<mods:title id="BD3C4BD6CA3640ADA1A9DCD4F5235B25">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="8BD12892F78E7AB9481C985CF661AD4B">
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<mods:date id="09420904A8F2A11B52D874B9DEC94781">2021</mods:date>
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<mods:detail id="F6AEAEE899205793283D3F0022B8DD4F" type="pubDate">
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<mods:number id="B214A5CC4E2551EDBBD250E8210B553F">2021-12-14</mods:number>
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</mods:detail>
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<mods:number id="A2230489CBFDF50790AC833605960CFB">4</mods:number>
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<mods:detail id="670622C9A5DAB0B13D078E1FF4619739" type="issue">
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<mods:number id="7171443D8EC122EA77E7B0E04EA15741">6</mods:number>
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<mods:start id="69680E8D0DC309124130971B36B8A34F">544</mods:start>
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<mods:classification id="5CC89107E72095C48E841C3B43957E89">journal article</mods:classification>
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<mods:identifier id="7B1FEC409B66FCFE40A96CB3D7AE811F" type="CLB-Dataset">2719</mods:identifier>
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<mods:identifier id="A4C49EC39F38765150BD3A74B4F4319F" type="DOI">10.11646/palaeoentomology.4.6.4</mods:identifier>
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<mods:identifier id="CAC6E97A65C5ACB59D08E1014B29AB53" type="GBIF-Dataset">09972b89-239e-49e6-af29-5bd9aa769f2d</mods:identifier>
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<mods:identifier id="D43BB7557CF96CD0B87C5123986F386D" type="ISSN">2624-2834</mods:identifier>
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<mods:identifier id="BA68C997ED208D5006D3BDBF16BC1CB9" type="Zenodo-Dep">5778566</mods:identifier>
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<mods:identifier id="F21BF0163BDEC2457C79A9B8CB61A987" type="ZooBank">FAB39DFF-4118-4AC4-870B-257458D56FA9</mods:identifier>
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</mods:mods>
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<treatment id="CE758787FFF0591AFC87FED7D22AFB00" ID-DOI="http://doi.org/10.5281/zenodo.5833905" ID-GBIF-Taxon="191975187" ID-Zenodo-Dep="5833905" LSID="urn:lsid:plazi:treatment:CE758787FFF0591AFC87FED7D22AFB00" httpUri="http://treatment.plazi.org/id/CE758787FFF0591AFC87FED7D22AFB00" lastPageId="2" lastPageNumber="546" pageId="1" pageNumber="545">
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<subSubSection id="0EC6651AFFF05919FC87FED7D3C3FEAD" box="[810,1199,270,297]" pageId="1" pageNumber="545" type="nomenclature">
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<paragraph id="46633691FFF05919FC87FED7D3C3FEAD" blockId="1.[810,1199,270,334]" box="[810,1199,270,297]" pageId="1" pageNumber="545">
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<heading id="1D2B81FDFFF05919FC87FED7D3C3FEAD" bold="true" box="[810,1199,270,297]" fontSize="10" level="2" pageId="1" pageNumber="545" reason="3">
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<emphasis id="74A8EA83FFF05919FC87FED7D3C3FEAD" bold="true" box="[810,1199,270,297]" pageId="1" pageNumber="545">
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<taxonomicName id="81DC4D12FFF05919FC87FED7D323FEAC" authority="Radchenko, 2021" authorityName="Radchenko" authorityYear="2021" box="[810,1103,270,297]" class="Insecta" family="Formicidae" genus="Drymomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="545" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="74A8EA83FFF05919FC87FED7D323FEAC" bold="true" box="[810,1103,270,297]" italics="true" pageId="1" pageNumber="545">Drymomyrmex rasnitsyni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6F9B57F8FFF05919FBFBFED7D3C3FEAD" box="[1110,1199,271,297]" pageId="1" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC6651AFFF05919FC87FEECD4F1FECA" box="[810,925,308,334]" pageId="1" pageNumber="545" type="description">
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<paragraph id="46633691FFF05919FC87FEECD4F1FECA" blockId="1.[810,1199,270,334]" box="[810,925,308,334]" pageId="1" pageNumber="545">
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(
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<figureCitation id="DEE72A14FFF05919FC9FFEECD4F8FECA" box="[818,916,308,334]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[810,909,1977,2001]" captionStartId-1="2.[136,235,705,729]" captionStartId-2="3.[136,235,1254,1278]" captionTargetBox-0="[818,1444,776,1953]" captionTargetBox-1="[149,759,186,661]" captionTargetBox-2="[439,1145,206,1211]" captionTargetId-0="figure-464@1.[818,1444,776,1953]" captionTargetId-1="figure-700@2.[139,774,172,681]" captionTargetId-2="figure-358@3.[431,1157,189,1230]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Photographs of Drymomyrmex rasnitsyni sp. nov., holotype, SIZK No. DO-136. A, Anterolateral view. B, Left dorsolateral view. C, Right dorsolateral view." captionText-1="FIGURE 2. Photograph of piece of amber with the holotype worker of Drymomyrmex rasnitsyni sp. nov." captionText-2="FIGURE 3. Line drawings of Drymomyrmex rasnitsyni sp. nov., holotype. A, Head, dorsal view. B and C, Right dorsolateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5778568" figureDoi-1="http://doi.org/10.5281/zenodo.5778570" figureDoi-2="http://doi.org/10.5281/zenodo.5778572" httpUri-0="https://zenodo.org/record/5778568/files/figure.png" httpUri-1="https://zenodo.org/record/5778570/files/figure.png" httpUri-2="https://zenodo.org/record/5778572/files/figure.png" pageId="1" pageNumber="545">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC6651AFFF05919FC87FE58D4DFFE62" pageId="1" pageNumber="545" type="materials_examined">
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<paragraph id="46633691FFF05919FC87FE58D4DFFE62" blockId="1.[810,1452,384,750]" pageId="1" pageNumber="545">
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<materialsCitation id="F6B43CCCFFF05919FC87FE58D4DFFE62" ID-GBIF-Occurrence="3429136301" collectionCode="SIZK" country="Ukraine" location="Osova" municipality="Dubrovitsy Distr." pageId="1" pageNumber="545" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
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<emphasis id="74A8EA83FFF05919FC87FE58D4F2FE1E" bold="true" box="[810,926,384,410]" pageId="1" pageNumber="545">
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<typeStatus id="99678833FFF05919FC87FE58D4F5FE1E" box="[810,921,384,410]" pageId="1" pageNumber="545" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Worker,
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<collectionCode id="20CDAE54FFF05919FBA6FE58D325FE1E" box="[1035,1097,384,410]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="1" pageNumber="545">SIZK</collectionCode>
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No. DO-136,
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<collectingCountry id="3ECB7601FFF05919FB58FE58D23EFE1E" box="[1269,1362,384,410]" name="Ukraine" pageId="1" pageNumber="545">Ukraine</collectingCountry>
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,
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<collectingRegion id="8418F873FFF05919FACCFE58D40BFE44" country="Ukraine" name="Rivne" pageId="1" pageNumber="545">Rovno Prov.</collectingRegion>
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,
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<collectingMunicipality id="A607ACEBFFF05919FCD5FE7DD32FFE3B" box="[888,1091,421,447]" pageId="1" pageNumber="545">Dubrovitsy Distr.</collectingMunicipality>
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, village
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<location id="4303604AFFF05919FB00FE7ED39BFE44" LSID="urn:lsid:plazi:treatment:CE758787FFF0591AFC87FED7D22AFB00:4303604AFFF05919FB00FE7ED39BFE44" box="[1197,1271,422,448]" country="Ukraine" municipality="Dubrovitsy Distr." name="Osova" pageId="1" pageNumber="545" stateProvince="Rivne">Osova</location>
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,
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<collectingRegion id="8418F873FFF05919FAA5FE7ED23FFE44" box="[1288,1363,422,448]" country="Ukraine" name="Rivne" pageId="1" pageNumber="545">Rovno</collectingRegion>
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amber, late Eocene.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC6651AFFF05919FCF2FE29D36CFDD2" pageId="1" pageNumber="545" type="etymology">
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<paragraph id="46633691FFF05919FCF2FE29D36CFDD2" blockId="1.[810,1452,384,750]" pageId="1" pageNumber="545">
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<emphasis id="74A8EA83FFF05919FCF2FE29D48AFD8F" bold="true" box="[863,998,497,523]" pageId="1" pageNumber="545">Etymology.</emphasis>
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The new species is named in honour of Professor Alexander P. Rasnitsyn, famous Russian paleoentomologist.
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC6651AFFF05919FCF2FDBAD23FFD6A" pageId="1" pageNumber="545" type="diagnosis">
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<paragraph id="46633691FFF05919FCF2FDBAD23FFD6A" blockId="1.[810,1452,384,750]" pageId="1" pageNumber="545">
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<emphasis id="74A8EA83FFF05919FCF2FDBAD4B5FDF8" bold="true" box="[863,985,610,636]" pageId="1" pageNumber="545">Diagnosis.</emphasis>
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Head elongated; antennae 11-segmented; mesonotum completely fused with metanotum and the latter does not form a distinctly separate sclerite; meso- and metatibiae with well developed simple spurs.
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</paragraph>
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</subSubSection>
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<caption id="12A36619FFF3591AFF25FD19D525FD79" ID-DOI="http://doi.org/10.5281/zenodo.5778570" ID-Zenodo-Dep="5778570" httpUri="https://zenodo.org/record/5778570/files/figure.png" pageId="2" pageNumber="546" startId="2.[136,235,705,729]" targetBox="[149,759,186,661]" targetPageId="2">
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||
<paragraph id="46633691FFF3591AFF25FD19D525FD79" blockId="2.[136,778,705,765]" pageId="2" pageNumber="546">
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<emphasis id="74A8EA83FFF3591AFF25FD19D66BFD5D" bold="true" box="[136,263,705,729]" pageId="2" pageNumber="546">FIGURE 2.</emphasis>
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Photograph of piece of amber with the holotype worker of
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<taxonomicName id="81DC4D12FFF3591AFF5FFD3ED69EFD79" authority="Radchenko, 2021" authorityName="Radchenko" authorityYear="2021" box="[242,498,742,765]" class="Insecta" family="Formicidae" genus="Drymomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="546" phylum="Arthropoda" rank="species" species="rasnitsyni" status="sp. nov.">
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<emphasis id="74A8EA83FFF3591AFF5FFD3ED69EFD79" box="[242,498,742,765]" italics="true" pageId="2" pageNumber="546">Drymomyrmex rasnitsyni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6F9B57F8FFF3591AFE54FD3DD525FD79" box="[505,585,741,765]" pageId="2" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<subSubSection id="0EC6651AFFF3591AFF10FCE4D578FC1A" pageId="2" pageNumber="546" type="materials_examined">
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<paragraph id="46633691FFF3591AFF10FCE4D578FC1A" blockId="2.[136,778,828,2078]" pageId="2" pageNumber="546">
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<emphasis id="74A8EA83FFF3591AFF10FCE4D692FCD2" bold="true" box="[189,510,828,854]" pageId="2" pageNumber="546">Type locality and horizon.</emphasis>
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<materialsCitation id="F6B43CCCFFF3591AFDA5FCE4D57FFCFE" ID-GBIF-Occurrence="3429136302" country="Ukraine" location="Dubrovitsy Distr." pageId="2" pageNumber="546" specimenCount="1" stateProvince="Rivne">
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<collectingCountry id="3ECB7601FFF3591AFDA5FCE4D509FCD2" box="[520,613,828,854]" name="Ukraine" pageId="2" pageNumber="546">Ukraine</collectingCountry>
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,
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<collectingRegion id="8418F873FFF3591AFDD9FCE4D46AFCD2" box="[628,774,828,854]" country="Ukraine" name="Rivne" pageId="2" pageNumber="546">Rovno Prov.</collectingRegion>
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,
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<location id="4303604AFFF3591AFF25FCB8D634FCFE" LSID="urn:lsid:plazi:treatment:CE758787FFF0591AFC87FED7D22AFB00:4303604AFFF3591AFF25FCB8D634FCFE" box="[136,344,864,890]" country="Ukraine" name="Dubrovitsy Distr." pageId="2" pageNumber="546" stateProvince="Rivne">Dubrovitsy Distr.</location>
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, village Osova
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</materialsCitation>
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,
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<collectingRegion id="8418F873FFF3591AFD84FCB8D518FCFE" box="[553,628,864,890]" country="Ukraine" name="Rivne" pageId="2" pageNumber="546">Rovno</collectingRegion>
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amber, late Eocene, Priabonian, 33.9–37.8 Ma.
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</paragraph>
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</subSubSection>
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<subSubSection id="0EC6651AFFF3591AFF10FC70D22AFB00" pageId="2" pageNumber="546" type="description">
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<paragraph id="46633691FFF3591AFF10FC70D6D0F95E" blockId="2.[136,778,828,2078]" pageId="2" pageNumber="546">
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<emphasis id="74A8EA83FFF3591AFF10FC70D623FC46" bold="true" box="[189,335,936,962]" pageId="2" pageNumber="546">Description.</emphasis>
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Total length
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<emphasis id="74A8EA83FFF3591AFE4AFC71D56DFC46" box="[487,513,937,962]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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.
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<quantity id="81249B74FFF3591AFDBCFC70D507FC46" box="[529,619,936,963]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="546" unit="mm" value="3.5">3.5 mm</quantity>
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. Despite head width not properly measurable, head definitely elongated, seems at least
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<emphasis id="74A8EA83FFF3591AFE94FC29D63FFB8E" box="[313,339,1009,1034]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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. 1.4 times as long as wide; sides of head very slightly convex, subparallel, occipital corners narrowly rounded, occipital margin straight. Antennae 11-segmented, inserted to head just near posterior margin of clypeus; scape relatively short, slightly does not reach occipital margin; funicular segments gradually enlarged from 2
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<superScript id="B1A99BD9FFF3591AFF78FB10D789FB52" attach="left" box="[213,229,1224,1238]" fontSize="6" pageId="2" pageNumber="546">nd</superScript>
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to apical, 1
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<superScript id="B1A99BD9FFF3591AFEC1FB10D61BFB52" attach="left" box="[364,375,1224,1238]" fontSize="6" pageId="2" pageNumber="546">st</superScript>
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segment
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<emphasis id="74A8EA83FFF3591AFE49FB11D692FB66" box="[484,510,1225,1250]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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. twice as long as wide, 2–5
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<superScript id="B1A99BD9FFF3591AFF1FFB34D7D3FB7E" attach="left" box="[178,191,1260,1274]" fontSize="6" pageId="2" pageNumber="546">th</superScript>
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transversal, 6–9
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<superScript id="B1A99BD9FFF3591AFE2FFB34D6E3FB7E" attach="left" box="[386,399,1260,1274]" fontSize="6" pageId="2" pageNumber="546">th</superScript>
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subsquare, 10
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<superScript id="B1A99BD9FFF3591AFD96FB34D524FB7E" attach="left" box="[571,584,1260,1274]" fontSize="6" pageId="2" pageNumber="546">th</superScript>
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the longest,
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<emphasis id="74A8EA83FFF3591AFD45FB35D46EFA82" box="[744,770,1261,1286]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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. 1.5 times as long as two preceding segments together. Eyes of moderate size (OI 0.26), situated approximately at midlength of sides of head, genae distinctly longer than maximum diameter of eye (GI 1.29). Ocelli absent. Surface of median portion of clypeus evenly convex, not truncated anteriorly over mandibles, its anterior margin feebly convex. Frontal carinae very short, not reaching level of anterior margin of eyes, clypeus not inserted between frontal carinae; anterior part of frons with not coarse longitudinal carina (instead of groove); frontal triangle small, but distinct. Mandibles triangular, but obscured, number of teeth cannot be counted. Maxillary and labial palps not visible.
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</paragraph>
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<paragraph id="46633691FFF3591AFF10F93CD224FE78" blockId="2.[136,778,828,2078]" lastBlockId="2.[810,1452,158,1157]" pageId="2" pageNumber="546">
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Mesosoma relatively stout,
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<emphasis id="74A8EA83FFF3591AFDA5F93DD54EF97A" box="[520,546,1765,1790]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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. 1.3 times as long as high, pro-mesonotal suture very clear and deep, mesonotum slightly raised above pronotum, inclined to propodeum; mesonotum fused with metanotum, so that metanotum does not form distinctly separate sclerite (seen from above); propodeal dorsum much lower than promesonotum (
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<emphasis id="74A8EA83FFF3591AFEEAF865D623F852" box="[327,335,1981,2006]" italics="true" pageId="2" pageNumber="546">i</emphasis>
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.
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<emphasis id="74A8EA83FFF3591AFEFBF865D60EF852" box="[342,354,1981,2006]" italics="true" pageId="2" pageNumber="546">e</emphasis>
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., mesosoma does not form regular arch in profile); propodeum gradually rounded, without teeth or tubercles, its dorsal surface somewhat shorter than declivous one; propodeal spiracles rounded, located quite close to declivity. Petiole with short posterior peduncle, its scale slightly inclined forward, thick, relatively low, with subparallel anterior and posterior surfaces and widely rounded dorsum; gaster partially overhangs posterior petiolar peduncle. Metacoxae widely separated (seen from below). Legs of moderate length, fore femora quite wide,
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<emphasis id="74A8EA83FFF3591AFC03FE43D4A4FE30" box="[942,968,411,436]" italics="true" pageId="2" pageNumber="546">ca</emphasis>
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. 3 times as long as width; middle and hind tibiae with well developed simple spur, which longer than maximum width of tibia; pretarsal claws simple.
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</paragraph>
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<paragraph id="46633691FFF3591AFCF2FDDED305FC70" blockId="2.[810,1452,158,1157]" pageId="2" pageNumber="546">Head dorsum and appendages with short and relatively sparse decumbent pubescence, length of appressed hairs subequal to distance between them, but such pubescence much denser in ventral and lateral surfaces of head, mesosoma and gaster, where hairs much longer than distance between them. Frons and occiput with infrequent quite long erect to suberect hairs, occipital margin and margin of temples also with very dense short subdecumbent hairs; mesosoma and petiole with rather long, often curved erect to suberect hairs, gaster with similar, but shorter hairs; antennae and legs with numerous quite short subdecumbent hairs, tarsi additionally with numerous coarse bristles. Body and appendages brownish-black.</paragraph>
|
||
<paragraph id="46633691FFF3591AFCF2FC26D3CEFB9C" blockId="2.[810,1452,158,1157]" box="[863,1186,1022,1048]" pageId="2" pageNumber="546">Gynes and males. Unknown.</paragraph>
|
||
<paragraph id="46633691FFF3591AFCF2FBFAD387FBE4" blockId="2.[810,1452,158,1157]" pageId="2" pageNumber="546">
|
||
<emphasis id="74A8EA83FFF3591AFCF2FBFAD362FBB8" bold="true" box="[863,1038,1058,1084]" pageId="2" pageNumber="546">Measurements</emphasis>
|
||
(in mm). HL 0.69, SL 0.49, OL 0.18, GL 0.23, ML 0.98, MH 0.43, HTL 0.74.
|
||
</paragraph>
|
||
<paragraph id="46633691FFF3591AFCF2FBB2D22AFB00" blockId="2.[810,1452,158,1157]" box="[863,1350,1130,1157]" pageId="2" pageNumber="546">Indices: SI 0.72, OI 0.26, GI 1.29, MI 2.27.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |