treatments-xml/data/7B/67/07/7B670705B95353228DE96C96DD49E7A3.xml

278 lines
30 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="B8AA3A52263377F296D9E603363C8959" ID-CLB-Dataset="307126" ID-DOI="10.3897/jhr.98.136200" ID-GBIF-Dataset="2f30ca71-e2ba-4645-9a2e-4f8c42de60c0" ID-ZooBank="77736D1C-30F9-4D6C-8310-3A04121CACE2" ID-publisher-id="136200" URI-arpha="CFF2E897-2633-5F4E-8369-35D589D56708" URI-zoobank="http://zoobank.org/77736D1C-30F9-4D6C-8310-3A04121CACE2" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1735917671054" checkinUser="pensoft" docAuthor="Wang, Zhen, Rasnitsyn, Alexandr P., Perkovsky, Evgeny E., Vilhelmsen, Lars &amp; Gao, Taiping" docDate="2025" docId="7B670705B95353228DE96C96DD49E7A3" docLanguage="en" docName="Journal of Hymenoptera Research 98: 1-18" docOrigin="Journal of Hymenoptera Research 98" docSource="https://jhr.pensoft.net/article/136200/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Siccibythus aristovi Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="18" masterDocId="CFF2E89726335F4E836935D589D56708" masterDocTitle="New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1735918735285" updateUser="ExternalLinkService">
<mods:mods id="5BC88A2F5F158DDFEB968871BC50114D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="83D2EABA1999E6B1210E78272E08EB35">
<mods:title id="0D2399C9F9642A3173045C4D4CD12749">New wasps of † Falsiformicidae from mid-Cretaceous Kachin amber</mods:title>
</mods:titleInfo>
<mods:name id="690A143DAA51B927C024B7045E6FEEDC" type="personal">
<mods:role id="795B99A322780F11ED7BC96219252AC2">
<mods:roleTerm id="6017B253FA180740581F8DE64C23A948">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E114E11E91587ED4009C9873D478CBCE">Wang, Zhen</mods:namePart>
<mods:nameIdentifier id="B14B775A93DE707D950935432CBE8C8B" type="ORCID">https://orcid.org/0009-0002-1981-8323</mods:nameIdentifier>
<mods:affiliation id="13D0D13DA87FA80686CE2375C7CE8223">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
</mods:name>
<mods:name id="70A037F8E184151829717320512FC413" type="personal">
<mods:role id="25AE45F8D05595C46FF4621FE012382E">
<mods:roleTerm id="AEEBF12AF57ECAAA3EA38DF22412DEA3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="663C4E4DA45127E96F5C746994FF273E">Rasnitsyn, Alexandr P.</mods:namePart>
<mods:affiliation id="8882B78D57AFFF1C969D07FEB97437EE">A. A. Borissiak Palaeontological Institute, Russian Academy of Sciences, Moscow, 117647, Russia</mods:affiliation>
</mods:name>
<mods:name id="E1414002E3B6B0104FFC1005A9FA51BE" type="personal">
<mods:role id="7A7144D5E65286FB58C8282610F92EBB">
<mods:roleTerm id="F2967C1356FB106807EE75C109C2488B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A72E187D1C1AD40FC279952B6B6174A1">Perkovsky, Evgeny E.</mods:namePart>
<mods:nameIdentifier id="B5FB78DE1AD9795BE42ED3ED3E122857" type="ORCID">0000-0002-7959-4379</mods:nameIdentifier>
<mods:affiliation id="F3D460AA4C6951513431886A547B0458">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
</mods:name>
<mods:name id="5BF0E70001F2230F65A13AA6441AFC25" type="personal">
<mods:role id="26DF2042A6B5E13154A55D0042718B9D">
<mods:roleTerm id="AC9B05E5FE289FB2A85B871FBEE8F1D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="717047321205BB31428B806DEB7E55A6">Vilhelmsen, Lars</mods:namePart>
<mods:nameIdentifier id="41A47D0AC292A3A36E30EAD7C8A80DC4" type="ORCID">0000-0002-5593-5722</mods:nameIdentifier>
<mods:affiliation id="0C10B1F231A225D60E2309A9B3080335">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Copenhagen, 2100, Denmark</mods:affiliation>
</mods:name>
<mods:name id="40224EF80F1560662580F3CAE3D64A89" type="personal">
<mods:role id="CFFA453323DAC9E7F38E7BA8CF209D30">
<mods:roleTerm id="9B87BAEF56A5CF78854D3A90CD4851B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5EB9AC8CBF79C060EE37856075CCEDB">Gao, Taiping</mods:namePart>
<mods:nameIdentifier id="2A892247B12189E897951B7230253ABD" type="ORCID">0000-0002-8118-8708</mods:nameIdentifier>
<mods:affiliation id="6370F4BFB850999B830969082E22549C">College of Life Sciences, Capital Normal University, Beijing, 100048, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="171E468349E109D04AEE1F757CB8FCDC">text</mods:typeOfResource>
<mods:relatedItem id="2FBE6BAFF581D6104C58615EB54FEC58" type="host">
<mods:titleInfo id="726C751BBBD9729EE464FA56632F8127">
<mods:title id="490B8D3B5FC7956A37F7C19AFF455463">Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part id="F5CB75F80C6B3D31EAA2427BB336819F">
<mods:date id="636BDFC943827128B9095D6D4D592605">2025</mods:date>
<mods:detail id="39C151A135DA64C5635647DCA17FA263" type="pubDate">
<mods:number id="9A31006A5DD8C48494902C356CAC988F">2025-01-03</mods:number>
</mods:detail>
<mods:detail id="EC054427725BF1C07BE1AED4B42471EE" type="volume">
<mods:number id="6CD84FFC8E7ADCFF9D24C7AF82637A08">98</mods:number>
</mods:detail>
<mods:extent id="CD8971B3B8D6FF44F647B3FAD6F63D70" unit="page">
<mods:start id="00676477CB96AC45F75D3E229842EA70">1</mods:start>
<mods:end id="818431DD1C3C989FF3623DD658A9E637">18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CE65B37A28E1F861A010FCF6CAAEA68C">journal article</mods:classification>
<mods:identifier id="768206A3FE910B8AF35A21A2D1EB252B" type="CLB-Dataset">307126</mods:identifier>
<mods:identifier id="2A442B0002D1474C2D47D8C83915EF58" type="DOI">10.3897/jhr.98.136200</mods:identifier>
<mods:identifier id="72779423488D8C3F25653FE83757E961" type="GBIF-Dataset">2f30ca71-e2ba-4645-9a2e-4f8c42de60c0</mods:identifier>
<mods:identifier id="465DAD4F9EEF02C902FF77E819B35AFB" type="ZooBank">77736D1C-30F9-4D6C-8310-3A04121CACE2</mods:identifier>
</mods:mods>
<treatment id="7B670705B95353228DE96C96DD49E7A3" ID-arpha="7B670705-B953-5322-8DE9-6C96DD49E7A3" ID-zoobank="https://zoobank.org/C0027405-25CC-48BB-AD21-8FCA112E21F0" LSID="urn:lsid:plazi:treatment:7B670705B95353228DE96C96DD49E7A3" LSID-ZBK="urn:lsid:zoobank.org:act:C0027405-25CC-48BB-AD21-8FCA112E21F0" httpUri="http://treatment.plazi.org/id/7B670705B95353228DE96C96DD49E7A3" scope_family="Falsiformicidae" scope_order="Hymenoptera">
<subSubSection id="CDBE9E3391390AFAEB15843DDDA73A92" type="nomenclature">
<paragraph id="FA1A7CFAAB8F0F7F3BEF763076FB84E8">
<taxonomicName id="1050EDADCE5205A07A0D36E3E1020B30" authority="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="FA1AEE3227921FA2492CAE606B70C362" italics="true">Siccibythus aristovi</emphasis>
Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen
</taxonomicName>
<taxonomicNameLabel id="58F2B1114EA1684BCE7C89F5004C0DD9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E34AE" type="diagnosis">
<paragraph id="C0CE1C096649597B1A870FB43EA8252B">
<heading id="A8CD3B84ABA4553E1BBFEEB9F0CC4FB3" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="07FAA2C035100AA8E51F3ED300979AEF">Fore wing with 2 Rs distal of r-rs at most about as long as r-rs; Rs + M and Cu both nebulous; A behind 1 cu-a not pigmented (except for very base). Fore tibial spur short, bifid apically. Basitarsus not longer than tarsomeres 24 combined. Pronotum short, propodeal horns long.</paragraph>
</subSubSection>
<subSubSection id="SECID0EB5AE" type="etymology">
<paragraph id="3A2C5B908FB64911ADD8E1A90900257F">
<heading id="B6F5FCD956CF3CCA74B523D4ADA1C60C" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="24B111D3FFC247AF0FA0F18D37312802">
The specific name “
<emphasis id="CD8525821D6801B228DE757893C3EA9B" italics="true">aristovi</emphasis>
” is dedicated to the memory of the late Dr. Daniil Sergeevich Aristov, for his outstanding contribution to palaeoentomology.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ER5AE" type="type material">
<paragraph id="6A67FDA9ABDC420344A33B47FB9800B7">
<heading id="6EAFD42892662108502502D88C7BD766" reason="title">Type material.</heading>
</paragraph>
<paragraph id="F7F3799DEA90C310DA25549ECC513411">
<materialsCitation id="1200163C3D7E144933CF76E1A0B175AB" country="Myanmar" location="Kachin" specimenCode="CNU-HYM-MA 2016214" specimenCount="1" specimenCount-female="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="23CBBD796B71D8420A68F1AA80BD494F" bold="true" italics="true">
<typeStatus id="ADD3EE3BA589341585273690D66BC93E" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="4B5B69BF926B2A2A2C0DA183C2A13F56" count="1" type="female">female</specimenCount>
. No.
<specimenCode id="DC0B7CDF0BA981550061EE7C1ADD9B2D">CNU-HYM-MA 2016214</specimenCode>
. A well-preserved and complete specimen (Figs
<figureCitation id="B1B2CF1FAB8FB3913B45C15FD2CDF6EC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A habitus in dorsal view B habitus in lateral view." figureDoi="10.3897/jhr.98.136200.figure5" httpUri="https://binary.pensoft.net/fig/1213368">5</figureCitation>
,
<figureCitation id="015120464B11CE1A2BE5A1D0FEAC3D51" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="FA85A54E913004BCE28E6258F95D15AE" ID-DOI="10.3897/jhr.98.136200.figure5" ID-arpha="23B8DC68-F7F1-5D46-94A4-B92BC5D5D76B" httpUri="https://binary.pensoft.net/fig/1213368" startId="F5">
<paragraph id="2094A10D15D8ABA20594E7C63D89A5D9">
<label id="2F9FD6E1D1F2039D3A1FBFF37EF08A15">Figure 5.</label>
</paragraph>
<paragraph id="72D34D4CCAF54A770BCA315645201E10">
Photographs of
<taxonomicName id="2D940FFD9D88738018770A13BFB808CD" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="256075F34C8B158D70D2F57AD857D652" italics="true">Siccibythus aristovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8B9CF498DBF7B61E1AE11B0A380D2920" rank="species">sp. nov.</taxonomicNameLabel>
(Female holotype; CNU-HYM-MA- 2016214)
<emphasis id="84C90237F123EA4100EBC9CD28A6EB2C" bold="true">A</emphasis>
habitus in dorsal view
<emphasis id="E1F4BB78E1D7ECFC0484AB3B746EF4D1" bold="true">B</emphasis>
habitus in lateral view.
</paragraph>
</caption>
<caption id="7438A2D4CF0C2481BC257CE76249A2CF" ID-DOI="10.3897/jhr.98.136200.figure6" ID-arpha="558F7053-A129-5650-99D8-8479B810E09A" httpUri="https://binary.pensoft.net/fig/1213369" startId="F6">
<paragraph id="858FBD0B7BFFA6769A4E7AF06EE056B7">
<label id="F8F22A0BC4DBB9E0D2FAC7DBDD2B6E9D">Figure 6.</label>
</paragraph>
<paragraph id="157B5187014E0A7695EAFB15800C85B6">
Photographs of
<taxonomicName id="AD2791CC4F375E78481B7CDB93F715B4" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="35413E489406873A430DCFDEF64EB1D6" italics="true">Siccibythus aristovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1EAC4F372FB775E203426194203FAA7" rank="species">sp. nov.</taxonomicNameLabel>
(Female holotype; CNU-HYM-MA- 2016214)
<emphasis id="D0C9DCB20006CC21AE41A34283AEE8E1" bold="true">A</emphasis>
head in lateral view
<emphasis id="2683DD1F1AC5B93FD9BD5FC7BD4EF748" bold="true">B</emphasis>
mouthparts
<emphasis id="2B3621D333F587F0E2F190A6866B2016" bold="true">C</emphasis>
mesosoma in lateral view, black arrow indicates the carinae
<emphasis id="C5171604F303287BA580EE75F5429487" bold="true">D</emphasis>
mesosoma in dorsal view
<emphasis id="7B449FD0ECCDC48167DC4A64B87BBE6F" bold="true">E</emphasis>
fore wing venation
<emphasis id="0554A827528FCBACEB09CAB27B80E1E5" bold="true">F</emphasis>
metasoma in lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EFBAG" type="type locality">
<paragraph id="5CF1C2D447366ACFDD2B0261A75E2A07">
<heading id="667E5F991882758B4B64B9EAA091B76B" reason="title">Type locality and horizon.</heading>
</paragraph>
<paragraph id="29B3A6E8E933494C0DB95B8640C87608">
The amber specimen was collected from
<collectingRegion id="00F0564B3DB959A9AACD7326A20DC0E2" country="Myanmar" name="Kachin">Kachin</collectingRegion>
, northern
<collectingCountry id="9F463E19AC482388677765E681184B11" name="Myanmar">Myanmar</collectingCountry>
, lowermost Cenomanian, mid-Cretaceous, which was dated at 98.79 ± 0.62 Ma (
<bibRefCitation id="33BE68BBF3224DF39C4D04C2306B41F4" DOI="10.1016/S1367-9120(02)00044-5" author="Cruickshank and Ko" firstAuthor="Cruickshank" journalOrPublisher="Journal of Asian Earth Sciences" pagination="441-455" refId="B3" refString="Cruickshank RD, Ko K (2003) Geology of an amber locality in the Hukawng Valley, Northern Myanmar. Journal of Asian Earth Sciences 21: 441455. https://doi.org/10.1016/S1367-9120 (02) 00044-5" title="Geology of an amber locality in the Hukawng Valley, Northern Myanmar." volume="21" year="2003">Cruickshank and Ko 2003</bibRefCitation>
;
<bibRefCitation id="6C482AF1AC389557F1209BE41BE8DAA4" DOI="10.1016/j.cretres.2012.03.014" author="Shi" etAl="et al." firstAuthor="Shi" journalOrPublisher="Cretaceous Research" pagination="155-163" refId="B25" refString="Shi G, Grimaldi DA, Harlow GE, Wang J, Wang J, Yang M, Lei W, Li Q, Li X (2012) Age constraint on Burmese amber based on U Pb dating of zircons. Cretaceous Research 37: 155163. https://doi.org/10.1016/j.cretres.2012.03.014" title="Age constraint on Burmese amber based on U - Pb dating of zircons." volume="37" year="2012">Shi et al. 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETBAG" type="description">
<paragraph id="19A98DEC328B2D4D63C61D80D237FDD6">
<heading id="1138E481BFF6647D54AC112F54D03B65" reason="title">Description.</heading>
</paragraph>
<paragraph id="02724523D63E8060355BE48887187CEA">
<emphasis id="BCDD5E2927D50971546CCC56744F35AA" bold="true">Female.</emphasis>
Body brown, metasoma dark brown. Head globose, with eyes bulging, occupying three-quarters of head in lateral view, almost reaching mandibular bases, occipital carina complete, reaching hypostomal carina well behind mandibular base, ocelli in small acute triangle well distant from eyes, interantennal space flat. Antenna narrow, with 12 antennomeres, each antennomere is approximately 12 times longer than wide, scape shorter than pedicel, pedicel longer than first flagellomere, flagellomeres slightly shortening gradually toward apex except for longer apical segment (Fig.
<figureCitation id="7902E25D3EB7E86E972CCA483AC4E087" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 A</figureCitation>
). Mandibles with three teeth, not overlapping; labrum not visible; maxillary palp short with six elongate palpomeres, labial palp short, with four slightly elongate to subquadrate palpomeres (Fig.
<figureCitation id="E6A459043219E2FE3E993BD1F8BC2374" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 B</figureCitation>
).
</paragraph>
<paragraph id="6D3793C7486B5C103E4440B072142D8E">
Pronotum with long dorsal part shorter than wide before lateral lobes, constricted near midlength, with fore declivitous part short, delimited by transverse groove posteriorly. Propleura long, slender, in lateral view, the length is 4 times its width, and abutting throughout. Mesoscutum with deep notauli widely spaced anteriorly, converging to almost meet posteriorly; transscutal articulation distinct; mesoscutellum rather short, wide apically, weakly convex. Meso- and metapleurae smooth. Metanotum short, with metascutellum very short and wide. Propodeum with dorsal and posterior surfaces meeting in angle, dorsal with longitudinal carina, and with transverse one midway between sublateral carinae, posterior surface with lateral longitudinal carinae, sublateral longitudinal carinae and supracoxal transverse carinae (Fig.
<figureCitation id="FE54A8E8338C4695A4E97926966F5579" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 C, D</figureCitation>
).
</paragraph>
<paragraph id="165E37707DCD62287E130078425F844D">
Fore wing basal veins (C, Sc + R, M + Cu, A, 1 Rs and 1 M), r-rs, short pterostigma, part of 2 Rs distal of r-rs and cu-a tubular, other veins nebulous. Pterostigma prominent, longer than wide; vein 1 Rs straight, shorter than 1 M, arise from Sc + R just proximal to pterostigma; 1 cu-a shorter than 1 M, parallel to 2 cu-a, placed slightly basally of 1 M; 1 Cu parallel to A; vein A extends distal up to 1 cu-a only; r-rs straight, arising beyond midlength of pterostigma; r-m not present (Fig.
<figureCitation id="53770044FADC31CCA21C326E9B60C2EC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 E</figureCitation>
). Hind wing with Sc + R; other veins not visible.
</paragraph>
<paragraph id="320039E7456960E2D4F61DC461BFE47A">Fore trochanter present, fore trochantellus ring-like; fore femur slightly swollen in middle, nearly three times as long as wide; fore tibia shorter than femur, calcar acute (not bifid); fore tarsus shorter than fore tibia, basitarsomere shorter than combined length of remaining tarsomeres. Mid trochanter present, mid trochantellus ring-like; mid femur slightly swollen in middle, four times as long as wide; mid tibia nearly as long as femur, six times as long as wide; mid tarsus shorter than mid tibia, basitarsomere shorter than combined length remaining tarsomeres. Hind trochanter short, bulging dorsally, hind trochantellus ring-like; hind femur slightly swollen in middle, nearly four times as long as wide; hind tibia slender than femur, longer than femur; hind tarsus nearly as long as hind tibia, remaining tarsomeres combined shorter than basitarsomere. Claw simple, arolium small. Tibial spur formula 1 / 2 / 2.</paragraph>
<paragraph id="06A1C006A8F41699B95069D266C10698">
First metasomal segment short, tergum node-like, triangular in lateral view, with anterior surface concave, second segment slightly longer and higher than following, following gradually smaller, apical (sixth) sternum high, apparently folded, with upper sides and apical orifice apparently not specialized, tergum 7 as visible (in lateral view) small but more or less sclerotized, with spiracle not enlarged (Fig.
<figureCitation id="51A5402712162F471B21C9208AA9D599" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of Siccibythus aristovi sp. nov. (Female holotype; CNU-HYM-MA- 2016214) A head in lateral view B mouthparts C mesosoma in lateral view, black arrow indicates the carinae D mesosoma in dorsal view E fore wing venation F metasoma in lateral view." figureDoi="10.3897/jhr.98.136200.figure6" httpUri="https://binary.pensoft.net/fig/1213369">6 F</figureCitation>
). Sting straight, apparently short, without teeth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWCAG" type="description">
<paragraph id="0196B03107F33418C9477AEB5EB3B9D9">
<heading id="142FD024AC6C4A251910F2AA8964D328" reason="title">Measurements (mm).</heading>
</paragraph>
<paragraph id="AF2F1D01F00804FDB31C7EFAFDD1B2A1">
<typeStatus id="60534AEFF89EB0922C76B9B52C35ABCF" type="holotype">Holotype</typeStatus>
female. Total body length (excluding sting and antenna) 4.14; head length 0.94, width 0.62, height 0.48; scape length 0.20, pedicel length 0.19, the first flagellum length 0.14, total flagellum length 1.11; fore wing length 1.95; hind wing length 1.20; fore coxa length 0.27, fore trochantellus 0.06, fore femur length 0.64, fore tibia length 0.49, fore tarsal length 0.64; mid coxa length 0.25, mid trochanter length 0.10, mid trochantellus 0.04, mid femur length 0.60, mid tibia length 0.64, mid tarsal length 0.39; hind coxa length 0.23, hind trochanter length 0.10, hind trochantellus 0.06, hind femur length 0.69, hind tibia length 0.58, hind tarsal length 0.79; the first metasomal segment length 0.29, metasoma length 1.66.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2CAG" type="remarks">
<paragraph id="1E9A11545C8FB4751359AE9A4EC2CACC">
<heading id="6D8784AFA07B709724D4369DF475B554" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="92EFCD66B272E4B2BBE5DF2DC33034E2">
The new species is classified in †
<taxonomicName id="16512B109E6E596E1A35960B6ACC879B" authority="Cockx &amp; McKellar, 2016" authorityName="Cockx &amp; McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">
<emphasis id="48716356E4F83DF8B0D83D6AE9996A0D" italics="true">Siccibythus</emphasis>
<bibRefCitation id="AA5EF34503A8AF3912AD19A31CC9BBC3" DOI="10.1016/j.cretres.2016.06.015" author="Cockx PFD &amp; McKellar RC" journalOrPublisher="Cretaceous Research" pagination="133-139" refId="B2" refString="Cockx PFD, McKellar RC (2016) First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar. Cretaceous Research 67: 133 - 139. https: // doi. org / 10.1016 / j. cretres. 2016.06. 015" title="First record of the family Scolebythidae (Hymenoptera) in mid-Cretaceous amber from Myanmar." volume="67" year="2016">Cockx &amp; McKellar, 2016</bibRefCitation>
</taxonomicName>
based on the fore wing with no cells closed with tubular veins in its middle and distal part (
<bibRefCitation id="EE75EF1F9B60707724DA9EF51B127BC3" DOI="10.11646/palaeoentomology.3.6.10" author="Rasnitsyn" etAl="et al." firstAuthor="Rasnitsyn" journalOrPublisher="Palaeoentomology" pagination="582-596" refId="B20" refString="Rasnitsyn AP, Zhang Q, Müller P, Zhang HC (2020) On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.). Palaeoentomology 3: 582596. https://doi.org/10.11646/palaeoentomology.3.6.10" title="On the identity and limits of Falsiformicidae (Insecta: Hymenoptera, Vespoidea s. l.)." volume="3" year="2020">Rasnitsyn et al. 2020</bibRefCitation>
). It is differentiated from
<taxonomicName id="4798421EFC510891D4BD29B9C78AB2D4" authorityName="Cockx &amp; McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
<emphasis id="7A2554C75ABE2D5BB364513CED4F6709" italics="true">Siccibythus musculosus</emphasis>
</taxonomicName>
,
<taxonomicName id="5AAB368797027804C4E381A761DD908D" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
<emphasis id="6AEBB685155A2105F4974B9D84E43374" italics="true">S. paulus</emphasis>
</taxonomicName>
,
<taxonomicName id="8364D6CC3400DA567C018B610706C77A" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
<emphasis id="2DB0BDC3C322A1C64FBA888AF917BFD9" italics="true">S. ohmkuhnlei</emphasis>
</taxonomicName>
and
<taxonomicName id="2674496A124A6E51EAEF368ABC9BB4EF" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="EF21A7EF736265A1ED9694D1757217CF" italics="true">S. robustus</emphasis>
</taxonomicName>
by fore wing with 2 Rs distal of r-rs shorter than r-rs, basitarsus not longer than tarsomeres 24 combined (vs. 2 Rs distal of r-rs several times as long as r-rs, basitarsus longer than tarsomeres 24 combined in
<taxonomicName id="65C1CC51DAED0057F54CF782C2E810EA" authorityName="Cockx &amp; McKellar" authorityYear="2016" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="musculosus">
<emphasis id="7837D722073DB4384EAB954D1FC84962" italics="true">S. musculosus</emphasis>
</taxonomicName>
,
<taxonomicName id="59C5BAFE0F1C29A968A0CE3440BA5E4D" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="paulus">
<emphasis id="7435FCD0AF5512462A0E3ECC85CB2396" italics="true">S. paulus</emphasis>
</taxonomicName>
,
<taxonomicName id="589A379A0616AC48DF96FA8817481D35" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ohmkuhnlei">
<emphasis id="7C83623B986214A6E899608E9F7721C0" italics="true">S. ohmkuhnlei</emphasis>
</taxonomicName>
and
<taxonomicName id="B93DEA4D1374D2C73BC09AF0440DC1BB" authorityName="Wang, Rasnitsyn, Perkovsky &amp; Vilhelmsen" authorityYear="2025" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="D5EC16E9A650603FE6A9E58218D2001A" italics="true">S. robustus</emphasis>
</taxonomicName>
). It is distinguished from
<taxonomicName id="676BF01993866DCD134526AA0906A9AB" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
<emphasis id="1628283EE7DE6D9B6415594625812700" italics="true">S. martynovae</emphasis>
</taxonomicName>
by fore wing veins Rs + M and Cu both being nebulous (vs. Rs + M and Cu are both tubular in
<taxonomicName id="63EBCCE5986DFDFB3C426BE37D504C7C" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="martynovae">
<emphasis id="C38EF60D9B5F86D7203068C5C5391B10" italics="true">S. martynovae</emphasis>
</taxonomicName>
). It is differentiated from
<taxonomicName id="2BA0E64431454C6FD5CD930B126BDC5D" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="pallidus">
<emphasis id="F0937FD25FCB9F255C0C0A9A53CC5F57" italics="true">S. pallidus</emphasis>
</taxonomicName>
by the following characters: 1) fore wing with 2 A not pigmented (except for very base) (vs. fore wing with 2 A pigmented); 2) pronotum constricted in the middle (vs. not constricted). It is distinguished from
<taxonomicName id="F883EF78E486B0F14ED8BCB03925DBA5" authorityName="Rasnitsyn, Zhang, Muller &amp; Zhang" authorityYear="2020" class="Insecta" family="Scolebythidae" genus="Siccibythus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="F552A4A1C3F59475FC5CA718388AF49C" italics="true">S. oculatus</emphasis>
</taxonomicName>
by pronotum short, propodeal horns long (vs. pronotum long, propodeal horns short).
</paragraph>
</subSubSection>
</treatment>
</document>