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<document id="28970B7F8E8529A6BF8BE4453D5AA2CA" ID-DOI="10.59893/abud.24(2).010" ID-ISSN="1407-8953" ID-Zenodo-Dep="14659857" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736974727826" checkinUser="felipe" docAuthor="Anichtchenko, Alexander" docDate="2024" docId="054E5D3A1173FFC8FF55FAD3113D2BE7" docLanguage="en" docName="ActaBiolUnivDaugavp.24.2.259-264.pdf" docOrigin="Acta Biologica Universitatis Daugavpiliensis 24 (2)" docSource="https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/" docStyle="DocumentStyle:DD7D35B4ACFEF3C20AEB77B8656EEADF.5:ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleId="DD7D35B4ACFEF3C20AEB77B8656EEADF" docStyleName="ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleVersion="5" docTitle="Amphimenes hartmanni Anichtchenko, 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="263" masterDocId="F97725421171FFCCFFC2FFDE15632B76" masterDocTitle="New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia" masterLastPageNumber="264" masterPageNumber="259" pageNumber="261" updateTime="1736975295492" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EDFC8A1C5505024157A63F1BF05EBB42">New Species Of Genus Amphimenes Bates, 1873 (Coleoptera, Carabidae, Lebiinae) From Laos And Malaysia</mods:title>
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<mods:name id="CF2E9AEE890C51E30C3E6A65312D5AFA" type="personal">
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<mods:namePart id="FD8B1E81F69FDAEDD6529408E074869B">Anichtchenko, Alexander</mods:namePart>
<mods:affiliation id="A94F081CD6BC33AD02A535AA396A9497">Corresponding author: Alexander Anichtchenko. Institute of Life Sciences and Technologies, Daugavpils University, Vienibas Str. 13, LV- 5401, Daugavpils, Latvia. E-mail: alexander. anishchenko @ du. lv. ORCID: https: // orcid. org / 0000 - 0001 - 8512 - 830 X</mods:affiliation>
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<mods:title id="DD99B08D3CE1FD263E0DA22475314045">Acta Biologica Universitatis Daugavpiliensis</mods:title>
</mods:titleInfo>
<mods:part id="7DC755F4B700811C08D3DE75384C78CA">
<mods:date id="CFB8E8D68130FA69F3C4CCABFCB53A25">2024</mods:date>
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<mods:number id="B31374E86DD269437086BC166D5F18E1">2024-12-30</mods:number>
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<mods:number id="76FEA205009747E8C0D4CC625D4D664C">24</mods:number>
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<mods:number id="7D63363DDD6F072DA03107E16F2CC682">2</mods:number>
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<mods:url id="6D1C2591D08A73BF42363436F51F9AB4">https://du.lv/en/research/institutes/institute-of-life-sciences-and-technologies/issued-journal/acta-biologica-universitatis-daugavpiliensis/acta-biologica-universitatis-daugavpiliensis-archives/acta-biologica-universitatis-daugavpiliensis-volume-24-no-2-2024/new-species-of-genus-amphimenes-bates-1873-coleoptera-carabidae-lebiinae-from-laos-and-malaysia/</mods:url>
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<mods:classification id="B60B2DCAF7299ACB88B1B3D95A3A8E79">journal article</mods:classification>
<mods:identifier id="07D860DBBCE6D4EF798A9C3CCFC74AA7" type="DOI">10.59893/abud.24(2).010</mods:identifier>
<mods:identifier id="8E4AA0E469FA591E1AFD799B092D5817" type="ISSN">1407-8953</mods:identifier>
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<treatment id="054E5D3A1173FFC8FF55FAD3113D2BE7" LSID="urn:lsid:plazi:treatment:054E5D3A1173FFC8FF55FAD3113D2BE7" httpUri="http://treatment.plazi.org/id/054E5D3A1173FFC8FF55FAD3113D2BE7" lastPageId="4" lastPageNumber="263" pageId="2" pageNumber="261">
<subSubSection id="C5FDBFA71173FFCEFF55FAD314442E30" pageId="2" pageNumber="261" type="nomenclature">
<paragraph id="8D58EC2C1173FFCEFF55FAD3175A2E50" blockId="2.[151,569,1293,1350]" box="[151,569,1293,1318]" pageId="2" pageNumber="261">
<heading id="D6105B401173FFCEFF55FAD3175A2E50" bold="true" box="[151,569,1293,1318]" fontSize="10" level="2" pageId="2" pageNumber="261" reason="5">
<emphasis id="BF93303E1173FFCEFF55FAD3175A2E50" bold="true" box="[151,569,1293,1318]" pageId="2" pageNumber="261">
<taxonomicName id="4AE797AF1173FFCEFF55FAD314952E53" authorityName="Anichtchenko" authorityYear="2024" box="[151,502,1293,1317]" class="Insecta" family="Carabidae" genus="Amphimenes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="hartmanni" status="sp. nov.">
<emphasis id="BF93303E1173FFCEFF55FAD314442E53" bold="true" box="[151,295,1293,1317]" italics="true" pageId="2" pageNumber="261">Amphimenes</emphasis>
(s. str.)
<emphasis id="BF93303E1173FFCEFE43FAD314952E53" bold="true" box="[385,502,1293,1317]" italics="true" pageId="2" pageNumber="261">hartmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4A08D451173FFCEFE3FFAD0175A2E50" box="[509,569,1294,1318]" pageId="2" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8D58EC2C1173FFCEFF55FAF014442E30" blockId="2.[151,569,1293,1350]" box="[151,295,1326,1350]" pageId="2" pageNumber="261">(Figs. 2, 5, 6)</paragraph>
</subSubSection>
<subSubSection id="C5FDBFA71173FFCEFF55FAB014EB2D10" pageId="2" pageNumber="261" type="materials_examined">
<paragraph id="8D58EC2C1173FFCEFF55FAB014EB2D10" blockId="2.[151,646,1390,1670]" pageId="2" pageNumber="261">
<emphasis id="BF93303E1173FFCEFF55FAB014662EF0" bold="true" box="[151,261,1390,1414]" pageId="2" pageNumber="261">Material.</emphasis>
<materialsCitation id="3D8FE6711173FFCEFECDFAB014EB2D10" collectingDate="2011-05-31" collectionCode="NME" country="Laos" elevation="1600" latitude="20.2" location="Phou Pan" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" pageId="2" pageNumber="261" specimenCount="2" stateProvince="Houaphan" typeStatus="holotype">
<typeStatus id="525C528E1173FFCEFECDFAB0141A2EF0" box="[271,377,1390,1414]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
, male: “NE –
<collectingCountry id="F5F0ACBC1173FFCEFDE3FAB0173A2EF3" box="[545,601,1390,1413]" name="Laos" pageId="2" pageNumber="261">Laos</collectingCountry>
, Pr.
<collectingRegion id="4F2322CE1173FFCEFF55FA5014652ED0" box="[151,262,1422,1446]" country="Laos" name="Houaphan" pageId="2" pageNumber="261">Hua Phan</collectingRegion>
/
<collectingMunicipality id="6D3C76561173FFCEFEE3FA5014FC2ED0" box="[289,415,1422,1446]" pageId="2" pageNumber="261">Ban Saleui</collectingMunicipality>
,
<location id="8838BAF71173FFCEFE6FFA50177B2ED3" LSID="urn:lsid:plazi:treatment:054E5D3A1173FFC8FF55FAD3113D2BE7:8838BAF71173FFCEFE6FFA50177B2ED3" box="[429,536,1422,1446]" country="Laos" latitude="20.2" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleui" name="Phou Pan" pageId="2" pageNumber="261" stateProvince="Houaphan">Phou Pan</location>
(Mt.) / ~
<geoCoordinate id="E8D38AEB1173FFCEFF55FA70158E2EB0" box="[151,237,1454,1478]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="261" precision="925" value="20.2">20°12N</geoCoordinate>
,
<geoCoordinate id="E8D38AEB1173FFCEFF3BFA7014342EB0" box="[249,343,1454,1478]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="2" pageNumber="261" precision="925" value="104.01667">104°01E</geoCoordinate>
/
<date id="F959CAEC1173FFCEFEAFFA7014B82EB0" box="[365,475,1454,1478]" pageId="2" pageNumber="261" value="2011-05-31">
<collectingDate id="E91D33041173FFCEFEAFFA7014B82EB0" box="[365,475,1454,1478]" pageId="2" pageNumber="261" value="2011-05-31">31.V.2011</collectingDate>
</date>
,
<quantity id="4A1F41C91173FFCEFE25FA7017112EB0" box="[487,626,1454,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="2" pageNumber="261" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">
<elevation id="06CA0B1F1173FFCEFE25FA7017112EB0" box="[487,626,1454,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="2" pageNumber="261" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">1300-1900m</elevation>
</quantity>
, / local collector”, “collection / NATURKUNDE -/ MUSEUM ERFURT [yellow label]”, “
<typeStatus id="525C528E1173FFCEFF63F9D014692D50" box="[161,266,1550,1574]" pageId="2" pageNumber="261" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4AE797AF1173FFCEFEE6F9D014D42D50" authorityName="BATES" authorityYear="1873" box="[292,439,1550,1574]" class="Insecta" family="Carabidae" genus="Amphimenes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="genus">Amphimenes</taxonomicName>
(s.str.) / hartmanni
<taxonomicNameLabel id="A4A08D451173FFCEFF76F9F0158B2D33" box="[180,232,1582,1605]" pageId="2" pageNumber="261" rank="species">sp.n.</taxonomicNameLabel>
/ des. Anichtchenko A. 24 [red label, handwritten]” (
<collectionCode id="EBF674E91173FFCEFEF9F990141F2D10" box="[315,380,1614,1638]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="2" pageNumber="261">NME</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5FDBFA71173FFCEFF55F9B016882D50" pageId="2" pageNumber="261" type="diagnosis">
<paragraph id="8D58EC2C1173FFCEFF55F9B017E62DF0" blockId="2.[151,646,1390,1670]" box="[151,645,1646,1670]" pageId="2" pageNumber="261">
<emphasis id="BF93303E1173FFCEFF55F9B014732DF0" bold="true" box="[151,272,1646,1670]" pageId="2" pageNumber="261">Diagnosis.</emphasis>
This new species belongs to the
</paragraph>
<paragraph id="8D58EC2C1173FFCEFD67FAD016882D50" blockId="2.[677,1173,1294,1670]" pageId="2" pageNumber="261">
“
<emphasis id="BF93303E1173FFCEFD73FAD016682E50" box="[689,779,1294,1318]" italics="true" pageId="2" pageNumber="261">piceolus</emphasis>
”-group sensu
<bibRefCitation id="E97691DD1173FFCEFC69FAD011112E50" author="Fedorenko D. N." box="[939,1138,1294,1318]" pageId="2" pageNumber="261" pagination="233 - 250" refId="ref2557" refString="Fedorenko D. N. 2019. A revision of Amphimenes (Coleoptera: Carabidae: Lebiini). Russian Entomological Journal 28 (3): 233 - 250." type="journal article" year="2019">Fedorenko (2019)</bibRefCitation>
. In general appearance, by the reduced hind-wings and the shape of aedeagus it is closely related to
<taxonomicName id="4AE797AF1173FFCEFD67FAB0116B2EF0" authority="Nakane, 1957" authorityName="Nakane" authorityYear="1957" box="[677,1032,1390,1414]" class="Insecta" family="Carabidae" genus="Amphimenes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="asahinai">
<emphasis id="BF93303E1173FFCEFD67FAB017DF2EF3" box="[677,700,1390,1413]" italics="true" pageId="2" pageNumber="261">A.</emphasis>
(s. str.)
<emphasis id="BF93303E1173FFCEFCD2FAB016082EF0" box="[784,875,1390,1414]" italics="true" pageId="2" pageNumber="261">asahinai</emphasis>
Nakane, 1957
</taxonomicName>
, known from
<collectingCountry id="F5F0ACBC1173FFCEFD67FA50179A2ED0" box="[677,761,1422,1446]" name="Taiwan" pageId="2" pageNumber="261">Taiwan</collectingCountry>
, from which it can be distinguished by the small size, the narrow lateral margins of pronotum and the shape of median lobe of aedeagus, curved before the middle in lateral view, and wide in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C5FDBFA71173FFCFFD67F9F011312833" lastPageId="3" lastPageNumber="262" pageId="2" pageNumber="261" type="description">
<paragraph id="8D58EC2C1173FFCEFD67F9F011F02DF0" blockId="2.[677,1173,1294,1670]" pageId="2" pageNumber="261">
<emphasis id="BF93303E1173FFCEFD67F9F016562D30" bold="true" box="[677,821,1582,1606]" pageId="2" pageNumber="261">Description.</emphasis>
Body length
<quantity id="4A1F41C91173FFCEFC16F9F0114D2D33" box="[980,1070,1582,1606]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="261" unit="mm" value="4.5">4.5 mm</quantity>
. Dorsum brown, mouthparts, legs and antennae light brown, clypeus and labrum brown (Fig. 2).
</paragraph>
<paragraph id="8D58EC2C1173FFCEFBAEF91A11F72DAA" blockId="2.[1132,1172,1732,1756]" box="[1132,1172,1732,1756]" pageId="2" pageNumber="261">261</paragraph>
<paragraph id="8D58EC2C1172FFCFFDC5FFA017BB2BE7" blockId="3.[519,728,126,145]" box="[519,728,126,145]" pageId="3" pageNumber="262">
<emphasis id="BF93303E1172FFCFFDC5FFA017BB2BE7" box="[519,728,126,145]" italics="true" pageId="3" pageNumber="262">Alexander Anichtchenko</emphasis>
</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3FF10174F2B90" blockId="3.[113,609,206,1062]" box="[113,556,206,230]" pageId="3" pageNumber="262">Gula, trochanters and coxa brownish-red.</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3FF3014082AB0" blockId="3.[113,609,206,1062]" pageId="3" pageNumber="262">Eyes flattened, temples about half as long as eye; posterior supraorbital seta situated in distance equal to radius of eye back. Labrum slightly incised. Frontal foveae shallow. Microreticulation consists of polygonal meshes. Antennae long, surpassing base of pronotum by two and half last joints.</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3FE1014B828B0" blockId="3.[113,609,206,1062]" pageId="3" pageNumber="262">Pronotum 1.25 times as wide as long, 1.42 times as wide as head, with front angles slightly protruding and rounded apically; sides slightly angulate in front of lateral pore, weakly sinuate before hind angles; not reflexed between lateral pore and anterior angle, very slightly reflexed towards hind angles; lateral margin narrow, gradually expanding from anterior angles towards hind angles. Hind angles obtuse. Base of pronotum slightly protruding backwards. Mid-line reaching anterior border, superficial on the disc. Basal foveae flat and indistinct. Microreticulation on the disc very strong, almost isodiametric, towards the sides meshes becomes stronger, and towards hind angles meshes turns into small tubercles.</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3FC1016412A90" blockId="3.[113,609,206,1062]" lastBlockId="3.[633,1135,206,1030]" pageId="3" pageNumber="262">Elytra oval, 1.32 times as long as wide, 1.66 times as wide as pronotum, broadest at about middle, with shoulders strongly rounded; not fused along midline; apical truncature moderately sinuate between a rounded outer angle and a pointed apex. Three discal setigerous pores: D1/EL=0.32, D2/EL=0.78 and D3/EL=0.94. All elytral striae deep, gently punctate, intervals moderately transversally striate, on the disc of elytra transverse striation becomes weak. Microreticulation strongly transverse. Hind wings reduced.</paragraph>
<paragraph id="8D58EC2C1172FFCFFD42FE3016A429D0" blockId="3.[633,1135,206,1030]" pageId="3" pageNumber="262">Ventral segments smooth, impunctate. Metepisternum short, subsquare. Last metatarsomere with two distal pairs of ventral setae. Claws with four denticles, basal one much smaller. Basal third of male profemur with obtuse and wide ventral tubercle.</paragraph>
<paragraph id="8D58EC2C1172FFCFFD42FD7011312833" blockId="3.[633,1135,206,1030]" pageId="3" pageNumber="262">Aedeagus (Figs. 5-6): median lobe elongate, in lateral view evenly rounded dorsally and bend- ed at an obtuse angle before the middle ventrally; apical lamella elongate, with a rounded tip. Endophallus without distinct structures.</paragraph>
</subSubSection>
<subSubSection id="C5FDBFA71172FFCFFD42FC90165F28B0" pageId="3" pageNumber="262" type="etymology">
<paragraph id="8D58EC2C1172FFCFFD42FC90165F28B0" blockId="3.[633,1135,206,1030]" pageId="3" pageNumber="262">
<emphasis id="BF93303E1172FFCFFD42FC9016632810" bold="true" box="[640,768,846,870]" pageId="3" pageNumber="262">Etymology.</emphasis>
This new species is named for Matthias Hartmann,
<taxonomicName id="4AE797AF1172FFCFFC99FCB016B328F0" authorityName="Linnaeus" authorityYear="1758" box="[859,976,878,902]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="3" pageNumber="262" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
specialist, and Managing Director at Naturkundemuseum Erfurt (
<collectingCountry id="F5F0ACBC1172FFCFFD05FC70165228B0" box="[711,817,942,966]" name="Germany" pageId="3" pageNumber="262">Germany</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C5FDBFA71172FFC8FD42FC10113D2BE7" lastPageId="4" lastPageNumber="263" pageId="3" pageNumber="262" type="distribution">
<paragraph id="8D58EC2C1172FFCFFD42FC1017B52F73" blockId="3.[633,1135,206,1030]" pageId="3" pageNumber="262">
<emphasis id="BF93303E1172FFCFFD42FC1016712890" bold="true" box="[640,786,974,998]" pageId="3" pageNumber="262">Distribution.</emphasis>
Known only from
<typeStatus id="525C528E1172FFCFFC25FC1011762890" box="[999,1045,974,998]" pageId="3" pageNumber="262">type</typeStatus>
locality in
<collectingCountry id="F5F0ACBC1172FFCFFD59FC3017B12F73" box="[667,722,1006,1029]" name="Laos" pageId="3" pageNumber="262">Laos</collectingCountry>
.
</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3F99A16CA2D0B" blockId="3.[113,1134,1604,1661]" pageId="3" pageNumber="262">
<emphasis id="BF93303E1172FFCFFFB3F99A15942D2A" bold="true" box="[113,247,1604,1628]" pageId="3" pageNumber="262">Figures 3-6.</emphasis>
Aedeagus structure of
<taxonomicName id="4AE797AF1172FFCFFE28F99B17112D2B" authorityName="BATES" authorityYear="1873" box="[490,626,1605,1629]" class="Insecta" family="Carabidae" genus="Amphimenes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="BF93303E1172FFCFFE28F99B17112D2B" box="[490,626,1605,1629]" italics="true" pageId="3" pageNumber="262">Amphimenes</emphasis>
</taxonomicName>
in lateral and dorsal view. 3-4 –
<taxonomicName id="4AE797AF1172FFCFFC0CF99B15F12D0A" authorityName="Anichtchenko" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amphimenes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="kmecoi" status="sp. n.">
<emphasis id="BF93303E1172FFCFFC0CF99B16862D2A" box="[974,997,1605,1628]" italics="true" pageId="3" pageNumber="262">A.</emphasis>
(s. str.)
<emphasis id="BF93303E1172FFCFFBF8F99B15F12D0A" italics="true" pageId="3" pageNumber="262">kmecoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4A08D451172FFCFFF5BF9BB15B12D0A" box="[153,210,1637,1660]" pageId="3" pageNumber="262" rank="species">sp. n.</taxonomicNameLabel>
5-6 –
<taxonomicName id="4AE797AF1172FFCFFED5F9BB14902D0B" authorityName="Anichtchenko" authorityYear="2024" box="[279,499,1637,1661]" class="Insecta" family="Carabidae" genus="Amphimenes" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="hartmanni" status="sp. n.">
<emphasis id="BF93303E1172FFCFFED5F9BB144D2D0A" box="[279,302,1637,1660]" italics="true" pageId="3" pageNumber="262">A.</emphasis>
(s. str.)
<emphasis id="BF93303E1172FFCFFE46F9BB14902D0B" box="[388,499,1637,1661]" italics="true" pageId="3" pageNumber="262">hartmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4A08D451172FFCFFE38F9BB17502D0A" box="[506,563,1637,1660]" pageId="3" pageNumber="262" rank="species">sp. n.</taxonomicNameLabel>
Images courtesy A. Anichtchenko.
</paragraph>
<paragraph id="8D58EC2C1172FFCFFFB3F91A15FA2DAA" blockId="3.[113,153,1732,1756]" box="[113,153,1732,1756]" pageId="3" pageNumber="262">262</paragraph>
<paragraph id="8D58EC2C1175FFC8FF0FFFA1113D2BE7" blockId="4.[205,1118,126,145]" box="[205,1118,126,145]" pageId="4" pageNumber="263">
<emphasis id="BF93303E1175FFC8FF0FFFA1113D2BE7" box="[205,1118,126,145]" italics="true" pageId="4" pageNumber="263">
New species of genus
<taxonomicName id="4AE797AF1175FFC8FE45FFA0173C2BE7" authority="Bates, 1873" authorityName="BATES" authorityYear="1873" box="[391,607,126,145]" class="Insecta" family="Carabidae" genus="Amphimenes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="263" phylum="Arthropoda" rank="genus">Amphimenes Bates, 1873</taxonomicName>
(
<taxonomicName id="4AE797AF1175FFC8FDA8FFA017AE2BE7" authorityName="Linnaeus" authorityYear="1758" box="[618,717,126,145]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="263" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
, carabidae,
<taxonomicName id="4AE797AF1175FFC8FCF5FFA016E02BE7" authorityName="Bonelli" authorityYear="1810" box="[823,899,126,145]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Lebiinae">Lebiinae</taxonomicName>
) from
<collectingCountry id="F5F0ACBC1175FFC8FC79FFA116872BE7" box="[955,996,127,145]" name="Laos" pageId="4" pageNumber="263">Laos</collectingCountry>
and
<collectingCountry id="F5F0ACBC1175FFC8FBCDFFA0113D2BE7" box="[1039,1118,126,145]" name="Malaysia" pageId="4" pageNumber="263">Malaysia</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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