696 lines
147 KiB
XML
696 lines
147 KiB
XML
<document id="8F2D114E182684861AB16362B01DFC1E" ID-DOI="10.5281/zenodo.280919" ID-GBIF-Dataset="5ae7d78b-18a9-4af9-b594-ea3d269e20c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="280919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460477598879" checkinUser="plazi" docAuthor="West, Rick C., Nunn, Steven C. & Hogg, Stephen" docDate="2012" docId="DE0287A9FF9B556FFF4A70E0FBC9479A" docLanguage="en" docName="zt03299p043.pdf" docOrigin="Zootaxa 3299" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Selenocosmia lyra West, Nunn & Hogg, 2012, gen. nov." docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="223BFFD1FF985569FFDD736BFFF94556" masterDocTitle="A new tarantula genus, Psednocnemis, from West Malaysia (Araneae: Theraphosidae), with cladistic analyses and biogeography of Selenocosmiinae Simon 1889" masterLastPageNumber="43" masterPageNumber="1" pageNumber="4" updateTime="1698307071297" updateUser="plazi">
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<mods:title id="521B136FF3024D717D65ABE913E4BDFE">A new tarantula genus, Psednocnemis, from West Malaysia (Araneae: Theraphosidae), with cladistic analyses and biogeography of Selenocosmiinae Simon 1889</mods:title>
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<mods:namePart id="8F23137169B0D56A64889CAEB933FCA6">West, Rick C.</mods:namePart>
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<mods:namePart id="AD1949B4DFBAE91048FEAEE95A3E2DF7">Nunn, Steven C.</mods:namePart>
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<mods:namePart id="748BA647FC11180D49A66E8E91CA90AF">Hogg, Stephen</mods:namePart>
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<mods:date id="5215D4C7DBFCF80C0A7515FB614670E5">2012</mods:date>
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<mods:number id="ABE65D2DA4F48AA32D7B49EE1899084E">3299</mods:number>
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<treatment id="DE0287A9FF9B556FFF4A70E0FBC9479A" ID-DOI="http://doi.org/10.5281/zenodo.6168296" ID-GBIF-Taxon="123201047" ID-Zenodo-Dep="6168296" LSID="urn:lsid:plazi:treatment:DE0287A9FF9B556FFF4A70E0FBC9479A" httpUri="http://treatment.plazi.org/id/DE0287A9FF9B556FFF4A70E0FBC9479A" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
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<subSubSection id="1EB16534FF9B556FFF4A70E0FBC9479A" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="561436BFFF9B556AFF4A70E0FF2146F5" blockId="3.[151,1436,907,1867]" box="[151,216,907,931]" pageId="3" pageNumber="4">
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<heading id="0D5C81D3FF9B556AFF4A70E0FF2146F5" bold="true" box="[151,216,907,931]" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
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<emphasis id="64DFEAADFF9B556AFF4A70E0FF2146F5" bold="true" box="[151,216,907,931]" pageId="3" pageNumber="4">
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<taxonomicName id="91AB4D3CFF9B556AFF4A70E0FF2D46F5" box="[151,212,907,931]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
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:
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="561436BFFF9B556AFF4A70C5FDAF437C" blockId="3.[151,1436,907,1867]" pageId="3" pageNumber="4">
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5. Female
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<taxonomicName id="91AB4D3CFF9B556AFEFC70C5FEA84690" box="[289,337,942,966]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
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: (0) rudimentary (
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<figureCitation id="CE902A3AFF9B556AFDC170C5FD944690" box="[540,621,942,966]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="3" pageNumber="4">Fig. 47</figureCitation>
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) or absent; (1) oval with proximally truncate form (
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<figureCitation id="CE902A3AFF9B556AFB6970C5FAED4690" box="[1204,1300,942,966]" captionStart="FIGURES 49 a – 50 b. 49 a, b" captionStartId="27.[151,264,1127,1150]" captionTargetBox="[151,1436,193,1104]" captionTargetId="figure@27.[151,1436,193,1106]" captionTargetPageId="27" captionText="FIGURES 49 a – 50 b. 49 a, b. Haplocosmia himalayana (Pocock 1899), holotype female, NHM 1899.8. 19.3. 49 a. Right maxillae (inverted), prolateral view. 49 b. Right chelicerae, prolateral view. 50 a, b. Chilobrachys sericeus (Thorell 1895), syntype male, NHM 1895.9. 21.21. 50 a. Left maxillae, prolateral view. 50 b. Left strikers, retrolateral view. Photos: R. Raven." httpUri="https://zenodo.org/record/280930/files/figure.png" pageId="3" pageNumber="4">Fig. 49a</figureCitation>
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); (2) almost circular (Fig. 52a); (3) reinform (
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<figureCitation id="CE902A3AFF9B556AFDE470B9FD6346BC" box="[569,666,978,1002]" captionStart="FIGURES 51 a – 52 b. 51 a, b" captionStartId="28.[151,264,1143,1166]" captionTargetBox="[151,1436,181,1125]" captionTargetId="figure@28.[151,1436,181,1126]" captionTargetPageId="28" captionText="FIGURES 51 a – 52 b. 51 a, b. Orphnaecus species, female, QM S 83783. 51 a. left maxillae, prolateral view. 51 b. Orphnaecus species, female, QM S 83783, strikers, retrolateral view. 52 a. Selenocosmia javanensis (Walckenaer 1837), female, QM S 83754, left maxillae, prolateral view. 52 b. S. javanensis female QM S 83754, strikers, retrolateral view. Photos: S. Nunn." httpUri="https://zenodo.org/record/280931/files/figure.png" pageId="3" pageNumber="4">Fig. 51a</figureCitation>
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); (4) 1–3 rows of heavy clavates (not reniform) (Fig. 50a). Length = 6, CI = 0.66, RI = 0.86. In those selenocosmiine species that show loss of prolateral maxillary
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<taxonomicName id="91AB4D3CFF9B556AFACC709DFAC64158" box="[1297,1343,1014,1038]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
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, several issues arise, namely this trait (loss of
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<taxonomicName id="91AB4D3CFF9B556AFDAE7771FD594164" box="[627,672,1050,1074]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
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) is the singular defining feature of
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<taxonomicName id="91AB4D3CFF9B556AFBE87777FB8E4165" box="[1077,1143,1052,1075]" class="Arachnida" family="Theraphosidae" genus="Yamia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFBE87777FB8E4165" box="[1077,1143,1052,1075]" italics="true" pageId="3" pageNumber="4">Yamia</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="323A4B4EFF9B556AFB577771FA614164" author="Schmidt" box="[1162,1432,1050,1074]" pageId="3" pageNumber="4" refString="Haupt, J. & Schmidt, G. (2004) Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae). Spixiana, 27, 199 - 204." type="journal article" year="2004">Schmidt & Haupt 2004</bibRefCitation>
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,
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<bibRefCitation id="323A4B4EFF9B556AFF187754FEB64100" author="Raven" box="[197,335,1086,1110]" pageId="3" pageNumber="4" refString="Raven, R. J. (2005) A new tarantula species from northern Australia (Araneae, Theraphosidae). Zootaxa, 1004, 15 - 28." type="journal article" year="2005">Raven 2005</bibRefCitation>
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,
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<bibRefCitation id="323A4B4EFF9B556AFE817755FDC24100" author="Zhu" box="[348,571,1086,1110]" pageId="3" pageNumber="4" refString="Zhu, M. S. & Zhang, R. (2008) Revision of the theraphosid spiders from China (Araneae: Mygalomorphae). Journal of Arachnology, 36, 425 - 447." type="journal article" year="2008">Zhu & Zhang 2008</bibRefCitation>
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). Yet, many undescribed
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<taxonomicName id="91AB4D3CFF9B556AFC847755FC1B4100" box="[857,994,1086,1110]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFC847755FC1B4100" box="[857,994,1086,1110]" italics="true" pageId="3" pageNumber="4">Orphnaecus</emphasis>
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</taxonomicName>
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species (West & Nunn, pers. obs.) and some Australian species (R. Raven, pers. comm. 2008) also show losses of maxillary
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<taxonomicName id="91AB4D3CFF9B556AFB597709FB4A412C" box="[1156,1203,1122,1146]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
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. Secondly (and most critical to character state coding),
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<taxonomicName id="91AB4D3CFF9B556AFD8677EDFD2241C8" box="[603,731,1158,1182]" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFD8677EDFD2241C8" box="[603,731,1158,1182]" italics="true" pageId="3" pageNumber="4">Phlogiellus</emphasis>
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</taxonomicName>
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species show wide ranges of lyrate bacillae counts, but no distinct division between presence/absence (some species such as
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<taxonomicName id="91AB4D3CFF9B556AFC7977C7FC034195" box="[932,1018,1195,1219]" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="baeri">
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<emphasis id="64DFEAADFF9B556AFC7977C7FC034195" box="[932,1018,1195,1219]" italics="true" pageId="3" pageNumber="4">P. baeri</emphasis>
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</taxonomicName>
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show 1–3 bacillate setae). Therefore, instead of using the ambiguous state “absent”, due to overly subjective character delineation combined with false multi-state character coding (
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<bibRefCitation id="323A4B4EFF9B556AFD817799FD0D405C" author="Sereno" box="[604,756,1266,1290]" pageId="3" pageNumber="4" refString="Sereno, P. C. (2007) Logical basis for morphological characters in phylogenetics. Cladistics, 23, 565 - 587." type="journal article" year="2007">Sereno 2007</bibRefCitation>
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), it becomes logical to include the rudimentary condition within the same transformational character state (0). This resolves those subjective character state tendencies, as noted by
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<bibRefCitation id="323A4B4EFF9B556AFE8F7650FE084004" author="Jenner" box="[338,497,1338,1362]" pageId="3" pageNumber="4" refString="Jenner, R. A. (2004) The scientific status of metazoan cladistics: why current practice must change. Zoologica Scripta, 33, 293 - 310." type="journal article" year="2004">Jenner (2004)</bibRefCitation>
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,
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<bibRefCitation id="323A4B4EFF9B556AFE227651FD634004" author="Sereno" box="[511,666,1338,1362]" pageId="3" pageNumber="4" refString="Sereno, P. C. (2007) Logical basis for morphological characters in phylogenetics. Cladistics, 23, 565 - 587." type="journal article" year="2007">Sereno (2007</bibRefCitation>
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,
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<bibRefCitation id="323A4B4EFF9B556AFD7A7651FD1A4004" author="Sereno" box="[679,739,1338,1362]" pageId="3" pageNumber="4" refString="Sereno, P. C. (2009) Comparative cladistics. Cladistics, 25, 624 - 659." type="journal article" year="2009">2009</bibRefCitation>
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),
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<bibRefCitation id="323A4B4EFF9B556AFD247651FC734004" author="Assis" box="[761,906,1338,1362]" pageId="3" pageNumber="4" refString="Assis, L. C. S. (2009) Coherence, correspondence, and the renaissance of morphology in phylogenetic systematics. Cladistics, 25, 528 - 544" type="journal article" year="2009">Assis (2009)</bibRefCitation>
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and
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<bibRefCitation id="323A4B4EFF9B556AFC197650FB694004" box="[964,1168,1338,1363]" pageId="3" pageNumber="4" refString="Vogt, L., Bartolomaeus, T. & Giribet, G. (2010) The linguistic problem of morphology: structure versus homology and the standardization of morphological data. Cladistics, 26, 301 - 325." type="journal article">
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Vogt
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<emphasis id="64DFEAADFF9B556AFBDF7657FBCC4005" box="[1026,1077,1339,1363]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
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. (2010)
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</bibRefCitation>
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which can threaten the value of morphological characters. This was so critical to
|
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<bibRefCitation id="323A4B4EFF9B556AFCBE7634FBF94020" author="Jenner" box="[867,1024,1374,1398]" pageId="3" pageNumber="4" refString="Jenner, R. A. (2004) The scientific status of metazoan cladistics: why current practice must change. Zoologica Scripta, 33, 293 - 310." type="journal article" year="2004">Jenner (2004)</bibRefCitation>
|
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and
|
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<bibRefCitation id="323A4B4EFF9B556AFBE67635FB194020" author="Sereno" box="[1083,1248,1374,1398]" pageId="3" pageNumber="4" refString="Sereno, P. C. (2009) Comparative cladistics. Cladistics, 25, 624 - 659." type="journal article" year="2009">Sereno (2009)</bibRefCitation>
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, they stated that such artificial/weak character state delineation can “
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<emphasis id="64DFEAADFF9B556AFCD276E8FE4C40E8" italics="true" pageId="3" pageNumber="4">threaten the very future of morphology based cladistics as a vigorous discipline</emphasis>
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” (surely a strong enough argument as to support our reasoning). Other genera display unique maxillary lyrate arrangements,
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<taxonomicName id="91AB4D3CFF9B556AFDA476A0FCEA40B5" box="[633,787,1483,1507]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFDA476A0FCEA40B5" box="[633,787,1483,1507]" italics="true" pageId="3" pageNumber="4">Selenocosmia</emphasis>
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</taxonomicName>
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have an almost circular patch of very short bacillae (Fig. 52a), some
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<taxonomicName id="91AB4D3CFF9B556AFE977685FE2A4350" box="[330,467,1518,1542]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFE977685FE2A4350" box="[330,467,1518,1542]" italics="true" pageId="3" pageNumber="4">Orphnaecus</emphasis>
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</taxonomicName>
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possess a reniform patch (
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<figureCitation id="CE902A3AFF9B556AFCDB7685FC914350" box="[774,872,1518,1542]" captionStart="FIGURES 51 a – 52 b. 51 a, b" captionStartId="28.[151,264,1143,1166]" captionTargetBox="[151,1436,181,1125]" captionTargetId="figure@28.[151,1436,181,1126]" captionTargetPageId="28" captionText="FIGURES 51 a – 52 b. 51 a, b. Orphnaecus species, female, QM S 83783. 51 a. left maxillae, prolateral view. 51 b. Orphnaecus species, female, QM S 83783, strikers, retrolateral view. 52 a. Selenocosmia javanensis (Walckenaer 1837), female, QM S 83754, left maxillae, prolateral view. 52 b. S. javanensis female QM S 83754, strikers, retrolateral view. Photos: S. Nunn." httpUri="https://zenodo.org/record/280931/files/figure.png" pageId="3" pageNumber="4">Fig. 51a</figureCitation>
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), while
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<taxonomicName id="91AB4D3CFF9B556AFC197685FBA24350" box="[964,1115,1518,1542]" class="Arachnida" family="Theraphosidae" genus="Chilobrachys" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="64DFEAADFF9B556AFC197685FBA24350" box="[964,1115,1518,1542]" italics="true" pageId="3" pageNumber="4">Chilobrachys</emphasis>
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</taxonomicName>
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<taxonomicName id="91AB4D3CFF9B556AFBB97685FB694350" box="[1124,1168,1518,1542]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
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comprises 1–3 rows of heavy claviform bacillae (Fig. 50a).
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</paragraph>
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<paragraph id="561436BFFF9B556AFF4A755DFCE54388" blockId="3.[151,1436,907,1867]" pageId="3" pageNumber="4">
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6. Setal fringe ventrally to prolateral maxillary suture: (0) absent; (1) fine pallid setae (
|
||
<figureCitation id="CE902A3AFF9B556AFB66755DFADC4318" box="[1211,1317,1590,1614]" captionStart="FIGURES 49 a – 50 b. 49 a, b" captionStartId="27.[151,264,1127,1150]" captionTargetBox="[151,1436,193,1104]" captionTargetId="figure@27.[151,1436,193,1106]" captionTargetPageId="27" captionText="FIGURES 49 a – 50 b. 49 a, b. Haplocosmia himalayana (Pocock 1899), holotype female, NHM 1899.8. 19.3. 49 a. Right maxillae (inverted), prolateral view. 49 b. Right chelicerae, prolateral view. 50 a, b. Chilobrachys sericeus (Thorell 1895), syntype male, NHM 1895.9. 21.21. 50 a. Left maxillae, prolateral view. 50 b. Left strikers, retrolateral view. Photos: R. Raven." httpUri="https://zenodo.org/record/280930/files/figure.png" pageId="3" pageNumber="4">Figs 49a</figureCitation>
|
||
, 50a); (2) scopulate, dark setae (
|
||
<figureCitation id="CE902A3AFF9B556AFE1F7531FDEE4324" box="[450,535,1626,1650]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="3" pageNumber="4">Fig. 47</figureCitation>
|
||
). Length = 3, CI = 0.66, RI = 0.8. The genera
|
||
<taxonomicName id="91AB4D3CFF9B556AFBF37530FB3F4325" box="[1070,1222,1627,1651]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFBF37530FB3F4325" box="[1070,1222,1627,1651]" italics="true" pageId="3" pageNumber="4">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91AB4D3CFF9B556AFB087530FA934325" box="[1237,1386,1627,1651]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFB087530FA934325" box="[1237,1386,1627,1651]" italics="true" pageId="3" pageNumber="4">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<taxonomicName id="91AB4D3CFF9B556AFEDB7515FE6443C0" box="[262,413,1662,1686]" class="Arachnida" family="Theraphosidae" genus="Chilobrachys" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFEDB7515FE6443C0" box="[262,413,1662,1686]" italics="true" pageId="3" pageNumber="4">Chilobrachys</emphasis>
|
||
</taxonomicName>
|
||
possess a setal fringe just ventral to the prolateral maxillary suture, but dorsal to the
|
||
<taxonomicName id="91AB4D3CFF9B556AFA917515FA8243C0" box="[1356,1403,1662,1686]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
|
||
. In
|
||
<taxonomicName id="91AB4D3CFF9B556AFF1875C8FEA743ED" box="[197,350,1699,1723]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFF1875C8FEA743ED" box="[197,350,1699,1723]" italics="true" pageId="3" pageNumber="4">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
this fringe is dark, almost scopulate (
|
||
<figureCitation id="CE902A3AFF9B556AFD2375C9FCA843EC" box="[766,849,1698,1722]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="3" pageNumber="4">Fig. 47</figureCitation>
|
||
), while in
|
||
<taxonomicName id="91AB4D3CFF9B556AFC1B75C8FBA243ED" box="[966,1115,1699,1723]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFC1B75C8FBA243ED" box="[966,1115,1699,1723]" italics="true" pageId="3" pageNumber="4">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CE902A3AFF9B556AFBB775C9FB3143EC" box="[1130,1224,1698,1722]" captionStart="FIGURES 49 a – 50 b. 49 a, b" captionStartId="27.[151,264,1127,1150]" captionTargetBox="[151,1436,193,1104]" captionTargetId="figure@27.[151,1436,193,1106]" captionTargetPageId="27" captionText="FIGURES 49 a – 50 b. 49 a, b. Haplocosmia himalayana (Pocock 1899), holotype female, NHM 1899.8. 19.3. 49 a. Right maxillae (inverted), prolateral view. 49 b. Right chelicerae, prolateral view. 50 a, b. Chilobrachys sericeus (Thorell 1895), syntype male, NHM 1895.9. 21.21. 50 a. Left maxillae, prolateral view. 50 b. Left strikers, retrolateral view. Photos: R. Raven." httpUri="https://zenodo.org/record/280930/files/figure.png" pageId="3" pageNumber="4">Fig. 49a</figureCitation>
|
||
) and
|
||
<taxonomicName id="91AB4D3CFF9B556AFAD875C8FA6543ED" box="[1285,1436,1699,1723]" class="Arachnida" family="Theraphosidae" genus="Chilobrachys" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFAD875C8FA6543ED" box="[1285,1436,1699,1723]" italics="true" pageId="3" pageNumber="4">Chilobrachys</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 50a) the fringe is formed from fine, pallid setae.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9B556AFF4A7580FB09421D" blockId="3.[151,1436,907,1867]" pageId="3" pageNumber="4">
|
||
7. Tubercles on prolateral maxillary surface: (0) absent; (1) present (
|
||
<figureCitation id="CE902A3AFF9B556AFC627581FBEF4254" box="[959,1046,1770,1794]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="3" pageNumber="4">Fig. 47</figureCitation>
|
||
). Length = 1, CI = 1, RI = 0. All
|
||
<taxonomicName id="91AB4D3CFF9B556AFF187465FEA44270" box="[197,349,1806,1830]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9B556AFF187465FEA44270" box="[197,349,1806,1830]" italics="true" pageId="3" pageNumber="4">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
possess 1–12 tubercles immediately distad to patch of rudimentary bacillae on prolateral maxillary surfaces (
|
||
<figureCitation id="CE902A3AFF9B556AFE7F7459FE0E421C" box="[418,503,1842,1866]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="3" pageNumber="4">Fig. 47</figureCitation>
|
||
). These tubercles are not known in any other selenocosmiine genus.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9B556DFF1A7411FD9B444A" blockId="3.[197,1436,1914,2011]" lastBlockId="4.[151,1436,151,895]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Sexual morphology (female):
|
||
<bibRefCitation id="323A4B4EFF9B556AFDC37411FCCE42C4" author="Perez-Miles" box="[542,823,1914,1939]" pageId="3" pageNumber="4" refString="Perez-Miles, F., Lucas, S. M., da Silva Jr., P. I. & Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
|
||
Pérez-Miles
|
||
<emphasis id="64DFEAADFF9B556AFD727417FD1842C5" box="[687,737,1915,1939]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
. (1996
|
||
</bibRefCitation>
|
||
: characters 13, 14) used spermathecal morphology in their cladistic analysis of
|
||
<taxonomicName id="91AB4D3CFF9B556AFDDC74F5FD5042E0" box="[513,681,1950,1974]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subFamily" subFamily="Theraphosinae">Theraphosinae</taxonomicName>
|
||
.
|
||
<bibRefCitation id="323A4B4EFF9B556AFD6674F5FB8442E0" author="Guadanucci" box="[699,1149,1950,1974]" pageId="3" pageNumber="4" refString="Guadanucci, J. P. L. (2005) Tarsal scopula significance in Ischnocolinae phylogenetics (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology, 33, 456 - 467." type="journal article" year="2005">Guadanucci (2005: characters 11, 12)</bibRefCitation>
|
||
also used spermathecal morphology in his analysis of Ischnocolinae.
|
||
<bibRefCitation id="323A4B4EFF9B556AFD0574A9FC0C428C" author="Perez-Miles" box="[728,1013,1986,2011]" pageId="3" pageNumber="4" refString="Perez-Miles, F., Gabriel, R., Miglio, L., Bonaldo, A., Gallon, R., Jimenez, J. J. & Bertani, R. (2008) Ami, a new theraphosid genus from Central and South America, with the description of six new species (Araneae: Mygalomorphae). Zootaxa, 1915, 54 - 68." type="journal article" year="2008">
|
||
Pérez-Miles
|
||
<emphasis id="64DFEAADFF9B556AFCB774AFFC67428D" box="[874,926,1987,2011]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
. (2008
|
||
</bibRefCitation>
|
||
: characters 8–11) used spermathecal morphology to define their new theraphosine genus
|
||
<taxonomicName id="91AB4D3CFF9C556DFCC773F3FBE545F9" authority="Perez-Miles 2008" authorityName="Perez-Miles" authorityYear="2008" box="[794,1052,151,176]" class="Arachnida" family="Theraphosidae" genus="Ami" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFCC773F3FCBE45E6" box="[794,839,152,176]" italics="true" pageId="4" pageNumber="5">Ami</emphasis>
|
||
<bibRefCitation id="323A4B4EFF9C556DFC8D73FCFBE545F9" author="Perez-Miles" box="[848,1052,151,175]" pageId="4" pageNumber="5" refString="Perez-Miles, F., Gabriel, R., Miglio, L., Bonaldo, A., Gallon, R., Jimenez, J. J. & Bertani, R. (2008) Ami, a new theraphosid genus from Central and South America, with the description of six new species (Araneae: Mygalomorphae). Zootaxa, 1915, 54 - 68." type="journal article" year="2008">Pérez-Miles 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
; West
|
||
<emphasis id="64DFEAADFF9C556DFBB773F2FB6445E6" box="[1130,1181,152,176]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. (2008: characters 23, 24) used spermathecal morphology in their analysis of
|
||
<taxonomicName id="91AB4D3CFF9C556DFC9573D7FC084582" box="[840,1009,188,212]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Theraphosidae</taxonomicName>
|
||
. Five characters were used based on spermathecal morphology, all of which (with the exception of character 8, states 0 and 1 = West
|
||
<emphasis id="64DFEAADFF9C556DFAC7738AFAB545AE" box="[1306,1356,224,248]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
|
||
. 2008: character 23, states 2 and 0) are new.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A724CFAA544FA" blockId="4.[151,1436,151,895]" pageId="4" pageNumber="5">
|
||
8. Spermathecae: (0) fused (
|
||
<figureCitation id="CE902A3AFF9C556DFE2D724CFDB04469" box="[496,585,295,319]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 28</figureCitation>
|
||
b, 30); (1) not fused (
|
||
<figureCitation id="CE902A3AFF9C556DFC9A724CFC654469" box="[839,924,295,319]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 31</figureCitation>
|
||
b). Length = 2, CI = 0.5, RI = 0.83. Within
|
||
<taxonomicName id="91AB4D3CFF9C556DFF187227FE7A4432" box="[197,387,332,356]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">Selenocosmiinae</taxonomicName>
|
||
examined, both fused (
|
||
<bibRefCitation id="323A4B4EFF9C556DFD557227FCF64432" author="Smith" box="[648,783,332,356]" pageId="4" pageNumber="5" refString="Smith, A. M. (2004) A new species of the arboreal theraphosid, genus Poecilotheria, from southern India (Araneae, Mygalomorphae, Theraphosidae) with notes on its conservation status. British Tarantula Society Journal, 19, 48 - 61." type="journal article" year="2004">Smith 2004</bibRefCitation>
|
||
: fig. 6; 2006: fig. 6) and unfused (
|
||
<bibRefCitation id="323A4B4EFF9C556DFB567227FAB64432" author="Kulczynski" box="[1163,1359,332,356]" pageId="4" pageNumber="5" refString="Kulczynski, W. (1908) Symbola ad faunam Aranearum Javae et Sumatrae cognoscendam. I. Mygalomorphae et Cribellatae. Bulletin international de l'Academie des sciences de Cracovie, 1908, 527 - 581." type="journal article" year="1908">Kulczyński 1908</bibRefCitation>
|
||
: fig. 1;
|
||
<bibRefCitation id="323A4B4EFF9C556DFF187204FE9C44D1" author="Schmidt" box="[197,357,367,391]" pageId="4" pageNumber="5" refString="Schmidt, G. (1999) Selenobrachys philippinus gen. et sp. n. (Araneae: Theraphosidae: Selenocosmiinae), eine neue Theraphosidae sp. von der Insel Negros (Philippinen). Arachnologisches Magazin, 7 (5 / 6), 1 - 13." type="journal article" year="1999">Schmidt 1999</bibRefCitation>
|
||
: fig. 5;
|
||
<bibRefCitation id="323A4B4EFF9C556DFE64721BFD3844D1" author="Haupt" box="[441,705,367,391]" pageId="4" pageNumber="5" refString="Haupt, J. & Schmidt, G. (2004) Description of the male and illustration of the female receptacula of Yamia watasei Kishida, 1920 (Arachnida, Araneae, Theraphosidae, Selenocosmiinae). Spixiana, 27, 199 - 204." type="journal article" year="2004">Haupt & Schmidt 2004</bibRefCitation>
|
||
: figs 7–9;
|
||
<bibRefCitation id="323A4B4EFF9C556DFCE87204FBE944D1" author="Zhu" box="[821,1040,367,391]" pageId="4" pageNumber="5" refString="Zhu, M. S. & Zhang, R. (2008) Revision of the theraphosid spiders from China (Araneae: Mygalomorphae). Journal of Arachnology, 36, 425 - 447." type="journal article" year="2008">Zhu & Zhang 2008</bibRefCitation>
|
||
: figs
|
||
<date id="2215107FFF9C556DFB917204FB9244D1" box="[1100,1131,367,391]" pageId="4" pageNumber="5">4 I</date>
|
||
,
|
||
<date id="2215107FFF9C556DFBA57204FB6144D1" box="[1144,1176,367,391]" pageId="4" pageNumber="5">6 I</date>
|
||
,
|
||
<date id="2215107FFF9C556DFB797204FB3A44D1" box="[1188,1219,367,391]" pageId="4" pageNumber="5">8 I</date>
|
||
, 9 F;
|
||
<bibRefCitation id="323A4B4EFF9C556DFADE721BFEF744FA" author="West" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a</bibRefCitation>
|
||
: figs 6, 23, 51, 70, 101; 2010b: figs 17, 37, 84, 141, 160, 181, 200, 201) spermathecae were noted.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A72DCFAFD476A" blockId="4.[151,1436,151,895]" pageId="4" pageNumber="5">
|
||
9. Lateral bilobe spermathecal morphology: (0) apically swollen (
|
||
<figureCitation id="CE902A3AFF9C556DFC7F72DCFC014499" box="[930,1016,439,463]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Fig. 38</figureCitation>
|
||
b); (1) not apically swollen (
|
||
<figureCitation id="CE902A3AFF9C556DFA9B72DCFA744499" box="[1350,1421,439,463]" captionStart="FIGURES 2 – 7" captionStartId="17.[151,264,1492,1515]" captionTargetBox="[151,1436,157,1470]" captionTargetId="figure@17.[151,1436,157,1471]" captionTargetPageId="17" captionText="FIGURES 2 – 7. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 2. Carapace, dorsal view. 3. Eyes, dorsal view. 4. Labium, ventral view. 5. Sternum, labium, maxilla and coxae, ventral view. 6. Spermatheca, ventral view. 7. Abdomen, dorsal view. Scale bar = 10 mm for 2, 5 and 7; 5 mm for 4; 2 mm for 3; 1 mm for 6. Photos: S. Nunn." httpUri="https://zenodo.org/record/280921/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
||
). Length = 2, CI = 0.5, RI = 0.75. In all
|
||
<taxonomicName id="91AB4D3CFF9C556DFDB172B7FCF544A2" box="[620,780,476,500]" class="Arachnida" family="Theraphosidae" genus="Psednocnemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFDB172B7FCF544A2" box="[620,780,476,500]" italics="true" pageId="4" pageNumber="5">Psednocnemis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFCC472B7FA6544A2" box="[793,1436,476,500]" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article">
|
||
West & Nunn 2010b, figs 37, 64, 141; herein, (
|
||
<figureCitation id="CE902A3AFF9C556DFAFE72B7FA9244A2" box="[1315,1387,476,500]" captionStart="FIGURES 2 – 7" captionStartId="17.[151,264,1492,1515]" captionTargetBox="[151,1436,157,1470]" captionTargetId="figure@17.[151,1436,157,1471]" captionTargetPageId="17" captionText="FIGURES 2 – 7. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 2. Carapace, dorsal view. 3. Eyes, dorsal view. 4. Labium, ventral view. 5. Sternum, labium, maxilla and coxae, ventral view. 6. Spermatheca, ventral view. 7. Abdomen, dorsal view. Scale bar = 10 mm for 2, 5 and 7; 5 mm for 4; 2 mm for 3; 1 mm for 6. Photos: S. Nunn." httpUri="https://zenodo.org/record/280921/files/figure.png" pageId="4" pageNumber="5">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="CE902A3AFF9C556DFAAB72B7FA6D44A2" box="[1398,1428,476,500]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">40</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
as well as
|
||
<taxonomicName id="91AB4D3CFF9C556DFE99716AFE3D474E" box="[324,452,512,536]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="64DFEAADFF9C556DFE99716AFE3D474E" box="[324,452,512,536]" italics="true" pageId="4" pageNumber="5">L. robustus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFE0B7294FD984741" author="Smith" box="[470,609,511,535]" pageId="4" pageNumber="5" refString="Smith, A. M. (1988) Lyrognathus robustus, a new species of theraphosid spider from Malaysia. British Tarantula Society Journal, 4 (2), 15 - 19." type="journal article" year="1988">Smith 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="323A4B4EFF9C556DFDAD716BFD144741" author="West" box="[624,749,511,535]" pageId="4" pageNumber="5" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991">West 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="323A4B4EFF9C556DFD21716BFC164741" author="West" box="[764,1007,511,535]" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a</bibRefCitation>
|
||
, fig. 101), lateral bilobes lack apical swelling, differing from all other selenocosmiine species with bilobed spermathecal morphology.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A712CFCA147D2" blockId="4.[151,1436,151,895]" pageId="4" pageNumber="5">
|
||
10. Bilobed spermathecae with medial and lateral lobes of similar length (
|
||
<figureCitation id="CE902A3AFF9C556DFC3B712CFBC54709" box="[998,1084,583,607]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Fig. 43</figureCitation>
|
||
): (0) absent (
|
||
<figureCitation id="CE902A3AFF9C556DFB0F712CFAD04709" box="[1234,1321,583,607]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 38</figureCitation>
|
||
b, 45); (1) present (
|
||
<figureCitation id="CE902A3AFF9C556DFEF87107FE2947D2" box="[293,464,620,644]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 40, 43, 44</figureCitation>
|
||
b). Length = 2, CI = 0.5, RI = 0.91.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A71E4FEF94642" blockId="4.[151,1436,151,895]" pageId="4" pageNumber="5">
|
||
11. Spermathecal lobes with mushroom cap ends (mushroomiform): (0) absent; (1) present. Length = 2, CI = 0.5, RI = 0.8. All
|
||
<taxonomicName id="91AB4D3CFF9C556DFE8771DFFE0D479A" box="[346,500,692,716]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFE8771DFFE0D479A" box="[346,500,692,716]" italics="true" pageId="4" pageNumber="5">Selenocosmia</emphasis>
|
||
</taxonomicName>
|
||
within this analysis (
|
||
<figureCitation id="CE902A3AFF9C556DFD3F71DFFCC0479A" box="[738,825,692,716]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 44</figureCitation>
|
||
b, 45),
|
||
<taxonomicName id="91AB4D3CFF9C556DFC5B71DEFC10479A" box="[902,1001,692,716]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="fuscus">
|
||
<emphasis id="64DFEAADFF9C556DFC5B71DEFC10479A" box="[902,1001,692,716]" italics="true" pageId="4" pageNumber="5">L. fuscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91AB4D3CFF9C556DFBFD71DEFB66479A" box="[1056,1183,692,716]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lessunda">
|
||
<emphasis id="64DFEAADFF9C556DFBFD71DEFB66479A" box="[1056,1183,692,716]" italics="true" pageId="4" pageNumber="5">L. lessunda</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFB7071DFFEBD47B9" author="West" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a: figs 51, 70</bibRefCitation>
|
||
), respectively, herein: (
|
||
<figureCitation id="CE902A3AFF9C556DFD8D71BCFD5A47B9" box="[592,675,727,751]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Fig. 38</figureCitation>
|
||
b) possess at least one lobe with mushroom cap (mushroomiform) ends.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A7074FEAE4629" blockId="4.[151,1436,151,895]" pageId="4" pageNumber="5">
|
||
12. Spermathecae tombstone-shaped (
|
||
<figureCitation id="CE902A3AFF9C556DFD957074FD674661" box="[584,670,799,823]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 36</figureCitation>
|
||
): (0) absent; (1) present. Length = 1, CI = 1, RI = 0. This trait was noted in an undescribed
|
||
<taxonomicName id="91AB4D3CFF9C556DFE04702FFD9B460A" box="[473,610,836,860]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFE04702FFD9B460A" box="[473,610,836,860]" italics="true" pageId="4" pageNumber="5">Orphnaecus</emphasis>
|
||
</taxonomicName>
|
||
species used in our analysis as well as
|
||
<taxonomicName id="91AB4D3CFF9C556DFBC2702FFAD7460A" box="[1055,1326,836,860]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="philippinus">
|
||
<emphasis id="64DFEAADFF9C556DFBC2702FFAD7460A" box="[1055,1326,836,860]" italics="true" pageId="4" pageNumber="5">Orphnaecus philippinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFAE0702FFEB04629" author="Schmidt" pageId="4" pageNumber="5" refString="Schmidt, G. (1999) Selenobrachys philippinus gen. et sp. n. (Araneae: Theraphosidae: Selenocosmiinae), eine neue Theraphosidae sp. von der Insel Negros (Philippinen). Arachnologisches Magazin, 7 (5 / 6), 1 - 13." type="journal article" year="1999">Schmidt 1999: fig. 5</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF1A70C4FE6B4162" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
Sexual morphology (male): Male palpal bulb characters are well established synapomorphies of and within
|
||
<taxonomicName id="91AB4D3CFF9C556DFF1870BFFE9346BA" box="[197,362,980,1004]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Theraphosinae">Theraphosinae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFEA170BFFDF346BA" author="Raven" box="[380,522,980,1004]" pageId="4" pageNumber="5" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven 1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="323A4B4EFF9C556DFDC470BFFCD546BA" author="Perez-Miles" box="[537,812,980,1004]" pageId="4" pageNumber="5" refString="Perez-Miles, F., Lucas, S. M., da Silva Jr., P. I. & Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
|
||
Pérez-Miles
|
||
<emphasis id="64DFEAADFF9C556DFD7670BEFD1F46BA" box="[683,742,980,1004]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
1996
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="323A4B4EFF9C556DFCE770BFFC2D46BA" author="Bertani" box="[826,980,980,1004]" pageId="4" pageNumber="5" refString="Bertani, R. (2000) Male palpal bulbs and homologous features in Theraphosinae (Araneae, Theraphosidae). Journal of Arachnology, 28, 29 - 42" type="journal article" year="2000">Bertani 2000</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="323A4B4EFF9C556DFC3770BFFB6846BA" author="Bertani" box="[1002,1169,980,1004]" pageId="4" pageNumber="5" refString="Bertani, R. (2000) Male palpal bulbs and homologous features in Theraphosinae (Araneae, Theraphosidae). Journal of Arachnology, 28, 29 - 42" type="journal article" year="2000">Bertani (2000)</bibRefCitation>
|
||
identified homologous keels within
|
||
<taxonomicName id="91AB4D3CFF9C556DFE8E709CFE024159" box="[339,507,1015,1039]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Theraphosinae">Theraphosinae</taxonomicName>
|
||
, we follow him in exploring embolus morphology in
|
||
<taxonomicName id="91AB4D3CFF9C556DFB91709CFAF44159" box="[1100,1293,1015,1039]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">Selenocosmiinae</taxonomicName>
|
||
, utilizing six characters, herein.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A7754FD3F412A" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
13. Terete embolus lanceolate (LTE) (
|
||
<figureCitation id="CE902A3AFF9C556DFD947754FD374101" box="[585,718,1087,1111]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 41, 44</figureCitation>
|
||
a): (0) absent (
|
||
<figureCitation id="CE902A3AFF9C556DFCAC7754FC304101" box="[881,969,1087,1111]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 28</figureCitation>
|
||
a, 31a, 32, 34, 35a, 37a, 38a); (1) present (
|
||
<figureCitation id="CE902A3AFF9C556DFF10770FFE80412A" box="[205,377,1124,1148]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 39, 41, 44</figureCitation>
|
||
a). Length = 1, CI = 1, RI = 0.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A77ECFB674079" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
14. Single retrolateral keel on embolus (R): (0) absent; (1) strong and heavy (
|
||
<figureCitation id="CE902A3AFF9C556DFBC977ECFB6141C9" box="[1044,1176,1159,1183]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 34, 35</figureCitation>
|
||
a); (2) weak (
|
||
<figureCitation id="CE902A3AFF9C556DFAEF77ECFA7341C9" box="[1330,1418,1159,1183]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 37</figureCitation>
|
||
a, 38a). Length = 2, CI = 1, RI = 0. In all
|
||
<taxonomicName id="91AB4D3CFF9C556DFDAA77C7FD0E4192" box="[631,759,1196,1220]" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFDAA77C7FD0E4192" box="[631,759,1196,1220]" italics="true" pageId="4" pageNumber="5">Phlogiellus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CE902A3AFF9C556DFCD877C7FCAE4192" box="[773,855,1196,1220]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 34</figureCitation>
|
||
) and
|
||
<taxonomicName id="91AB4D3CFF9C556DFC4877C7FBE74192" box="[917,1054,1196,1220]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFC4877C7FBE74192" box="[917,1054,1196,1220]" italics="true" pageId="4" pageNumber="5">Orphnaecus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CE902A3AFF9C556DFBF077C7FB874192" box="[1069,1150,1196,1220]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 35</figureCitation>
|
||
a) examined, a strong and heavy single keel (similar position to R, or retrolateral keel of
|
||
<bibRefCitation id="323A4B4EFF9C556DFC4477A4FBD641B1" author="Bertani" box="[921,1071,1231,1255]" pageId="4" pageNumber="5" refString="Bertani, R. (2000) Male palpal bulbs and homologous features in Theraphosinae (Araneae, Theraphosidae). Journal of Arachnology, 28, 29 - 42" type="journal article" year="2000">Bertani 2000</bibRefCitation>
|
||
) runs along the entire embolus, twisting 90 degrees distally.
|
||
<taxonomicName id="91AB4D3CFF9C556DFDDE779FFD76405A" box="[515,655,1268,1292]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFDDE779FFD76405A" box="[515,655,1268,1292]" italics="true" pageId="4" pageNumber="5">Lyrognathus</emphasis>
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="91AB4D3CFF9C556DFD33779EFC92405A" box="[750,875,1268,1292]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="robustus">
|
||
<emphasis id="64DFEAADFF9C556DFD33779EFC92405A" box="[750,875,1268,1292]" italics="true" pageId="4" pageNumber="5">L. robustus</emphasis>
|
||
</taxonomicName>
|
||
) also possess a single retrolateral (RR) keel (
|
||
<bibRefCitation id="323A4B4EFF9C556DFABF779FFE894079" author="West" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a</bibRefCitation>
|
||
: figs 5, 40 and, herein,
|
||
<figureCitation id="CE902A3AFF9C556DFDA8767CFD334079" box="[629,714,1303,1327]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 37</figureCitation>
|
||
a, 38a), although it is far less pronounced.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A7657FDDA4351" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
15. Basal embolus with distinct heavy retrolateral lobe (BL): (0) absent; (1) present (
|
||
<figureCitation id="CE902A3AFF9C556DFB8E7657FB2C4002" box="[1107,1237,1340,1364]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 34, 35</figureCitation>
|
||
a). Length = 1, CI = 1, RI = 0. In all
|
||
<taxonomicName id="91AB4D3CFF9C556DFE44760BFDE0402E" box="[409,537,1376,1400]" class="Arachnida" family="Theraphosidae" genus="Phlogiellus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFE44760BFDE0402E" box="[409,537,1376,1400]" italics="true" pageId="4" pageNumber="5">Phlogiellus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFDF47634FB624021" box="[553,1179,1375,1399]" pageId="4" pageNumber="5" refString="Zhu, M. S. & Zhang, R. (2008) Revision of the theraphosid spiders from China (Araneae: Mygalomorphae). Journal of Arachnology, 36, 425 - 447." type="journal article">
|
||
Zhu & Zhang 2008: figs 8 B,
|
||
<date id="2215107FFF9C556DFC5F7634FC584021" box="[898,929,1375,1399]" pageId="4" pageNumber="5">9 I</date>
|
||
and, herein, (
|
||
<figureCitation id="CE902A3AFF9C556DFB9D7634FB6A4021" box="[1088,1171,1375,1399]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 34</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
and
|
||
<taxonomicName id="91AB4D3CFF9C556DFB08760BFAA7402E" box="[1237,1374,1376,1400]" class="Arachnida" family="Theraphosidae" genus="Orphnaecus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFB08760BFAA7402E" box="[1237,1374,1376,1400]" italics="true" pageId="4" pageNumber="5">Orphnaecus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CE902A3AFF9C556DFAB37634FF1A40CA" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 35</figureCitation>
|
||
a), retrolateral embolus keel terminates basally as an enlarged lobe (BL). A similar lobe is present in
|
||
<taxonomicName id="91AB4D3CFF9C556DFF1876CCFEA840E9" box="[197,337,1447,1471]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Lyrognathus</taxonomicName>
|
||
(RBL) (
|
||
<figureCitation id="CE902A3AFF9C556DFE6C76CCFDF040E9" box="[433,521,1447,1471]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Figs 37</figureCitation>
|
||
a, 38a), but is far smaller, not so distinct. Additionally, an apophysis is present in
|
||
<taxonomicName id="91AB4D3CFF9C556DFF1876A7FE1F40B2" box="[197,486,1484,1508]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="himalayana">
|
||
<emphasis id="64DFEAADFF9C556DFF1876A7FE1F40B2" box="[197,486,1484,1508]" italics="true" pageId="4" pageNumber="5">Haplocosmia himalayana</emphasis>
|
||
</taxonomicName>
|
||
, however, this structure originates from the tegulum, not from the embolus and thus not applicable to this character.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A757FFEC84322" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
16. Embolus with distal (A) and sub-distal (SA) keels (
|
||
<figureCitation id="CE902A3AFF9C556DFCDC757FFCAF437A" box="[769,854,1556,1580]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 31</figureCitation>
|
||
a, 32): (0) absent; (1) present. Length = 1, CI = 1, RI = 0. All
|
||
<taxonomicName id="91AB4D3CFF9C556DFEFA7553FE474306" box="[295,446,1592,1616]" class="Arachnida" family="Theraphosidae" genus="Chilobrachys" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFEFA7553FE474306" box="[295,446,1592,1616]" italics="true" pageId="4" pageNumber="5">Chilobrachys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91AB4D3CFF9C556DFE2A7553FD754306" box="[503,652,1592,1616]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFE2A7553FD754306" box="[503,652,1592,1616]" italics="true" pageId="4" pageNumber="5">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
used in this analysis (
|
||
<figureCitation id="CE902A3AFF9C556DFC57755CFC184319" box="[906,993,1591,1615]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Figs 31</figureCitation>
|
||
a, 32, respectively) possess both A and SA keels.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556DFF4A7514FD954389" blockId="4.[151,1436,943,2012]" pageId="4" pageNumber="5">
|
||
17. Distal spiral curl on embolus (AC): (0) absent; (1) present (
|
||
<figureCitation id="CE902A3AFF9C556DFCBA7514FC4543C1" box="[871,956,1663,1687]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Fig. 39</figureCitation>
|
||
). Length = 1, CI = 1, RI = 0. In all known male
|
||
<taxonomicName id="91AB4D3CFF9C556DFEDE75CFFE5A43EA" box="[259,419,1700,1724]" class="Arachnida" family="Theraphosidae" genus="Psednocnemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFEDE75CFFE5A43EA" box="[259,419,1700,1724]" italics="true" pageId="4" pageNumber="5">Psednocnemis</emphasis>
|
||
</taxonomicName>
|
||
(herein,
|
||
<figureCitation id="CE902A3AFF9C556DFDD775CFFDA443EA" box="[522,605,1700,1724]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="4" pageNumber="5">Fig. 39</figureCitation>
|
||
;
|
||
<bibRefCitation id="323A4B4EFF9C556DFDB775CEFCA143EA" author="West" box="[618,856,1700,1724]" pageId="4" pageNumber="5" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn 2010b</bibRefCitation>
|
||
: figs 58, 116, 135), the embolus terminates distally with distinct distal “spiral curl” (AC).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9C556CFF4A7587FBFA4582" blockId="4.[151,1436,943,2012]" lastBlockId="5.[197,1436,151,213]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
18. Embolus with multiple distal keels: (0) present; (1) absent. Length = 1, CI = 1, RI = 0. Among all species used in this analysis, those with more than one keel (or what could be interpreted as a keel) were scored with the state “present”. The outgroup taxon
|
||
<taxonomicName id="91AB4D3CFF9C556DFD82745FFCD7421A" box="[607,814,1844,1868]" class="Arachnida" family="Barychelidae" genus="Reichlingia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="annae">
|
||
<emphasis id="64DFEAADFF9C556DFD82745FFCD7421A" box="[607,814,1844,1868]" italics="true" pageId="4" pageNumber="5">Reichlingia annae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFCE1745FFBCC421A" author="Reichling" box="[828,1077,1844,1868]" pageId="4" pageNumber="5" refString="Reichling, S. B. (1997) A diminutive new species of Acanthopelma from Belize (Araneae: Theraphosidae). Bulletin of the British Arachnological Society, 10, 337 - 340." type="journal article" year="1997">Reichling 1997: fig. 1</bibRefCitation>
|
||
) possesses a ventral crest (VC) and prolateral inferior (PI) keel. In
|
||
<taxonomicName id="91AB4D3CFF9C556DFD907433FD1F4226" box="[589,742,1880,1904]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFD907433FD1F4226" box="[589,742,1880,1904]" italics="true" pageId="4" pageNumber="5">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9C556DFD29743CFC824239" author="Smith" box="[756,891,1879,1903]" pageId="4" pageNumber="5" refString="Smith, A. M. (2004) A new species of the arboreal theraphosid, genus Poecilotheria, from southern India (Araneae, Mygalomorphae, Theraphosidae) with notes on its conservation status. British Tarantula Society Journal, 19, 48 - 61." type="journal article" year="2004">Smith 2004</bibRefCitation>
|
||
: fig. 13, 14;
|
||
<bibRefCitation id="323A4B4EFF9C556DFBDB743CFB754239" author="Smith" box="[1030,1164,1879,1903]" pageId="4" pageNumber="5" refString="Smith, A. M. (2006) A new species of Poecilotheria from northern peninsular India (Araneae, Mygalomorphae, Theraphosidae) with notes on its distribution and conservation status. British Tarantula Society Journal, 21, 83 - 94." type="journal article" year="2006">Smith 2006</bibRefCitation>
|
||
: figs 13, 14; herein,
|
||
<figureCitation id="CE902A3AFF9C556DFAB2743CFF1B42C2" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="4" pageNumber="5">Fig. 28</figureCitation>
|
||
a) three keels are distinct, one reverse twisting retrolateral (RCR) keel, as well as prolateral superior (PS) and prolateral inferior (PI) keels, corkscrewing along the entire embolus (in the opposite direction to other selenocosmiine species with embolus keels).
|
||
<taxonomicName id="91AB4D3CFF9C556DFD1774AFFC98428A" box="[714,865,1988,2012]" class="Arachnida" family="Theraphosidae" genus="Chilobrachys" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9C556DFD1774AFFC98428A" box="[714,865,1988,2012]" italics="true" pageId="4" pageNumber="5">Chilobrachys</emphasis>
|
||
</taxonomicName>
|
||
displays prolateral superior (PS) and inferior (PI) keels, sub-distal (SA) and distal (A) keels (
|
||
<figureCitation id="CE902A3AFF9D556CFD6D73FCFCF845F9" box="[688,769,151,175]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="5" pageNumber="6">Fig. 31</figureCitation>
|
||
a) and in
|
||
<taxonomicName id="91AB4D3CFF9D556CFCB373F3FBFA45E6" box="[878,1027,152,176]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFCB373F3FBFA45E6" box="[878,1027,152,176]" italics="true" pageId="5" pageNumber="6">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
, male embolus is flattened, twisted, with strong distal (A), sub-distal (SA), and retrolateral (R) keels (
|
||
<figureCitation id="CE902A3AFF9D556CFC7F73D7FC0F4582" box="[930,1014,188,212]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="5" pageNumber="6">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A726EFF2F444B" blockId="5.[151,1437,261,2012]" box="[151,214,261,285]" pageId="5" pageNumber="6">Legs:</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A724CFBF74741" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
19. Scopula division on tar. III: (0) divided; (1) undivided. Length = 2, CI = 0.5, RI = 0.89. The conditions “divided/undivided tarsal scopula” have been used extensively throughout history of classification of Mygalomorphae but, herein, is applied only to tar. III. Raven discussed tarsal scopula division in regard to “Anterior/Posterior Gradations” (
|
||
<bibRefCitation id="323A4B4EFF9D556CFD9C72FFFD2844FA" author="Raven" box="[577,721,404,428]" pageId="5" pageNumber="6" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven 1985</bibRefCitation>
|
||
);
|
||
<bibRefCitation id="323A4B4EFF9D556CFD3572FFFC3A44FA" author="Guadanucci" box="[744,963,404,428]" pageId="5" pageNumber="6" refString="Guadanucci, J. P. L. (2005) Tarsal scopula significance in Ischnocolinae phylogenetics (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology, 33, 456 - 467." type="journal article" year="2005">Guadanucci (2005)</bibRefCitation>
|
||
followed from Raven, both concluded a dependency for tarsal scopula division with most developed scopulation in anterior legs, grading to weaker conditions in hind legs. Furthermore,
|
||
<bibRefCitation id="323A4B4EFF9D556CFDAA72B7FCA844A2" author="Guadanucci" box="[631,849,476,500]" pageId="5" pageNumber="6" refString="Guadanucci, J. P. L. (2005) Tarsal scopula significance in Ischnocolinae phylogenetics (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology, 33, 456 - 467." type="journal article" year="2005">Guadanucci (2005)</bibRefCitation>
|
||
concluded tarsal scopula division is not dependant upon size and is useful in phylogenetic analysis (contra
|
||
<bibRefCitation id="323A4B4EFF9D556CFCEA7294FBF84741" author="Perez-Miles" box="[823,1025,511,535]" pageId="5" pageNumber="6" refString="Perez-Miles, F. (1994) Tarsal scopula division in Theraphosinae (Araneae, Theraphosidae): its systematic significance. Journal of Arachnology, 22, 46 - 53." type="journal article" year="1994">Pérez-Miles 1994</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A714FFE4047B9" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
20. Scopula division on met. IV: (0) divided; (1) undivided. Length = 1, CI = 1, RI = 0.
|
||
<bibRefCitation id="323A4B4EFF9D556CFBAE714EFEBA4709" author="West" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn (2010b, figs 19, 69, 74)</bibRefCitation>
|
||
, following
|
||
<bibRefCitation id="323A4B4EFF9D556CFE1B712CFD8B4709" author="Pocock" box="[454,626,583,607]" pageId="5" pageNumber="6" refString="Pocock, R. I. (1895 a) On a new and natural grouping of some of the Oriental genera of Mygalomorphae, with descriptions of new genera and species. The Annals and Magazine of Natural History, (6) 15, 165 - 184." type="journal article" year="1895" yearSuffix="a">Pocock (1895a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="323A4B4EFF9D556CFDA2712CFD3D4709" author="Pocock" box="[639,708,583,607]" pageId="5" pageNumber="6" refString="Pocock, R. I. (1900 a) The fauna of British India, including Ceylon and Burma. Arachnida. London, 279 pp." type="book" year="1900" yearSuffix="a">1900a</bibRefCitation>
|
||
) used undivided met. IV scopula to diagnose most
|
||
<taxonomicName id="91AB4D3CFF9D556CFACD7123FA654736" box="[1296,1436,584,608]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFACD7123FA654736" box="[1296,1436,584,608]" italics="true" pageId="5" pageNumber="6">Lyrognathus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<bibRefCitation id="323A4B4EFF9D556CFEFB7107FE3E47D2" author="Gallon" box="[294,455,620,644]" pageId="5" pageNumber="6" refString="Gallon, R. C. (2003) A new African arboreal genus and species of theraphosidae spider (Araneae, Theraphosidae, Stromatopelminae) which lacks spermathecae. Bulletin of the British Arachnological Society, 12, 405 - 411." type="journal article" year="2003">Gallon (2003)</bibRefCitation>
|
||
considered undivided met. IV scopula apomorphic for Eumenophorinae.
|
||
<bibRefCitation id="323A4B4EFF9D556CFACB7107FEF647F1" author="Guadanucci" pageId="5" pageNumber="6" refString="Guadanucci, J. P. L. (2005) Tarsal scopula significance in Ischnocolinae phylogenetics (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology, 33, 456 - 467." type="journal article" year="2005">Guadanucci (2005)</bibRefCitation>
|
||
concluded, in his work on tarsal scopula division, that dependency of tarsal scopula condition between legs means the more entire scopulate condition, the more apomorphic states it will show. This would also apply to metatarsal scopula.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A7197FE754629" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
21. Scopula on met. IV retrolaterally stronger: (0) absent; (1) present. Length = 1, CI = 1, RI = 0.
|
||
<bibRefCitation id="323A4B4EFF9D556CFB227196FEE74661" author="West" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn (2010b)</bibRefCitation>
|
||
first noted the presence of scopula on retrolateral side of division on met IV as almost entire, with prolateral side only extending 1/3 to 2/3 basally in
|
||
<taxonomicName id="91AB4D3CFF9D556CFD27702EFC50460A" box="[762,937,837,860]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFD27702EFC50460A" box="[762,937,837,860]" italics="true" pageId="5" pageNumber="6">Coremiocnemis</emphasis>
|
||
</taxonomicName>
|
||
(group 1) species (
|
||
<bibRefCitation id="323A4B4EFF9D556CFBA0702FFE794629" author="West" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn 2010b: figs 21, 87, 163, 206</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A70E7FD9E4159" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
22. Met. IV scopula divided by strong diagonal line: (0) absent; (1) present. Length = 1, CI = 1, RI = 0. Both
|
||
<taxonomicName id="91AB4D3CFF9D556CFF1870DBFE1D469E" box="[197,484,944,968]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kotacana">
|
||
<emphasis id="64DFEAADFF9D556CFF1870DBFE1D469E" box="[197,484,944,968]" italics="true" pageId="5" pageNumber="6">Coremiocnemis kotacana</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="91AB4D3CFF9D556CFD9570DBFD0E4691" box="[584,759,944,967]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFD9570DBFD0E4691" box="[584,759,944,967]" italics="true" pageId="5" pageNumber="6">Coremiocnemis</emphasis>
|
||
</taxonomicName>
|
||
sp. from Borneo, used within our data matrix, possess met IV scopula divided and separated distally by strong dividing lines; there is no gradual diminishing of scopula (
|
||
<bibRefCitation id="323A4B4EFF9D556CFF11709CFDA34159" author="West" box="[204,602,1015,1040]" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn 2010b: figs 165, 206</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A7777FE2A4192" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
23. Scopula on tarsi laterally extended: (0) absent (
|
||
<figureCitation id="CE902A3AFF9D556CFD137777FCB74162" box="[718,846,1052,1076]" captionStart="FIGURES 28 – 36. 28 a – b" captionStartId="24.[151,262,1692,1715]" captionTargetBox="[167,1432,190,1691]" captionTargetId="figure@24.[151,1436,177,1693]" captionTargetPageId="24" captionText="FIGURES 28 – 36. 28 a – b: a. Poecilotheria tigrinawesseli Smith 2006, male, embolus, retrolateral view. b. P. tigrinawesseli, female, spermatheca, dorsal view. 29. Phlogiellus watasei (Kishida 1920), female, dorsal habitus. 30. Chilobrachys fimbriatus Pocock 1899, holotype female, NHM 1899.11. 2.22, spermatheca, dorsal view. 31 a – b: a. Chilobrachys andersoni (Pocock 1895), paratype male, NHM 1895.9. 21.21, embolus, retrolateral view. b. Chilobrachys andersoni, holotype female, NHM 1891.10. 15.6, spermatheca, dorsal view. 32. Haplocosmia nepalensis Schmidt & von Wirth 1996, male, QM S 83827, embolus, retrolateral view. 33. Phlogiellus atriceps Pocock 1897, paratype female, NHM 1897.11. 1.6 – 7, spermatheca, dorsal view. 34. Phlogiellus baeri (Simon 1877), male, QM S 83789, embolus, retrolateral view. 35 a – c: a. Orphnaecus pellitus Simon 1892, syntype male, MNHN AR 4678, embolus, retrolateral view. b. Orphnaecus pellitus, paratype female, MNHN AR 4678, spermatheca, dorsal view. c. Orphnaecus pellitus, female, dorsal habitus. 36. Orphnaecus philippinus (Schmidt 1999), female, QM S 83833, spermatheca, dorsal view. Abbreviations: A—apical keel, BL—basal lobe, PI—prolateral inferior keel, PS—prolateral superior keel, Rretrolateral keel, RCR—reverse corkscrew retrolateral keel, SA—sub-apical keel. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280927/files/figure.png" pageId="5" pageNumber="6">Figs 29, 35</figureCitation>
|
||
c, 38c, 42,); (1) present (
|
||
<figureCitation id="CE902A3AFF9D556CFB827777FB4A4162" box="[1119,1203,1052,1076]" captionStart="FIGURES 46 – 48" captionStartId="26.[151,264,1605,1628]" captionTargetBox="[151,1436,193,1584]" captionTargetId="figure@26.[151,1436,193,1584]" captionTargetPageId="26" captionText="FIGURES 46 – 48. Poecilotheria species. 46. P. metallica Pocock 1899, female, in life, Nallamala Range, Pradesh, India, dorsal view. 47. P. f o r m o s a Pocock 1899, holotype female, NHM 1898.10.10.1 – 2, right maxillae (inverted), prolateral view. 48. P. metallica, female, QM S 88462, left strikers, retrolateral view. Photos: 46 R. West; 47 D. Court; 48 S. Nunn." httpUri="https://zenodo.org/record/280929/files/figure.png" pageId="5" pageNumber="6">Fig. 46</figureCitation>
|
||
). Length = 1, CI = 1, RI = 0. Following West
|
||
<emphasis id="64DFEAADFF9D556CFE09772AFDFC410E" box="[468,517,1088,1112]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||
. (2008: character 9, state 3).
|
||
<bibRefCitation id="323A4B4EFF9D556CFC90772BFB204101" author="Raven" box="[845,1241,1087,1111]" pageId="5" pageNumber="6" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven (1985: table 3, character 17)</bibRefCitation>
|
||
utilized a similar character in his cladistic analysis of
|
||
<taxonomicName id="91AB4D3CFF9D556CFD55770FFCD4412A" box="[648,813,1124,1148]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Theraphosidae</taxonomicName>
|
||
subfamilies, “
|
||
<emphasis id="64DFEAADFF9D556CFC02770EFEEF41F6" italics="true" pageId="5" pageNumber="6">tarsi broader than metatarsi: absent/ present</emphasis>
|
||
”, a trait he found synapomorphic for
|
||
<taxonomicName id="91AB4D3CFF9D556CFD3677ECFC7C41C9" box="[747,901,1159,1183]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Aviculariinae">Aviculariinae</taxonomicName>
|
||
. This character is noted in all
|
||
<taxonomicName id="91AB4D3CFF9D556CFAD977E3FA6541F6" box="[1284,1436,1160,1184]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFAD977E3FA6541F6" box="[1284,1436,1160,1184]" italics="true" pageId="5" pageNumber="6">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
considered for analyses.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A77BBFEA241B1" blockId="5.[151,1437,261,2012]" box="[151,347,1231,1256]" pageId="5" pageNumber="6">Leg morphology:</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A779FFBF04021" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
24. Width of Tib. IV in relation to Tib. I: (0) similar to, or smaller than (
|
||
<figureCitation id="CE902A3AFF9D556CFC1E779FFBEE405A" box="[963,1047,1268,1292]" captionStart="FIGURES 14 – 20" captionStartId="19.[151,264,1446,1469]" captionTargetBox="[151,1436,155,1424]" captionTargetId="figure@19.[151,1436,155,1424]" captionTargetPageId="19" captionText="FIGURES 14 – 20. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 14. Left intercheliceral pegs, prolateral view. 15. Left proximomedial retrolateral cheliceral spine cluster. 16. Close up of intercheliceral pegs. 17. Left patella and tibia IV, dorsal view. 18. Left metatarsus and tarsus IV, prolateral view. 19. Left metatarsus and tarsus IV, retrolateral view. 20. Left metatarsus and tarsus IV, ventral view. Scale bar = 10 mm for 17 – 19; 5 mm for 20; 2 mm for 14 and 15; 1 mm for 16. Photos: S. Nunn." httpUri="https://zenodo.org/record/280923/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
|
||
); (1) incrassate, much thicker than (
|
||
<figureCitation id="CE902A3AFF9D556CFF10767CFED84079" box="[205,289,1303,1327]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="5" pageNumber="6">Fig. 38</figureCitation>
|
||
c). Length = 1, CI = 1, RI = 0.
|
||
<bibRefCitation id="323A4B4EFF9D556CFDAA767CFCD84079" author="Pocock" box="[631,801,1303,1327]" pageId="5" pageNumber="6" refString="Pocock, R. I. (1895 a) On a new and natural grouping of some of the Oriental genera of Mygalomorphae, with descriptions of new genera and species. The Annals and Magazine of Natural History, (6) 15, 165 - 184." type="journal article" year="1895" yearSuffix="a">Pocock (1895a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="323A4B4EFF9D556CFCF1767CFC8B4079" author="Pocock" box="[812,882,1303,1327]" pageId="5" pageNumber="6" refString="Pocock, R. I. (1900 a) The fauna of British India, including Ceylon and Burma. Arachnida. London, 279 pp." type="book" year="1900" yearSuffix="a">1900a</bibRefCitation>
|
||
) used incrassate tib. IV to diagnose
|
||
<taxonomicName id="91AB4D3CFF9D556CFAD47673FA6C4066" box="[1289,1429,1304,1328]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFAD47673FA6C4066" box="[1289,1429,1304,1328]" italics="true" pageId="5" pageNumber="6">Lyrognathus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="323A4B4EFF9D556CFF187657FEA74002" author="Smith" box="[197,350,1340,1364]" pageId="5" pageNumber="6" refString="Smith, A. M. (1988) Lyrognathus robustus, a new species of theraphosid spider from Malaysia. British Tarantula Society Journal, 4 (2), 15 - 19." type="journal article" year="1988">Smith (1988)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="323A4B4EFF9D556CFEB17656FE0F4002" author="West" box="[364,502,1340,1364]" pageId="5" pageNumber="6" refString="West, R. C. (1991) A new theraphosid spider from the Cameron Highlands, Malaysia. Memoirs of the Queensland Museum, 30, 615 - 619." type="journal article" year="1991">West (1991)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="323A4B4EFF9D556CFDEC7656FCD24002" author="West" box="[561,811,1340,1364]" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn (2010a</bibRefCitation>
|
||
: fig. 38c) followed
|
||
<bibRefCitation id="323A4B4EFF9D556CFBD37657FB3D4002" author="Pocock" box="[1038,1220,1340,1364]" pageId="5" pageNumber="6" refString="Pocock, R. I. (1895 a) On a new and natural grouping of some of the Oriental genera of Mygalomorphae, with descriptions of new genera and species. The Annals and Magazine of Natural History, (6) 15, 165 - 184." type="journal article" year="1895" yearSuffix="a">Pocock (1895a)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="323A4B4EFF9D556CFB0C7657FEF34021" author="Perez-Miles" pageId="5" pageNumber="6" refString="Perez-Miles, F., Lucas, S. M., da Silva Jr., P. I. & Bertani, R. (1996) Systematic revision and cladistic analysis of Theraphosinae (Araneae: Theraphosidae). Mygalomorph, 1, 33 - 68." type="journal article" year="1996">
|
||
Pérez-Miles
|
||
<emphasis id="64DFEAADFF9D556CFABF7656FA654002" box="[1378,1436,1340,1364]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
|
||
(1996
|
||
</bibRefCitation>
|
||
: character 16) used this character in their analysis of
|
||
<taxonomicName id="91AB4D3CFF9D556CFC807634FBFF4021" box="[861,1030,1375,1399]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Theraphosinae">Theraphosinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A76EFFE0640B2" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
25. Leg spines on tibia: (0) present; (1) absent. Length = 1, CI = 1, RI = 0.
|
||
<bibRefCitation id="323A4B4EFF9D556CFC2076EEFB6E40CA" author="Raven" box="[1021,1175,1412,1436]" pageId="5" pageNumber="6" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven (1985)</bibRefCitation>
|
||
and West
|
||
<emphasis id="64DFEAADFF9D556CFACC76EEFABA40CA" box="[1297,1347,1412,1436]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
|
||
. (2008: character 11) discussed leg spination in
|
||
<taxonomicName id="91AB4D3CFF9D556CFD5C76CCFCEC40E9" box="[641,789,1447,1471]" class="Arachnida" family="Barychelidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Barychelidae</taxonomicName>
|
||
and considered reductions in leg spination to be a derived condition in
|
||
<taxonomicName id="91AB4D3CFF9D556CFE8F76A7FE0240B2" box="[338,507,1484,1508]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Theraphosidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A7684FEB24319" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
26. Number of clavate trichobothria on tar. IV: (0) under 25; (1) 25 and over. Length = 2, CI = 0.5, RI = 0.91.
|
||
<bibRefCitation id="323A4B4EFF9D556CFF18757EFD93437A" author="Raven" box="[197,618,1556,1580]" pageId="5" pageNumber="6" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven (1985: table 1, character 37)</bibRefCitation>
|
||
considered reduction/loss of tarsal trichobothria a synapomorphy for Atypoidina.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A7537FACE4252" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
27. Position of clavates on tar. IV: (0) proximal 2/3 to 3/
|
||
<quantity id="91539B5AFF9D556CFCC57537FCBD4322" box="[792,836,1628,1652]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="6" unit="in" value="4.0">4 in</quantity>
|
||
straight line; (1) proximal 1/2 or less in straight line; (2) In proximal 1/
|
||
<quantity id="91539B5AFF9D556CFE4F7514FE4643C1" box="[402,447,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="6" unit="in" value="3.0">3 in</quantity>
|
||
"U" shape. Length = 3, CI = 0.66, RI = 0.91.
|
||
<bibRefCitation id="323A4B4EFF9D556CFC0B75EBFB8943C1" author="Raven" box="[982,1136,1663,1687]" pageId="5" pageNumber="6" refString="Raven, R. J. (1985) The spider infraorder mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182, 1 - 180." type="journal article" year="1985">Raven (1985)</bibRefCitation>
|
||
discussed variation of leg trichobothria in Mygalomorphae and used those variations in his systematic revision of the infraorder. Among
|
||
<taxonomicName id="91AB4D3CFF9D556CFF1875ACFE7A4389" box="[197,387,1735,1759]" class="Arachnida" family="Theraphosidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Selenocosmiinae">Selenocosmiinae</taxonomicName>
|
||
examined, several clavate trichobothrial conditions were noted. Additionally, Guadanucci (in prep.) noted the clavate arrangement in
|
||
<taxonomicName id="91AB4D3CFF9D556CFD5F7587FCE24252" box="[642,795,1772,1796]" class="Arachnida" family="Theraphosidae" genus="Poecilotheria" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFD5F7587FCE24252" box="[642,795,1772,1796]" italics="true" pageId="5" pageNumber="6">Poecilotheria</emphasis>
|
||
</taxonomicName>
|
||
(Guadanucci, in prep.: fig. 167) as our state (2).
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556CFF4A7464FF0742C2" blockId="5.[151,1437,261,2012]" pageId="5" pageNumber="6">
|
||
28. Brush of penicillate setae on retrolateral tib. IV (
|
||
<figureCitation id="CE902A3AFF9D556CFD317464FCB94271" box="[748,832,1807,1831]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="5" pageNumber="6">Fig. 38</figureCitation>
|
||
c): (0) absent; (1) present. Length = 1, CI = 1, RI = 0.
|
||
<bibRefCitation id="323A4B4EFF9D556CFF18745EFE42421A" author="West" box="[197,443,1844,1868]" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn (2010a)</bibRefCitation>
|
||
used the presence of penicillate setae along retrolateral surfaces of tib. IV to diagnose all
|
||
<taxonomicName id="91AB4D3CFF9D556CFF187433FEA84226" box="[197,337,1880,1904]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFF187433FEA84226" box="[197,337,1880,1904]" italics="true" pageId="5" pageNumber="6">Lyrognathus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9D556CFE82743CFCBD4239" author="West" box="[351,836,1879,1903]" pageId="5" pageNumber="6" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a: figs 8, 20, 53, 71, 98</bibRefCitation>
|
||
). We introduce this character for analysis for the first time.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9D556FFF4A74CBFE7A45A1" blockId="5.[151,1437,261,2012]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
29. Brush of recurved setae along retrolateral met. IV (
|
||
<figureCitation id="CE902A3AFF9D556CFCDE74CBFCA142EE" box="[771,856,1952,1976]" captionStart="FIGURES 37 – 45. 37 a – b" captionStartId="25.[151,264,1661,1684]" captionTargetBox="[163,1431,198,1651]" captionTargetId="figure@25.[151,1436,193,1659]" captionTargetPageId="25" captionText="FIGURES 37 – 45. 37 a – b: a. Lyrognathus crotalus Pocock 1895, male, QM S 88361, embolus, retrolateral view. b. Lyrognathus crotalus, female, QM S 88362, spermatheca, dorsal view. 38 a – c: a. Lyrognathus lessunda West & Nunn 2010, paratype male, QM S 74066, embolus, retrolateral view. b. Lyrognathus lessunda, holotype female, QM S 74067, spermatheca, dorsal view. c. Lyrognathus lessunda, holotype female, QM S 74067, habitus, dorsal view. 39. Psednocnemis davidgohi sp. nov., paratype male, QM S 88022, embolus, retrolateral view. 40. Psednocnemis jeremyhuffi (West & Nunn 2010), paratype female, QM S 88071, spermatheca, dorsal view. 41. Coremiocnemis obscura West & Nunn 2010, male, QM S 88360, embolus, retrolateral view. 42. Coremiocnemis hoggi West & Nunn 2010, paratype female, QM S 74074, dorsal habitus. 43. Coremiocnemis kotacana West & Nunn 2010, holotype female, QM S 83759, spermatheca, dorsal view. 44 a – b: a. Selenocosmia javanensis (Walckenaer 1837), male, NHMW 173 embolus, retrolateral view. b. Selenocosmia javanensis, female, QM S 88355, spermatheca, dorsal view. 45. Selenocosmia species (A), female, RMBR ZRC. ARA. 02, spermatheca, dorsal view. Abbreviations: AC—apical spiral curl, LTE—lanceolate terete embolus, RBL—reduced basal lobe, RR—reduced retrolateral keel, TE—terete embolus. Illustrations: S. Nunn." httpUri="https://zenodo.org/record/280928/files/figure.png" pageId="5" pageNumber="6">Fig. 42</figureCitation>
|
||
): (0) absent; (1) present. Length = 1, CI = 1, RI = 0. West & Nunn noted in
|
||
<taxonomicName id="91AB4D3CFF9D556CFE1B74AEFD8C428A" box="[454,629,1989,2012]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9D556CFE1B74AEFD8C428A" box="[454,629,1989,2012]" italics="true" pageId="5" pageNumber="6">Coremiocnemis</emphasis>
|
||
</taxonomicName>
|
||
(group 1) the presence of recurved setae in brushform along retrolateral surfaces of met. IV (
|
||
<bibRefCitation id="323A4B4EFF9E556FFE6D73FCFC6745F9" author="West" box="[432,926,151,175]" pageId="6" pageNumber="7" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn 2010b: figs 21, 87, 165, 206</bibRefCitation>
|
||
). The description of
|
||
<taxonomicName id="91AB4D3CFF9E556FFB5173F3FAD545E6" box="[1164,1324,152,176]" class="Arachnida" family="Theraphosidae" genus="Psednocnemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="64DFEAADFF9E556FFB5173F3FAD545E6" box="[1164,1324,152,176]" italics="true" pageId="6" pageNumber="7">Psednocnemis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FEC57D6FF9E556FFAE873F3FA6545F9" box="[1333,1436,152,176]" pageId="6" pageNumber="7" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
indicates this characteristic is diagnostic to
|
||
<taxonomicName id="91AB4D3CFF9E556FFD7073D6FCA54582" box="[685,860,189,212]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFD7073D6FCA54582" box="[685,860,189,212]" italics="true" pageId="6" pageNumber="7">Coremiocnemis</emphasis>
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="91AB4D3CFF9E556FFC6173D6FB574582" authority="Raven 2005" authorityName="Raven" authorityYear="2005" box="[956,1198,188,212]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tropix">
|
||
<emphasis id="64DFEAADFF9E556FFC6173D6FBE64582" box="[956,1055,189,212]" italics="true" pageId="6" pageNumber="7">C. tropix</emphasis>
|
||
<bibRefCitation id="323A4B4EFF9E556FFBFB73D6FB574582" author="Raven" box="[1062,1198,188,212]" pageId="6" pageNumber="7" refString="Raven, R. J. (2005) A new tarantula species from northern Australia (Araneae, Theraphosidae). Zootaxa, 1004, 15 - 28." type="journal article" year="2005">Raven 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
) which is tested here for the first time.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9E556FFF4A726FFB76444A" blockId="6.[151,1436,151,285]" box="[151,1167,260,285]" pageId="6" pageNumber="7">30. Leg width formula IV, III, I, II: (0) absent; (1) present. Length = 2, CI = 0.5, RI = 0.83.</paragraph>
|
||
<paragraph id="561436BFFF9E556FFF4A7227FEEE4432" blockId="6.[151,1437,332,716]" box="[151,279,332,356]" pageId="6" pageNumber="7">Chelicerae:</paragraph>
|
||
<paragraph id="561436BFFF9E556FFF4A7204FB6D4499" blockId="6.[151,1437,332,716]" pageId="6" pageNumber="7">
|
||
31. Intercheliceral setal fringe proximodorsally: (0) absent (
|
||
<figureCitation id="CE902A3AFF9E556FFCE77204FC7744D1" box="[826,910,367,391]" captionStart="FIGURES 14 – 20" captionStartId="19.[151,264,1446,1469]" captionTargetBox="[151,1436,155,1424]" captionTargetId="figure@19.[151,1436,155,1424]" captionTargetPageId="19" captionText="FIGURES 14 – 20. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 14. Left intercheliceral pegs, prolateral view. 15. Left proximomedial retrolateral cheliceral spine cluster. 16. Close up of intercheliceral pegs. 17. Left patella and tibia IV, dorsal view. 18. Left metatarsus and tarsus IV, prolateral view. 19. Left metatarsus and tarsus IV, retrolateral view. 20. Left metatarsus and tarsus IV, ventral view. Scale bar = 10 mm for 17 – 19; 5 mm for 20; 2 mm for 14 and 15; 1 mm for 16. Photos: S. Nunn." httpUri="https://zenodo.org/record/280923/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
|
||
); (1) present (
|
||
<figureCitation id="CE902A3AFF9E556FFBF67204FB8744D1" box="[1067,1150,367,391]" captionStart="FIGURES 49 a – 50 b. 49 a, b" captionStartId="27.[151,264,1127,1150]" captionTargetBox="[151,1436,193,1104]" captionTargetId="figure@27.[151,1436,193,1106]" captionTargetPageId="27" captionText="FIGURES 49 a – 50 b. 49 a, b. Haplocosmia himalayana (Pocock 1899), holotype female, NHM 1899.8. 19.3. 49 a. Right maxillae (inverted), prolateral view. 49 b. Right chelicerae, prolateral view. 50 a, b. Chilobrachys sericeus (Thorell 1895), syntype male, NHM 1895.9. 21.21. 50 a. Left maxillae, prolateral view. 50 b. Left strikers, retrolateral view. Photos: R. Raven." httpUri="https://zenodo.org/record/280930/files/figure.png" pageId="6" pageNumber="7">Fig. 49</figureCitation>
|
||
b). Length = 1, CI = 1, RI = 0. In
|
||
<taxonomicName id="91AB4D3CFF9E556FFECB72FFFE5244FA" box="[278,427,404,428]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFECB72FFFE5244FA" box="[278,427,404,428]" italics="true" pageId="6" pageNumber="7">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
, a distinct fringe of long pallid setae is present, intermixed with intercheliceral pegs. This fringe has not been seen outside
|
||
<taxonomicName id="91AB4D3CFF9E556FFDEF72D3FD3E4486" box="[562,711,440,464]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFDEF72D3FD3E4486" box="[562,711,440,464]" italics="true" pageId="6" pageNumber="7">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
, and is considered here for the first time.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9E556FFF4A72B7FC99476A" blockId="6.[151,1437,332,716]" pageId="6" pageNumber="7">
|
||
32. Intercheliceral pegs with prolateral keeled ridge (
|
||
<figureCitation id="CE902A3AFF9E556FFD2D72B7FCBA44A2" box="[752,835,476,500]" captionStart="FIGURES 49 a – 50 b. 49 a, b" captionStartId="27.[151,264,1127,1150]" captionTargetBox="[151,1436,193,1104]" captionTargetId="figure@27.[151,1436,193,1106]" captionTargetPageId="27" captionText="FIGURES 49 a – 50 b. 49 a, b. Haplocosmia himalayana (Pocock 1899), holotype female, NHM 1899.8. 19.3. 49 a. Right maxillae (inverted), prolateral view. 49 b. Right chelicerae, prolateral view. 50 a, b. Chilobrachys sericeus (Thorell 1895), syntype male, NHM 1895.9. 21.21. 50 a. Left maxillae, prolateral view. 50 b. Left strikers, retrolateral view. Photos: R. Raven." httpUri="https://zenodo.org/record/280930/files/figure.png" pageId="6" pageNumber="7">Fig. 49</figureCitation>
|
||
b): (0) absent; (1) present. Length = 1, CI = 1, RI = 0. In
|
||
<taxonomicName id="91AB4D3CFF9E556FFF38716BFE83474E" box="[229,378,512,536]" class="Arachnida" family="Theraphosidae" genus="Haplocosmia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFF38716BFE83474E" box="[229,378,512,536]" italics="true" pageId="6" pageNumber="7">Haplocosmia</emphasis>
|
||
</taxonomicName>
|
||
, and only within that selenocosmiine genus, intercheliceral pegs possess a distinct prolateral keel. As in character 31, this trait is tested for the first time.
|
||
</paragraph>
|
||
<paragraph id="561436BFFF9E556FFF4A712CFBC9479A" blockId="6.[151,1437,332,716]" pageId="6" pageNumber="7">
|
||
33. Spine cluster on retrolateral proximomedial chelicerae (
|
||
<figureCitation id="CE902A3AFF9E556FFCE6712CFC764709" box="[827,911,583,607]" captionStart="FIGURES 14 – 20" captionStartId="19.[151,264,1446,1469]" captionTargetBox="[151,1436,155,1424]" captionTargetId="figure@19.[151,1436,155,1424]" captionTargetPageId="19" captionText="FIGURES 14 – 20. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 14. Left intercheliceral pegs, prolateral view. 15. Left proximomedial retrolateral cheliceral spine cluster. 16. Close up of intercheliceral pegs. 17. Left patella and tibia IV, dorsal view. 18. Left metatarsus and tarsus IV, prolateral view. 19. Left metatarsus and tarsus IV, retrolateral view. 20. Left metatarsus and tarsus IV, ventral view. Scale bar = 10 mm for 17 – 19; 5 mm for 20; 2 mm for 14 and 15; 1 mm for 16. Photos: S. Nunn." httpUri="https://zenodo.org/record/280923/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
): (0) absent; (1) present. Length = 2, CI = 0.5, RI = 0.94. West & Nunn first noted retrolateral cheliceral spines in
|
||
<taxonomicName id="91AB4D3CFF9E556FFC657107FBBD47D2" box="[952,1092,620,644]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFC657107FBBD47D2" box="[952,1092,620,644]" italics="true" pageId="6" pageNumber="7">Lyrognathus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9E556FFB8F7107FEC447F1" author="West" pageId="6" pageNumber="7" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a: figs 26, 60, 80, 96</bibRefCitation>
|
||
),
|
||
<taxonomicName id="91AB4D3CFF9E556FFE8971FBFDFA47F1" box="[340,515,656,679]" class="Arachnida" family="Theraphosidae" genus="Coremiocnemis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFE8971FBFDFA47F1" box="[340,515,656,679]" italics="true" pageId="6" pageNumber="7">Coremiocnemis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9E556FFDC971E4FB6647F1" author="West" box="[532,1183,655,679]" pageId="6" pageNumber="7" refString="West, R. C. & Nunn, S. C. (2010 b) A taxonomic revision of the tarantula spider genus Coremiocnemis Simon 1892 (Araneae, Theraphosidae), with further notes on the Selenocosmiinae. Zootaxa, 2443, 1 - 64." type="journal article" year="2010" yearSuffix="b">West & Nunn 2010b: figs 9, 47, 69, 107, 170, 189, 215</bibRefCitation>
|
||
) and in
|
||
<taxonomicName id="91AB4D3CFF9E556FFADF71FBFA6547FE" box="[1282,1436,656,680]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFADF71FBFA6547FE" box="[1282,1436,656,680]" italics="true" pageId="6" pageNumber="7">Selenocosmia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="323A4B4EFF9E556FFF1171DFFE41479A" author="West" box="[204,440,692,716]" pageId="6" pageNumber="7" refString="West, R. C. & Nunn, S. C. (2010 a) A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae. Zootaxa, 2362, 1 - 43." type="journal article" year="2010" yearSuffix="a">West & Nunn 2010a</bibRefCitation>
|
||
). The same feature is present in
|
||
<taxonomicName id="91AB4D3CFF9E556FFCFD71DFFC39479A" box="[800,960,692,716]" class="Arachnida" family="Theraphosidae" genus="Psednocnemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="64DFEAADFF9E556FFCFD71DFFC39479A" box="[800,960,692,716]" italics="true" pageId="6" pageNumber="7">Psednocnemis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CE902A3AFF9E556FFC0D71DFFBDA479A" box="[976,1059,692,716]" captionStart="FIGURES 14 – 20" captionStartId="19.[151,264,1446,1469]" captionTargetBox="[151,1436,155,1424]" captionTargetId="figure@19.[151,1436,155,1424]" captionTargetPageId="19" captionText="FIGURES 14 – 20. Psednocnemis davidgohi sp. nov., holotype female, QM S 88303. 14. Left intercheliceral pegs, prolateral view. 15. Left proximomedial retrolateral cheliceral spine cluster. 16. Close up of intercheliceral pegs. 17. Left patella and tibia IV, dorsal view. 18. Left metatarsus and tarsus IV, prolateral view. 19. Left metatarsus and tarsus IV, retrolateral view. 20. Left metatarsus and tarsus IV, ventral view. Scale bar = 10 mm for 17 – 19; 5 mm for 20; 2 mm for 14 and 15; 1 mm for 16. Photos: S. Nunn." httpUri="https://zenodo.org/record/280923/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |