treatments-xml/data/DB/1A/DF/DB1ADF00F93457ECA7A3B7E8116910DB.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1120.87067" ID-Pensoft-Pub="1313-2970-1120-47" ID-Pensoft-UUID="8013346308A35B09859E2A0B0F137A0E" ID-ZooBank="398862C6FCA84B0FB084F34D8B86EA1B" ModsDocID="1313-2970-1120-47" checkinTime="1662465956718" checkinUser="pensoft" docAuthor="Lin, Shi-Shi, Li, Yuan-Hang, Su, Hong-Lin, Yi, Hui, Pan, Zhong, Sun, Yan-Jun, Zeng, Zhao-Chi &amp; Wang, Jian" docDate="2022" docId="DB1ADF00F93457ECA7A3B7E8116910DB" docLanguage="en" docName="ZooKeys 1120: 47-66" docOrigin="ZooKeys 1120" docPubDate="2022-09-05" docSource="http://dx.doi.org/10.3897/zookeys.1120.87067" docTitle="Odorrana concelata Wang, Zeng, &amp; Lin 2022, sp. nov." docType="treatment" docUuid="63E81BE8-F60F-47F4-B49A-3403C4AB82D3" docUuidSource="ZooBank" docVersion="2" id="8013346308A35B09859E2A0B0F137A0E" lastPageNumber="47" masterDocId="8013346308A35B09859E2A0B0F137A0E" masterDocTitle="Discovery of a new limestone karst-restricted odorous frog from northern Guangdong, China (Anura, Ranidae, Odorrana)" masterLastPageNumber="66" masterPageNumber="47" pageNumber="47" updateTime="1662466258105" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Discovery of a new limestone karst-restricted odorous frog from northern Guangdong, China (Anura, Ranidae, Odorrana)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Shi-Shi</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Yuan-Hang</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Su, Hong-Lin</mods:namePart>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yi, Hui</mods:namePart>
<mods:affiliation>Department of Ecology and Environment of Guangdong Province, Guangzhou 510630, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pan, Zhong</mods:namePart>
<mods:affiliation>Guangzhou Shengheng Forestry Co., LTD, Guangzhou 510663, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sun, Yan-Jun</mods:namePart>
<mods:affiliation>Shenzhen Landscape Co., LTD, Shenzhen 518001, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zeng, Zhao-Chi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4054-8399</mods:nameIdentifier>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
<mods:nameIdentifier type="email">zengzhch5@mail2.sysu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Jian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4249-7767</mods:nameIdentifier>
<mods:affiliation>Guangdong Polytechnic of Environmental Protection Engineering, Foshan 528216, China</mods:affiliation>
<mods:nameIdentifier type="email">wangj-1994@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1120</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>47</mods:start>
<mods:end>66</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1120.87067</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1120.87067</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1120-47</mods:identifier>
<mods:identifier type="ZooBank">398862C6FCA84B0FB084F34D8B86EA1B</mods:identifier>
<mods:identifier type="Pensoft-UUID">8013346308A35B09859E2A0B0F137A0E</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:63E81BE8-F60F-47F4-B49A-3403C4AB82D3" httpUri="http://treatment.plazi.org/id/DB1ADF00F93457ECA7A3B7E8116910DB" lastPageNumber="47" pageId="0" pageNumber="47">
<subSubSection pageId="0" pageNumber="47" type="nomenclature">
<paragraph pageId="0" pageNumber="47">
<taxonomicName LSID="https://zoobank.org/63E81BE8-F60F-47F4-B49A-3403C4AB82D3" authority="Wang, Zeng, &amp; Lin" authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata" status="sp. nov.">Odorrana concelata Wang, Zeng, &amp; Lin</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="47">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="description">
<paragraph pageId="0" pageNumber="47">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphological features of the male holotype GEP a 055 in life: A dorsolateral view B ventral view C ventral view of hand, showing nuptial pads on fingers I, II and III D ventral view of foot." figureDoi="10.3897/zookeys.1120.87067.figure3" httpUri="https://binary.pensoft.net/fig/738804" pageId="0" pageNumber="47">Moss-speckled Odorous Frog (in English) / Tai Ban Chou Wa (苔斑臭蛙 in Chinese) Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the female paratype GEP a 050 in life: A dorsolateral view B ventral view C ventral view of hand D ventral view of foot." figureDoi="10.3897/zookeys.1120.87067.figure4" httpUri="https://binary.pensoft.net/fig/738805" pageId="0" pageNumber="47">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microhabitat of Odorrana concelata sp. nov. (A, B) and the uncaptured individuals of juvenile (C), female (D), and male (E) in situ." figureDoi="10.3897/zookeys.1120.87067.figure5" httpUri="https://binary.pensoft.net/fig/738806" pageId="0" pageNumber="47">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="holotype">
<paragraph pageId="0" pageNumber="47">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="47">
GEP a055, adult male, collected by Shi-Shi Lin, Hong-Lin Su and Yuan-Hang Li on 20 April 2022 from Longlinchang Village (
<geoCoordinate degrees="24" direction="north" minutes="04" orientation="latitude" precision="15" seconds="47" value="24.079723">24°04'47&quot;N</geoCoordinate>
,
<geoCoordinate degrees="112" direction="east" minutes="40" orientation="longitude" precision="15" seconds="37" value="112.67694">112°40'37&quot;E</geoCoordinate>
; ca. 280 m a.s.l.), Jintan Town, Qingyuan City, Guangdong, China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="paratypes">
<paragraph pageId="0" pageNumber="47">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="47">Three adult males, GEP a052-054, and two adult females, GEP a050-051, the same collection data as the holotype.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="etymology">
<paragraph pageId="0" pageNumber="47">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="47">
The specific epithet,
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
, is a feminine adjective that means disguised, in reference to the highly concealed coloration of the new species in its mossy habitat.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="diagnosis">
<paragraph pageId="0" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="47">
(1) Small body size, SVL 34.0-36.8 mm in males (
<emphasis italics="true" pageId="0" pageNumber="47">n</emphasis>
= 4), SVL 41.4-46.0 mm in females (
<emphasis italics="true" pageId="0" pageNumber="47">n</emphasis>
= 2); (2) dorsolateral folds absent; (3) relative finger lengths II &lt;I &lt;IV &lt;III; (4) pectoral spines absent; (5) vocal sacs absent; (6) nuptial pads present on base of finger I, medially along inner side of fingers II and III in males; (7) eggs of females uniformed beige; (8) dorsum with mixed irregular grass green speckles and brown mottling, ventral skin of body greyish white with light brown mottling.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="comparisons">
<paragraph pageId="0" pageNumber="47">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. is phylogenetically closest to the clade composed of
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. liboensis" pageId="0" pageNumber="47" rank="species" species="liboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. liboensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum-likelihood and Bayesian inference phylogenies. Numbers at nodes indicate the bootstrap support (BS) / Bayesian posterior (BPP) for the topology. ' - ' means BS &lt;70 or BPP &lt;0.90." figureDoi="10.3897/zookeys.1120.87067.figure2" httpUri="https://binary.pensoft.net/fig/738803" pageId="0" pageNumber="47">2</figureCitation>
). However, the new taxon can be distinguished by possessing a smaller body size, SVL 34.0-36.8 mm in males and 41.4-46.0 mm in females (vs. SVL 40.7-49.8 mm in males and 51.1-60.1 mm in females of
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
; SVL 47.1-49.9 mm in males and 55.8-58.2 mm in females of
<taxonomicName lsidName="O. liboensis" pageId="0" pageNumber="47" rank="species" species="liboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. liboensis</emphasis>
</taxonomicName>
); presence of pineal body (vs. absent in
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. liboensis" pageId="0" pageNumber="47" rank="species" species="liboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. liboensis</emphasis>
</taxonomicName>
); presence of nuptial pads on base of finger I, medially along inner side of fingers II and III (vs. presence of nuptial pad on finger I in males of both
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. liboensis" pageId="0" pageNumber="47" rank="species" species="liboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. liboensis</emphasis>
</taxonomicName>
); relative finger lengths II &lt;I &lt;IV &lt;III (vs. I = II &lt;IV &lt;III in
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
); absence of conical spines on upper lip except skin of commissure of jaw (vs. presence of conical spines on entire upper lip in
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
); tibiotarsal articulation reaches to nostril (vs. reaches to anterior of eye in
<taxonomicName lsidName="O. lipuensis" pageId="0" pageNumber="47" rank="species" species="lipuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lipuensis</emphasis>
</taxonomicName>
); presence of tiny conical spines on temporal region except tympanum, skin of commissure of jaw, upper edge of eyelid, and along dorsolateral sides of body (vs. absent in
<taxonomicName lsidName="O. liboensis" pageId="0" pageNumber="47" rank="species" species="liboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. liboensis</emphasis>
</taxonomicName>
).
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. further differs from another karst-dweller
<taxonomicName lsidName="O. wuchuanensis" pageId="0" pageNumber="47" rank="species" species="wuchuanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. wuchuanensis</emphasis>
</taxonomicName>
by the smaller body size (vs. 71.1-76.5 mm in males and 75.8-99.6 mm in females), and absence pectoral spines (vs. present).
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. can be easily distinguished from
<taxonomicName lsidName="O. absita" pageId="0" pageNumber="47" rank="species" species="absita">
<emphasis italics="true" pageId="0" pageNumber="47">O. absita</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. amamiensis" pageId="0" pageNumber="47" rank="species" species="amamiensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. amamiensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. anlungensis" pageId="0" pageNumber="47" rank="species" species="anlungensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. anlungensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. aureola" pageId="0" pageNumber="47" rank="species" species="aureola">
<emphasis italics="true" pageId="0" pageNumber="47">O. aureola</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. bacboensis" pageId="0" pageNumber="47" rank="species" species="bacboensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. bacboensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. banaorum" pageId="0" pageNumber="47" rank="species" species="banaorum">
<emphasis italics="true" pageId="0" pageNumber="47">O. banaorum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. bolavensis" pageId="0" pageNumber="47" rank="species" species="bolavensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. bolavensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. cangyuanensis" pageId="0" pageNumber="47" rank="species" species="cangyuanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. cangyuanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. chapaensis" pageId="0" pageNumber="47" rank="species" species="chapaensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. chapaensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. chloronota" pageId="0" pageNumber="47" rank="species" species="chloronota">
<emphasis italics="true" pageId="0" pageNumber="47">O. chloronota</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. dulongensis" pageId="0" pageNumber="47" rank="species" species="dulongensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. dulongensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. exiliversabilis" pageId="0" pageNumber="47" rank="species" species="exiliversabilis">
<emphasis italics="true" pageId="0" pageNumber="47">O. exiliversabilis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. fengkaiensis" pageId="0" pageNumber="47" rank="species" species="fengkaiensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. fengkaiensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. geminata" pageId="0" pageNumber="47" rank="species" species="geminata">
<emphasis italics="true" pageId="0" pageNumber="47">O. geminata</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. gigatympana" pageId="0" pageNumber="47" rank="species" species="gigatympana">
<emphasis italics="true" pageId="0" pageNumber="47">O. gigatympana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. grahami" pageId="0" pageNumber="47" rank="species" species="grahami">
<emphasis italics="true" pageId="0" pageNumber="47">O. grahami</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. graminea" pageId="0" pageNumber="47" rank="species" species="graminea">
<emphasis italics="true" pageId="0" pageNumber="47">O. graminea</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. hainanensis" pageId="0" pageNumber="47" rank="species" species="hainanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. hejiangensis" pageId="0" pageNumber="47" rank="species" species="hejiangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. hejiangensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. huanggangensis" pageId="0" pageNumber="47" rank="species" species="huanggangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. huanggangensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. indeprensa" pageId="0" pageNumber="47" rank="species" species="indeprensa">
<emphasis italics="true" pageId="0" pageNumber="47">O. indeprensa</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. ichangensis" pageId="0" pageNumber="47" rank="species" species="ichangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. ichangensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. ishikawae" pageId="0" pageNumber="47" rank="species" species="ishikawae">
<emphasis italics="true" pageId="0" pageNumber="47">O. ishikawae</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. jingdongensis" pageId="0" pageNumber="47" rank="species" species="jingdongensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. junlianensis" pageId="0" pageNumber="47" rank="species" species="junlianensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. khalam" pageId="0" pageNumber="47" rank="species" species="khalam">
<emphasis italics="true" pageId="0" pageNumber="47">O. khalam</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. kweichowensis" pageId="0" pageNumber="47" rank="species" species="kweichowensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. kweichowensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. lungshengensis" pageId="0" pageNumber="47" rank="species" species="lungshengensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. lungshengensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. macrotympana" pageId="0" pageNumber="47" rank="species" species="macrotympana">
<emphasis italics="true" pageId="0" pageNumber="47">O. macrotympana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. morafkai" pageId="0" pageNumber="47" rank="species" species="morafkai">
<emphasis italics="true" pageId="0" pageNumber="47">O. morafkai</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. nanjiangensis" pageId="0" pageNumber="47" rank="species" species="nanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. nanjiangensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. nasica" pageId="0" pageNumber="47" rank="species" species="nasica">
<emphasis italics="true" pageId="0" pageNumber="47">O. nasica</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. nasuta" pageId="0" pageNumber="47" rank="species" species="nasuta">
<emphasis italics="true" pageId="0" pageNumber="47">O. nasuta</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. orba" pageId="0" pageNumber="47" rank="species" species="orba">
<emphasis italics="true" pageId="0" pageNumber="47">O. orba</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. sangzhiensis" pageId="0" pageNumber="47" rank="species" species="sangzhiensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. sangzhiensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. schmackeri" pageId="0" pageNumber="47" rank="species" species="schmackeri">
<emphasis italics="true" pageId="0" pageNumber="47">O. schmackeri</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. swinhoana" pageId="0" pageNumber="47" rank="species" species="swinhoana">
<emphasis italics="true" pageId="0" pageNumber="47">O. swinhoana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. tianmuii" pageId="0" pageNumber="47" rank="species" species="tianmuii">
<emphasis italics="true" pageId="0" pageNumber="47">O. tianmuii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. tiannanensis" pageId="0" pageNumber="47" rank="species" species="tiannanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. tiannanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. tormota" pageId="0" pageNumber="47" rank="species" species="tormota">
<emphasis italics="true" pageId="0" pageNumber="47">O. tormota</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. trankieni" pageId="0" pageNumber="47" rank="species" species="trankieni">
<emphasis italics="true" pageId="0" pageNumber="47">O. trankieni</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. utsunomiyaorum" pageId="0" pageNumber="47" rank="species" species="utsunomiyaorum">
<emphasis italics="true" pageId="0" pageNumber="47">O. utsunomiyaorum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. versabilis" pageId="0" pageNumber="47" rank="species" species="versabilis">
<emphasis italics="true" pageId="0" pageNumber="47">O. versabilis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. yentuensis" pageId="0" pageNumber="47" rank="species" species="yentuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. yentuensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. yizhangensis" pageId="0" pageNumber="47" rank="species" species="yizhangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. yizhangensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. yunnanensis</emphasis>
</taxonomicName>
, by the absence of vocal sacs (vs. present; internal vocal sacs present in
<taxonomicName lsidName="O. grahami" pageId="0" pageNumber="47" rank="species" species="grahami">
<emphasis italics="true" pageId="0" pageNumber="47">O. grahami</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. hainanensis" pageId="0" pageNumber="47" rank="species" species="hainanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. hainanensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. jingdongensis" pageId="0" pageNumber="47" rank="species" species="jingdongensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. jingdongensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. junlianensis" pageId="0" pageNumber="47" rank="species" species="junlianensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. junlianensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. yunnanensis" pageId="0" pageNumber="47" rank="species" species="yunnanensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. yunnanensis</emphasis>
</taxonomicName>
); and from
<taxonomicName lsidName="O. absita" pageId="0" pageNumber="47" rank="species" species="absita">
<emphasis italics="true" pageId="0" pageNumber="47">O. absita</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. amamiensis" pageId="0" pageNumber="47" rank="species" species="amamiensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. amamiensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. banaorum" pageId="0" pageNumber="47" rank="species" species="banaorum">
<emphasis italics="true" pageId="0" pageNumber="47">O. banaorum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. bolavensis" pageId="0" pageNumber="47" rank="species" species="bolavensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. bolavensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. exiliversabilis" pageId="0" pageNumber="47" rank="species" species="exiliversabilis">
<emphasis italics="true" pageId="0" pageNumber="47">O. exiliversabilis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. gigatympana" pageId="0" pageNumber="47" rank="species" species="gigatympana">
<emphasis italics="true" pageId="0" pageNumber="47">O. gigatympana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. graminea" pageId="0" pageNumber="47" rank="species" species="graminea">
<emphasis italics="true" pageId="0" pageNumber="47">O. graminea</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. indeprensa" pageId="0" pageNumber="47" rank="species" species="indeprensa">
<emphasis italics="true" pageId="0" pageNumber="47">O. indeprensa</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. hosii" pageId="0" pageNumber="47" rank="species" species="hosii">
<emphasis italics="true" pageId="0" pageNumber="47">O. hosii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. khalam" pageId="0" pageNumber="47" rank="species" species="khalam">
<emphasis italics="true" pageId="0" pageNumber="47">O. khalam</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. livida" pageId="0" pageNumber="47" rank="species" species="livida">
<emphasis italics="true" pageId="0" pageNumber="47">O. livida</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. leporipes" pageId="0" pageNumber="47" rank="species" species="leporipes">
<emphasis italics="true" pageId="0" pageNumber="47">O. leporipes</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. monjerai" pageId="0" pageNumber="47" rank="species" species="monjerai">
<emphasis italics="true" pageId="0" pageNumber="47">O. monjerai</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. narina" pageId="0" pageNumber="47" rank="species" species="narina">
<emphasis italics="true" pageId="0" pageNumber="47">O. narina</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. nasica" pageId="0" pageNumber="47" rank="species" species="nasica">
<emphasis italics="true" pageId="0" pageNumber="47">O. nasica</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. nasuta" pageId="0" pageNumber="47" rank="species" species="nasuta">
<emphasis italics="true" pageId="0" pageNumber="47">O. nasuta</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. orba" pageId="0" pageNumber="47" rank="species" species="orba">
<emphasis italics="true" pageId="0" pageNumber="47">O. orba</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. supranarina" pageId="0" pageNumber="47" rank="species" species="supranarina">
<emphasis italics="true" pageId="0" pageNumber="47">O. supranarina</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. tormota" pageId="0" pageNumber="47" rank="species" species="tormota">
<emphasis italics="true" pageId="0" pageNumber="47">O. tormota</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. trankieni" pageId="0" pageNumber="47" rank="species" species="trankieni">
<emphasis italics="true" pageId="0" pageNumber="47">O. trankieni</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. utsunomiyaorum" pageId="0" pageNumber="47" rank="species" species="utsunomiyaorum">
<emphasis italics="true" pageId="0" pageNumber="47">O. utsunomiyaorum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. versabilis" pageId="0" pageNumber="47" rank="species" species="versabilis">
<emphasis italics="true" pageId="0" pageNumber="47">O. versabilis</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="O. yentuensis" pageId="0" pageNumber="47" rank="species" species="yentuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. yentuensis</emphasis>
</taxonomicName>
, by the absence of dorsolateral folds (vs. present).
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. differs from the remaining seven congeners by the marked differences in dorsal and ventral coloration; the smaller body size, SVL 34.0-36.8 mm in males and 41.4-46.0 mm in females (vs. 57.2 mm in male and 66.0-71.4 mm in females in
<taxonomicName lsidName="O. kuangwuensis" pageId="0" pageNumber="47" rank="species" species="kuangwuensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. kuangwuensis</emphasis>
</taxonomicName>
, 78.0-88.0 mm in males and 93.0-113.0 mm in females in
<taxonomicName lsidName="O. margaretae" pageId="0" pageNumber="47" rank="species" species="margaretae">
<emphasis italics="true" pageId="0" pageNumber="47">O. margaretae</emphasis>
</taxonomicName>
, 85.8-91.6 mm in males and 108.7-110.1 mm in females in
<taxonomicName lsidName="O. mutschmanni" pageId="0" pageNumber="47" rank="species" species="mutschmanni">
<emphasis italics="true" pageId="0" pageNumber="47">O. mutschmanni</emphasis>
</taxonomicName>
, 80.0 mm in males and 84.3-106.0 mm in females in
<taxonomicName lsidName="O. mawphlangensis" pageId="0" pageNumber="47" rank="species" species="mawphlangensis">
<emphasis italics="true" pageId="0" pageNumber="47">O. mawphlangensis</emphasis>
</taxonomicName>
, 44.0-55.0 mm in males and 86.0-97.0 mm in females in
<taxonomicName lsidName="O. rotodora" pageId="0" pageNumber="47" rank="species" species="rotodora">
<emphasis italics="true" pageId="0" pageNumber="47">O. rotodora</emphasis>
</taxonomicName>
, 66.6 mm in male in
<taxonomicName lsidName="O. sinica" pageId="0" pageNumber="47" rank="species" species="sinica">
<emphasis italics="true" pageId="0" pageNumber="47">O. sinica</emphasis>
</taxonomicName>
, and 74.4-124.4 mm in males and 94.6-137.4 mm in females in
<taxonomicName lsidName="O. splendida" pageId="0" pageNumber="47" rank="species" species="splendida">
<emphasis italics="true" pageId="0" pageNumber="47">O. splendida</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="description">
<paragraph pageId="0" pageNumber="47">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="47">Adult male. Body slender and small, SVL 36.8 mm. Head length larger than head width, HDW/HDL ratio 0.88; snout short, rounded in dorsal view, projecting beyond lower jaw, snout length larger than eye diameter, SNT/ED ratio 1.35; canthus rostralis distinct; nostril rounded, located laterally, closer to tip of snout than eye; internasal distance larger than interorbital distance, IND/IOD ratio 1.09; loreal region slightly concave and oblique; eye large and prominent; tympanum rounded, large, TD/ED ratio 0.86, edge of tympanum slightly elevated relative to tympanum; strong vomerine ridges bearing vomerine teeth; tongue deeply notched distally; pupil horizontally oval; pineal body present, small; vocal sac absent.</paragraph>
<paragraph pageId="0" pageNumber="47">Forelimbs slender, HND/SVL ratio 0.28, RAD/SVL ratio 0.22; fingers slender, relative finger lengths II &lt;I &lt;IV &lt;III; tips of fingers expanded into disc, all with circummarginal grooves, horizontal grooves present, without webbing and lateral fringes; subarticular tubercles prominent: 1, 1, 2, 2; inner metacarpal tubercle oval, elongate; medium and outer metacarpal tubercles oval; nuptial pads present on base of finger I, medially along inner side of fingers II and III.</paragraph>
<paragraph pageId="0" pageNumber="47">Hindlimbs slender, FTL/SVL ratio 0.70, TIB/SVL ratio 0.50; heels overlapping when thighs are appressed at right angles with respect to body; tibiotarsal articulation reaches to nostril when leg stretched forward; relative toe lengths I &lt;II &lt;III &lt;V &lt;IV; toes entirely webbed; tips of toes expanded into disc with circummarginal grooves; subarticular tubercles prominent: 1, 1, 2, 3, 2; inner metatarsal tubercle oval, elongate, almost equal length to first toe; outer metatarsal tubercle absent.</paragraph>
<paragraph pageId="0" pageNumber="47">Dorsal skin relatively smooth, granular; skin of loreal region smooth; weak supratympanic fold from posterior corner of eye to posterior edge of tympanum; dorsolateral folds absent; tiny conical spines present on temporal region except tympanum, skin of commissure of jaw, upper edge of eyelid, and along dorsolateral sides of body. Ventral skin smooth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="coloration of holotype in life">
<paragraph pageId="0" pageNumber="47">Coloration of holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="47">Skin of dorsal body, dorsal limbs and flanks with irregular moss-green speckles and brown mottling; dorsal skin of limbs with distinct brown transverse bands; ventral skin of body greyish white with light brown mottling; ventral skin of forelimb greyish white, ventral skin of hindlimb purplish brown. Iris black, with irregular gold-green reticulated mottles; pineal body light green; tympanum dark brown; nuptial pad creamy white.</paragraph>
<caption doi="10.3897/zookeys.1120.87067.figure3" httpUri="https://binary.pensoft.net/fig/738804" pageId="0" pageNumber="47" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 3.</emphasis>
Morphological features of the male holotype GEP a055 in life:
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
dorsolateral view
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
ventral view of hand, showing nuptial pads on fingers I, II and III
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
ventral view of foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="coloration of holotype in preservative">
<paragraph pageId="0" pageNumber="47">Coloration of holotype in preservative.</paragraph>
<paragraph pageId="0" pageNumber="47">Skin of dorsal body, dorsal limbs and flanks greyish brown, with brown mottling and dark brown transverse bands, moss-green speckles absent; ventral skin of body greyish white with brown mottling; ventral skin of thighs greyish white, ventral skin of shank and foot dark grey with dark brown mottling.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="variation">
<paragraph pageId="0" pageNumber="47">Variations.</paragraph>
<paragraph pageId="0" pageNumber="47">
Mensural data of the type series are listed in Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements (minimum-maximum (mean + / - SD); in mm) of Odorrana concelata sp. nov." httpUri="http://table.plazi.org/id/0ADD15FD3B6A9241E03560587B9B69FE" pageId="0" pageNumber="47" tableUuid="0ADD15FD3B6A9241E03560587B9B69FE">3</tableCitation>
. Most of the paratypes are similar to the holotype in morphology and color pattern, except for the following: (1) skin of dorsal trunk lacking tiny spines (vs. present in the male paratype GEP a052); (2) sparse spines on temporal region except tympanum, skin of commissure of jaw, upper edge of eyelid, and along dorsolateral sides of body; nuptial pads absent; and larger body size in female paratypes (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Morphological features of the female paratype GEP a 050 in life: A dorsolateral view B ventral view C ventral view of hand D ventral view of foot." figureDoi="10.3897/zookeys.1120.87067.figure4" httpUri="https://binary.pensoft.net/fig/738805" pageId="0" pageNumber="47">4</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="0ADD15FD3B6A9241E03560587B9B69FE" httpUri="http://table.plazi.org/id/0ADD15FD3B6A9241E03560587B9B69FE" pageId="0" pageNumber="47" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Table 3.</emphasis>
Measurements (minimum-maximum (mean
<normalizedToken originalValue="±">+/-</normalizedToken>
SD); in mm) of
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<table pageId="0" pageNumber="47">
<tr pageId="0" pageNumber="47">
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">Voucher</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a052</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a053</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a054</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a055</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">Range</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">Voucher</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a050</th>
<th colspan="1" pageId="0" pageNumber="47" rowspan="1">GEP a051</th>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">Sex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Male</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
Males (
<emphasis italics="true" pageId="0" pageNumber="47">n</emphasis>
= 4)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">Sex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Female</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">Female</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">34.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">35.7</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">35.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">36.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
34.0-36.8 (35.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">SVL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">46.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">41.4</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
11.8-12.8 (12.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HDL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">15.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">13.1</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
10.9-11.3 (11.1
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HDW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">14.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.2</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">SNT</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">5.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">5.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
5.1-5.3 (5.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">SNT</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">5.6</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">IND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
3.2-3.6 (3.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">IND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.5</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">IOD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
3.0-3.1 (3.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">IOD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.3</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">ED</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
3.8-4.1 (4.0
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">ED</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.5</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">4.2</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">TD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
3.2-3.4 (3.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">TD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">3.2</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">10.2</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
9.6-10.2 (9.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">HND</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.9</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12.3</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">RAD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">7.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">7.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">8.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
7.1-8.0 (7.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.4)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">RAD</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9.1</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">FTL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">24.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">24.8</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">24.7</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">25.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
24.0-25.6 (24.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">FTL</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">32.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">29.6</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">TIB</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">17.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">17.6</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">18.0</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">18.5</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
17.3-18.5 (7.8
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6)
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">TIB</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">23.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">20.8</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zookeys.1120.87067.figure4" httpUri="https://binary.pensoft.net/fig/738805" pageId="0" pageNumber="47" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 4.</emphasis>
Morphological features of the female paratype GEP a050 in life:
<emphasis bold="true" pageId="0" pageNumber="47">A</emphasis>
dorsolateral view
<emphasis bold="true" pageId="0" pageNumber="47">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
ventral view of hand
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
ventral view of foot.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="distribution">
<paragraph pageId="0" pageNumber="47">Distribution and habits.</paragraph>
<paragraph pageId="0" pageNumber="47">
Currently,
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. is known only from its type locality (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Odorrana concelata sp. nov. (solid circle), O. lipuensis (solid squares), and O. liboensis (solid triangle)." figureDoi="10.3897/zookeys.1120.87067.figure1" httpUri="https://binary.pensoft.net/fig/738802" pageId="0" pageNumber="47">1</figureCitation>
, solid circle). The nocturnal karst-dweller inhabits mossy rocks and damp forest floors in subtropical evergreen broad-leaved forests and secondary forests at elevations between 200-300 m (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microhabitat of Odorrana concelata sp. nov. (A, B) and the uncaptured individuals of juvenile (C), female (D), and male (E) in situ." figureDoi="10.3897/zookeys.1120.87067.figure5" httpUri="https://binary.pensoft.net/fig/738806" pageId="0" pageNumber="47">5A, B</figureCitation>
). They are completely hidden in their habitat by their coloration (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microhabitat of Odorrana concelata sp. nov. (A, B) and the uncaptured individuals of juvenile (C), female (D), and male (E) in situ." figureDoi="10.3897/zookeys.1120.87067.figure5" httpUri="https://binary.pensoft.net/fig/738806" pageId="0" pageNumber="47">5D, E</figureCitation>
). During breeding season (March to June), they congregate in and around the small and steep moss-covered waterfalls which flows out of karst caves (ca. 1-2 m width). Juveniles were observed in June (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Microhabitat of Odorrana concelata sp. nov. (A, B) and the uncaptured individuals of juvenile (C), female (D), and male (E) in situ." figureDoi="10.3897/zookeys.1120.87067.figure5" httpUri="https://binary.pensoft.net/fig/738806" pageId="0" pageNumber="47">5C</figureCitation>
). No individuals were found during surveys in mid-July.
</paragraph>
<caption doi="10.3897/zookeys.1120.87067.figure5" httpUri="https://binary.pensoft.net/fig/738806" pageId="0" pageNumber="47" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 5.</emphasis>
Microhabitat of
<taxonomicName authorityName="Wang, Zeng, &amp; Lin" authorityYear="2022" class="Amphibia" family="Ranidae" genus="Odorrana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odorrana concelata" order="Anura" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="concelata">
<emphasis italics="true" pageId="0" pageNumber="47">Odorrana concelata</emphasis>
</taxonomicName>
sp. nov. (
<emphasis bold="true" pageId="0" pageNumber="47">A, B</emphasis>
) and the uncaptured individuals of juvenile (
<emphasis bold="true" pageId="0" pageNumber="47">C</emphasis>
), female (
<emphasis bold="true" pageId="0" pageNumber="47">D</emphasis>
), and male (
<emphasis bold="true" pageId="0" pageNumber="47">E</emphasis>
) in situ.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>