628 lines
97 KiB
XML
628 lines
97 KiB
XML
<document ID-CLB-Dataset="9126" ID-DOI="http://dx.doi.org/10.3897/zookeys.1008.58854" ID-GBIF-Dataset="b31dfe66-0bbe-4226-8aa7-941526c56be1" ID-PMC="PMC7790807" ID-Pensoft-Pub="1313-2970-1008-1" ID-Pensoft-UUID="285C40BD37805C919C7CFFB981C9CF79" ID-PubMed="33505185" ID-ZooBank="6BF494BEA31B42D788AF2AB04EEB4725" ModsDocID="1313-2970-1008-1" checkinTime="1609463513021" checkinUser="pensoft" docAuthor="Huang, Chao, Ahyong, Shane T. & Shih, Hsi-Te" docDate="2020" docId="DB07660729E05031B65744A6C14D4B3D" docLanguage="en" docName="ZooKeys 1008: 1-15" docOrigin="ZooKeys 1008" docPubDate="2020-12-31" docSource="http://dx.doi.org/10.3897/zookeys.1008.58854" docTitle="Phasmon typhlops Huang & Ahyong & Shih 2020, sp. nov." docType="treatment" docUuid="FEE83A41-2669-4B3F-B3A0-5E1F15F1DEB1" docUuidSource="ZooBank" docVersion="6" id="285C40BD37805C919C7CFFB981C9CF79" lastPageNumber="1" masterDocId="285C40BD37805C919C7CFFB981C9CF79" masterDocTitle="The second known stygomorphic freshwater crab from China, Phasmon typhlops gen. nov. et sp. nov. (Crustacea, Decapoda, Potamidae), diverged at the beginning of the Late Miocene" masterLastPageNumber="15" masterPageNumber="1" pageNumber="1" updateTime="1732840732287" updateUser="ExternalLinkService">
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<mods:titleInfo id="E31B0D81EE09FF31949D3EBD41846F78">
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<mods:title id="0F154B8CDF2FFACC1ADA5A6588A8580F">The second known stygomorphic freshwater crab from China, Phasmon typhlops gen. nov. et sp. nov. (Crustacea, Decapoda, Potamidae), diverged at the beginning of the Late Miocene</mods:title>
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<mods:name id="5C102C039777C7FCD35A457CAA3FCE7E" type="personal">
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<mods:roleTerm id="C8BD3C3934B6D3A134D7A9B580E325FE">Author</mods:roleTerm>
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<mods:namePart id="12FA9FC784068D1427F4FB8FA2C9D11B">Huang, Chao</mods:namePart>
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<mods:affiliation id="FBD00813964ED0A0E39CD9802E9B5C7C">Unaffiliated, Guangdong, China</mods:affiliation>
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</mods:name>
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<mods:name id="6D34BFA5ECD8EDBF066A043D95153090" type="personal">
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<mods:roleTerm id="171F8A32CD1533F8366D2618C749FCDE">Author</mods:roleTerm>
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<mods:namePart id="836E429EAE344B0A619D99E62A056B8B">Ahyong, Shane T.</mods:namePart>
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<mods:affiliation id="69FD8A7BBC417EF98E9D38F3B8FF990B">Australian Museum, 1 William St, Sydney NSW 2010, Australia & School of Biological, Earth and Environmental Sciences, University of New South Wales, Kensington, NSW 2052, Australia</mods:affiliation>
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<mods:roleTerm id="9BD29DE33A97EAC03DAB9171F5125C74">Author</mods:roleTerm>
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<mods:namePart id="1C7498398AACEA835C1D3A25C3D77F4B">Shih, Hsi-Te</mods:namePart>
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<mods:nameIdentifier id="97056CC3B6A91F0430B27DE48FCDF0EF" type="ORCID">https://orcid.org/0000-0003-1317-8783</mods:nameIdentifier>
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<mods:affiliation id="9D9F753F9FD49060BD75FB6CEAC61B34">Department of Life Science and Research Center for Global Change Biology, National Chung Hsing University, Taichung 402, Taiwan</mods:affiliation>
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<mods:nameIdentifier id="E4C926FA83A5FAFD6CDB77B24892283E" type="email">htshih@dragon.nchu.edu.tw</mods:nameIdentifier>
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<mods:title id="AB82505C070CA54854A3AA3A106E366B">ZooKeys</mods:title>
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<mods:part id="A758F6FBBEE92DF80C6917DE11279426">
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<mods:date id="8D54D1CEC787F5F26DE375D480698E55">2020</mods:date>
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<mods:number id="C1F7196D49BB8D0D676302BD578AD830">2020-12-31</mods:number>
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<mods:number id="FCCD38E826EAF5ACF4CF1617BF71880E">1008</mods:number>
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<mods:url id="0A42072B91B4C331A2B32EB131690ED8">http://dx.doi.org/10.3897/zookeys.1008.58854</mods:url>
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<mods:classification id="B1B090FBEE82FCD01A6C09476050122F">journal article</mods:classification>
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<mods:identifier id="2DBBA0E9F64E9A56BDAB45EDBB30C20F" type="DOI">http://dx.doi.org/10.3897/zookeys.1008.58854</mods:identifier>
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<mods:identifier id="680019FF2ADF508CCA0CCD825007C2FE" type="Pensoft-Pub">1313-2970-1008-1</mods:identifier>
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<subSection id="D8701E56DB82C8A4FB2AD45AFC8473E2" pageId="0" pageNumber="1" type="taxonomy">
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<treatment id="DB07660729E05031B65744A6C14D4B3D" ID-GBIF-Taxon="176352596" LSID="urn:lsid:zoobank.org:act:FEE83A41-2669-4B3F-B3A0-5E1F15F1DEB1" httpUri="http://treatment.plazi.org/id/DB07660729E05031B65744A6C14D4B3D" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Potamidae" scope_order="Decapoda" scope_subFamily="Potamiscinae">
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<subSubSection id="03E440131D7939C15EAC152BCD33F9C0" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="123452832E8461388DF857B62C604509" pageId="0" pageNumber="1">
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<taxonomicName id="55667902E085EF400FCEE381C04E0097" ID-CoL="84H42" LSID="http://zoobank.org/FEE83A41-2669-4B3F-B3A0-5E1F15F1DEB1" authority="Huang & Ahyong & Shih, 2020" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops" status="sp. nov.">Phasmon typhlops</taxonomicName>
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<taxonomicNameLabel id="B3261CEE47EA8C217AD5BFE6CE9C1D6A" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="51C2CF9636CBACB071DAD19B4736ED5F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">Figs 1</figureCitation>
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<figureCitation id="B5694808E8140E89DD9B765BC71844E4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">, 2</figureCitation>
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<figureCitation id="D296E37674A8F202ED27D5A9D911BD84" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">, 3</figureCitation>
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<figureCitation id="47F32EA5DA74A0D80AC62F8574926A82" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phasmon typhlops gen. nov. et sp. nov., gonopods: left G 1, abdominal view (A) left G 1, sternal view (B) left G 2, mesial view (C) left G 2, pleonal view (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C6596BF85CC4104DB5EB267C7C59D0F5" pageId="0" pageNumber="1" type="type material">
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<paragraph id="067041CB1C9A839DCD852421005AF6B7" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="DC784B27E8FA2DEB125BCAC91ED400E3" pageId="0" pageNumber="1">
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<emphasis id="BB1CA056A143C03D2D72367A1EE45C0D" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: SYSBM 001982, male (14.4
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<normalizedToken id="F6858D3B480A109D2C807FBD707AA707" originalValue="×">x</normalizedToken>
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9.0 mm), Leiping Town, Daxin County, Chongzuo City, Guangxi Province, China,
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<geoCoordinate id="CD22CEAAD15E082ADABEF25C96731547" degrees="22.65" direction="north" orientation="latitude" precision="555" value="22.65">22.65°N</geoCoordinate>
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, 107.10° E, subterranean karst stream in cave, baited trap, coll. local collector, December 2019.
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<emphasis id="4D44A4814DD4F449154B206C667ED9B6" bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
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: AM P.105524, female (22.1
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<normalizedToken id="BBF090973E77F165C53539E9DB23E5E0" originalValue="×">x</normalizedToken>
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13.7 mm), Leiping Town, Daxin County, Chongzuo City, Guangxi Province, China, karst spring, baited trap, coll. local collector, September 2018.
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</paragraph>
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<caption id="669B4940C09330E80EF8F0366BDF0720" doi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
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<paragraph id="860C92C75316567E018909FA04825092" pageId="0" pageNumber="1">
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<emphasis id="2251E3A41E28F3165F8D5BF3ADE8FEDA" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
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<taxonomicName id="D1889E4C7D55CFA338BCDF5B1F4A1057" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
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<emphasis id="2147CC551FC9C70450BEEE9114D1786F" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
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</taxonomicName>
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gen. nov. et sp. nov., male holotype (14.4
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<normalizedToken id="62D73762889814E2735FD2093BCC7698" originalValue="×">x</normalizedToken>
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||
9.0 mm), SYSBM 001982 (
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<emphasis id="B748AFEF328681C2686CF7EF5CE0E1CD" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
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) female paratype (22.1
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<normalizedToken id="2C4F9A91EEF88F6884F31416D0DFBEB8" originalValue="×">x</normalizedToken>
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13.7 mm), AM P.105524 (
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<emphasis id="73ABA8A97B5A610F4528791D08592E9B" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
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). Cephalothorax, anterior view (
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<emphasis id="8ACAA618E283F21D68F41ACF80A3D966" bold="true" pageId="0" pageNumber="1">A</emphasis>
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) anterior thoracic sternum (
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<emphasis id="8BF85B2C417CEB53CF04602C2E88A842" bold="true" pageId="0" pageNumber="1">B</emphasis>
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) anterior thoracic sternum and pleon, ventral view (
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<emphasis id="B0C380737D57225BCE5654AC883BF462" bold="true" pageId="0" pageNumber="1">C</emphasis>
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) sterno-pleonal cavity with G1
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<emphasis id="E6920B547EB56A498613899C36A4742A" italics="true" pageId="0" pageNumber="1">in situ</emphasis>
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, ventral view (
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<emphasis id="00CFF250FAED98121BC1174B017F9A80" bold="true" pageId="0" pageNumber="1">D</emphasis>
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) pleon, ventral view (
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<emphasis id="192409780B46659457E575BA2B5B4D48" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) vulvae, ventral view (
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<emphasis id="83A3FD7A896FF8A65210EFDA44C05615" bold="true" pageId="0" pageNumber="1">F</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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||
<subSubSection id="E4E816F82A8E48EB498D0B9463EE797B" pageId="0" pageNumber="1" type="description">
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<paragraph id="E8853B0E5C857C5E7C1680341D0B928A" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="CE589130FB6197DAC8513BF98CAD6959" pageId="0" pageNumber="1">
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Carapace broad, about 1.6 times as wide as long; fronto-orbital width about twice width of posterior margin; regions indistinct, dorsal surface slightly convex; surface finely pitted (Fig.
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<figureCitation id="858451CD0EFA8E0C2DF588D345133CB2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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). Frontal margin weakly sinuous, continuous with supraorbital margin, forming almost straight transverse margin in dorsal view (Figs
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<figureCitation id="54348462E341CC94C9DF97A4F4882047" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A, B</figureCitation>
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). Epigastric cristae and postorbital cristae almost indiscernible (Figs
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<figureCitation id="1B77DEDFF7C7E449C073987929B261F3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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,
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<figureCitation id="1EBA8B957574CED8A50A0DA0B0CE21F7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
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). Branchial regions slightly swollen (Figs
|
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<figureCitation id="D6C3560A137CA1435157368CA3202B42" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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,
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<figureCitation id="C030A26697062FE9CC0046DF6810E222" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
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||
). Cervical groove shallow (Fig.
|
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<figureCitation id="45DC9E5E63174623D1C99D2495C1D0F4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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). Mesogastric region slightly convex (Fig.
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<figureCitation id="DFAA4DCBE0AF98A0D11653FFD51FF3AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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). External orbital angle obsolete, outer margin convex, almost indistinguishable from anterolateral margin (Figs
|
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<figureCitation id="3E957052E1348706ABB257D6AB9D6465" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
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,
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<figureCitation id="E716C0CE7F92168A5F04A25053397417" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
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). Epibranchial tooth granular, inconspicuous (Fig.
|
||
<figureCitation id="EB2ABA5B6EA84C83C72A5C2497768C6C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
). Anterolateral margin lined with 15-20 small, single or partially fused granules. Posterolateral margin posteriorly convergent (Fig.
|
||
<figureCitation id="8D91E23B15F135200DB46D020453702F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
); posterolateral surface generally smooth (Fig.
|
||
<figureCitation id="781690F1577E6FD2F8DE70B0928A91AD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
). Orbits shallow; supraorbital margins weakly cristate, infraorbital margins lined with granules (Fig.
|
||
<figureCitation id="D6F558415CB5100F3C8F1F8B62893F3F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
). Eyes almost immobile, greatly reduced, tapering, length about half orbital width; peduncle short, stout; cornea vestigial, surface without facets, unpigmented (Figs
|
||
<figureCitation id="1BA86F085159FE7A2281F81705F557B6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
,
|
||
<figureCitation id="61B7FF0486E6CB985B4C5E434DB31166" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3F</figureCitation>
|
||
). Sub-orbital, pterygostomial and sub-hepatic regions generally smooth, pitted (Fig.
|
||
<figureCitation id="705CCC061437129D8E40748B90B61FF7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
). Antennules large, folded within broad fossae; antennae very short (Fig.
|
||
<figureCitation id="1C016F2DA7FF1FFF4A48AA30E61C2E5F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
). Median lobe of epistome posterior margin broadly triangular, lateral margins sinuous (Fig.
|
||
<figureCitation id="5AFD4C1FEE9A8CD14E3B5915DB563389" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="F5861032F61A3C92A8FEFD98EA3DD0C8" doi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||
<paragraph id="B91279609E2BE515BB5FD50CC55EA195" pageId="0" pageNumber="1">
|
||
<emphasis id="7FE6BBB24163CACEBC4ABA7866655081" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||
<taxonomicName id="4C19B60ED870ABF0D37EF367CD8DB871" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="37C7EA10B54D4443DC01CA7ADF9EF9CB" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov., male holotype (14.4
|
||
<normalizedToken id="3B3DAA7CCA3E8F8CD5E17DC555221207" originalValue="×">x</normalizedToken>
|
||
9.0 mm), SYSBM 001982. Left maxilliped 3 (
|
||
<emphasis id="142B7C2797A924B7046545AFFCC5EE72" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
) left G2, pleonal view (
|
||
<emphasis id="B79896C2169B703EF1386A2828D3CD3A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
) left G1, pleonal view (
|
||
<emphasis id="75B2C5C666046B08460B107D7E6E2261" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
) major cheliped (
|
||
<emphasis id="713232FF38F578465B866E11542BC1DC" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
) minor cheliped (
|
||
<emphasis id="9F65BFBB9C189D78E797E9F3BBB8C558" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) colour in life (
|
||
<emphasis id="D67C6486A8E5F155EF0E5FF0555F5DD4" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
). Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C7D25AE3695C0C6D9727F80A7272EF96" pageId="0" pageNumber="1">
|
||
Maxilliped 3 merus subtrapezoidal, with median depression, width about 1.2
|
||
<normalizedToken id="2E2D611643C3456595783107AC832485" originalValue="×">x</normalizedToken>
|
||
length; ischium subtrapezoidal with shallow median sulcus, distomesial margin rounded, width about 0.6
|
||
<normalizedToken id="F5E5D357B8B908EC810DBAA3EA4972BA" originalValue="×">x</normalizedToken>
|
||
length. Exopod reaching proximal one-third of merus; flagellum longer than half ischium length (Fig.
|
||
<figureCitation id="6D2DBFB806A3C3367D3E41ACA51941E4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="805BC8548721EE700F1621296A599D19" pageId="0" pageNumber="1">
|
||
Chelipeds (pereiopod 1) subequal (Figs
|
||
<figureCitation id="6CAC372D9302EC3D7A62A540B141AAE3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
,
|
||
<figureCitation id="CA3578D7F045D13C2E9909C2A089CDDD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||
). Merus trigonal in cross section; margins slightly crenulated, surface generally smooth (Figs
|
||
<figureCitation id="DACCB4C6A9ACF684981E039322236EC1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
,
|
||
<figureCitation id="B13028D51E41AE64B05FF19E6A866EEA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
). Carpus with sharp spine at inner-distal angle (Fig.
|
||
<figureCitation id="A2D859B589426F313A6FB57EE7AA88E4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1 A</figureCitation>
|
||
). Major cheliped palm length about 1.5
|
||
<normalizedToken id="C0BE4D7A93364C279B02A9339843FEA1" originalValue="×">x</normalizedToken>
|
||
height; dactylus 0.9
|
||
<normalizedToken id="0324B0FA36CF032659B3700CD4A670E1" originalValue="×">x</normalizedToken>
|
||
palm length (male) (Fig.
|
||
<figureCitation id="6543FDB3EC9FB08A0833C5E5C1FCE93C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||
), as long as palm (female). Palm surface pitted (Fig.
|
||
<figureCitation id="BFE10A75AA680CE0703484115FD1B2D6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||
). Dactylus as long as pollex (Fig.
|
||
<figureCitation id="0BD8F9A883D3F62B30A5FCD52A137377" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||
). Occlusal margin of fingers with 18-20 irregular blunt teeth, without gape when closed (Fig.
|
||
<figureCitation id="D34A2BB726AE7214583404FF272AC647" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C1DE4B04EC77DEFF71B3102DDF6F8379" pageId="0" pageNumber="1">
|
||
Ambulatory legs (pereiopods 2-5) slender with very sparse short setae (Fig.
|
||
<figureCitation id="C0A3B543F05FB8C11BAF38496491E566" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
). Pereiopod 3 merus 0.9
|
||
<normalizedToken id="9DB87E7BF098B10485F438E112A24717" originalValue="×">x</normalizedToken>
|
||
CL (male) (Fig.
|
||
<figureCitation id="87A43BE82BF0B0EA697EAB4F43D2FCB9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
), 0.8
|
||
<normalizedToken id="E200510534C136688509162107F0395C" originalValue="×">x</normalizedToken>
|
||
CL (female) (Fig.
|
||
<figureCitation id="27BAFFCEDC3233ECE76B1C13A0FBA689" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1B</figureCitation>
|
||
). Pereiopod 5 propodus length 2.8
|
||
<normalizedToken id="BEA1133DD619D342965E7F6797DB6A53" originalValue="×">x</normalizedToken>
|
||
height (male) (Fig.
|
||
<figureCitation id="0F6EC77C0B6BA18ADFF6E216B662F878" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
), 3.4 height (female) (Fig.
|
||
<figureCitation id="D5FEF2C3BDE5145C8C502BBA7C75C02E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1B</figureCitation>
|
||
), shorter than dactylus; dactylus length 6.1
|
||
<normalizedToken id="AC81E2864E298AB8A15720765A85466D" originalValue="×">x</normalizedToken>
|
||
height (male) (Fig.
|
||
<figureCitation id="A7D8198BEA51914D2E980559499444D1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1A</figureCitation>
|
||
), 6.2
|
||
<normalizedToken id="0E6067359EC33D898C05CA9DB56D3022" originalValue="×">x</normalizedToken>
|
||
height (female) (Fig.
|
||
<figureCitation id="881D770218B6DD5FB079FF0ED13BF24D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BAF87D9E3D0905234526EA2D07B8F78F" pageId="0" pageNumber="1">
|
||
Male thoracic sternum generally smooth, pitted; sternites 1-4 width about 2.3
|
||
<normalizedToken id="94BE643FD0305C51B79FC77C586E3850" originalValue="×">x</normalizedToken>
|
||
length; sternites 1, 2 forming indistinguishably fused, broad triangle; fused sternites 1, 2 demarcated from sternite 3 by shallow transverse sulcus; sternites 3, 4 fused without indication of demarcation except for shallow lateral notch (Fig.
|
||
<figureCitation id="94E8538C24C2D6A4B9DB7ECE15F929FF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). Male sterno-pleonal cavity reaching anteriorly slightly beyond level of cheliped coxa articular condyle (Fig.
|
||
<figureCitation id="9B40828799312BC19468B8D2B72BB1E3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2B</figureCitation>
|
||
); deep median longitudinal groove between sternites 7, 8 (Fig.
|
||
<figureCitation id="5A7BD9312A49E829DB60635010311C79" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2D</figureCitation>
|
||
). Male pleonal locking tubercle positioned at mid-length of sternite 5 (Fig.
|
||
<figureCitation id="972E57637030FB7C67F098574F3171E6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2D</figureCitation>
|
||
). Female vulvae reaching sutures of sternites 5/6 anteriorly but not posteriorly to sutures of sternites 6/7, positioned widely apart from each other (Fig.
|
||
<figureCitation id="86AE021F73D28D3FED8DF26027C0D65A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8CBAA706D92FB5AF77E0CBF58E690EF2" pageId="0" pageNumber="1">
|
||
Male pleon broadly triangular; somites 3-6 progressively narrower; somite 6 width approximately 2.7
|
||
<normalizedToken id="E013640D914808712BF3E38258CB2B0D" originalValue="×">x</normalizedToken>
|
||
length; telson width 1.6
|
||
<normalizedToken id="0BBBB3620936D7992F5045A311FB2A90" originalValue="×">x</normalizedToken>
|
||
length; lateral margins slightly convex, apex rounded (Fig.
|
||
<figureCitation id="93BFEA7A583BCB9AFBB03153EC8FFC4D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2C</figureCitation>
|
||
). Female pleon subovate (Fig.
|
||
<figureCitation id="758476E90B775CC1BBB6171A63F2BDCF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="124CC6D41DA45B82AE89004B9F5CBA04" pageId="0" pageNumber="1">
|
||
G1 tapering, slightly sinuous, tip exceeding pleonal locking tubercle but not reaching suture between thoracic sternites 4/5
|
||
<emphasis id="A4CFCE856A31E9B5C71D7D281C2739B4" italics="true" pageId="0" pageNumber="1">in situ</emphasis>
|
||
(Fig.
|
||
<figureCitation id="9988BDA7B6456E0BFD7DD3B4CEB59C7F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2D</figureCitation>
|
||
); proximal segment length about 2.3
|
||
<normalizedToken id="04234E777CB0331E848491186A34EAFE" originalValue="×">x</normalizedToken>
|
||
length of distal segment (Figs
|
||
<figureCitation id="B03B2C1D428C4E5239BEB1EA82086F1D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3C</figureCitation>
|
||
,
|
||
<figureCitation id="76BA48C09822BE6BCF4D00EF590BD4A9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phasmon typhlops gen. nov. et sp. nov., gonopods: left G 1, abdominal view (A) left G 1, sternal view (B) left G 2, mesial view (C) left G 2, pleonal view (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1">4A, B</figureCitation>
|
||
). Distal segment slender, tapering anteriorly, slightly inclined towards midline; tip pointed upwards in dissected view (Figs
|
||
<figureCitation id="3C420C069D4490E10B1BCFBF6643C572" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3C</figureCitation>
|
||
,
|
||
<figureCitation id="74FD039830A8DB5D734CD33864F19F52" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phasmon typhlops gen. nov. et sp. nov., gonopods: left G 1, abdominal view (A) left G 1, sternal view (B) left G 2, mesial view (C) left G 2, pleonal view (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1">4A, B</figureCitation>
|
||
). G2 slender, almost straight, proximal portion with distal two-thirds subcylindrical, length about 2.4
|
||
<normalizedToken id="8AFEFFC002D379A94CF7DD2F170D6344" originalValue="×">x</normalizedToken>
|
||
length of distal portion (Figs
|
||
<figureCitation id="257C075FC4948E79BB6628713CE36484" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3B</figureCitation>
|
||
,
|
||
<figureCitation id="56420BA6B52CD59C1C23DDBB207BCF6D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phasmon typhlops gen. nov. et sp. nov., gonopods: left G 1, abdominal view (A) left G 1, sternal view (B) left G 2, mesial view (C) left G 2, pleonal view (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1">4C, D</figureCitation>
|
||
); distal portion flattened, apex acute, proximally with small triangular lobe.
|
||
</paragraph>
|
||
<caption id="D69C57A8514FC071C5093FDD58C1B7B3" doi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
||
<paragraph id="BFA34F375F8030E9B26745CBC2A312DE" pageId="0" pageNumber="1">
|
||
<emphasis id="236289334391252855586ED9DAE089EC" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||
<taxonomicName id="C3E0529C5DE95F679CD246E317385CF0" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="52CCAAD9F441B1522A93D0BD3A8CF08D" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov., gonopods: left G1, abdominal view (
|
||
<emphasis id="3B3AD12B4D1C2562C9D8B125F812E3F3" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
) left G1, sternal view (
|
||
<emphasis id="CBD6E6B9C2513A12681FB3EC42FF3ABC" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
) left G2, mesial view (
|
||
<emphasis id="0102E1936F391235079F55CC259EB1C3" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
) left G2, pleonal view (
|
||
<emphasis id="40C99C442A06CD7A7753BF345C46D79A" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
). Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="49AE6DD6F358A131944A4CD4D198D8D6" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="3EE81A3C093A83FAFDBB27E630061653" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="FFE8685CA0DB1AF0E7CAAA7796EEB483" pageId="0" pageNumber="1">
|
||
The species name is derived from the Greek words
|
||
<normalizedToken id="94A6B2F727F99EF09900C67304A27047" originalValue="“typhlos”">"typhlos"</normalizedToken>
|
||
and
|
||
<normalizedToken id="39EFEA6B0741026904F83BBE10017916" originalValue="“ops”">"ops"</normalizedToken>
|
||
, meaning
|
||
<normalizedToken id="D1FC57A1158B9B4B5DB2B0F7DBF09BEB" originalValue="“blind”">"blind"</normalizedToken>
|
||
and
|
||
<normalizedToken id="E7FAD3D09329653C2B4A1087A5394D1C" originalValue="“eyes”">"eyes"</normalizedToken>
|
||
, respectively. It refers to the greatly reduced and non-functional eyes of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A6F1A227A4391BC993E070994088D4F3" pageId="0" pageNumber="1" type="colour in life">
|
||
<paragraph id="F2E6A66693128CAF21DB4EE18C1F7450" pageId="0" pageNumber="1">Colour in life.</paragraph>
|
||
<paragraph id="108BBF2770B7E7BFE16A65B30E661895" pageId="0" pageNumber="1">
|
||
Pale yellowish-white all over (Fig.
|
||
<figureCitation id="AFD822F57AC12D73076B301CFFE0942B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9230E23E3EA05E6CAF21BC549F9F2429" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="C68905A28688AA525B7E332AE6E318F9" pageId="0" pageNumber="1">Habitat.</paragraph>
|
||
<paragraph id="98DBCCFC8052DECAC0FA72E438500A0F" pageId="0" pageNumber="1">
|
||
<taxonomicName id="487B4FAA53403778811CC869F6141581" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="D25D526B732B46C37C97C95C8BC54E7B" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. occurs in subterranean karst streams, but little is currently known about its precise habitat. According to the collector, subterranean streams in the dark zone of caves appear to be the primary habitat of
|
||
<taxonomicName id="CD2CCCA638480008354F621735656D94" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="0C48ADC569CE12C0C171274B9DAC8179" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., where it has been found in shallow and still water as well as flowing streamways. However, some specimens have also been captured at night from a karstic spring that is immediately connected to the more extensive subterranean karst system. We only examined the two type specimens, of which the holotype was collected from the former habitat and the paratype from the latter. An epigean species,
|
||
<taxonomicName id="640C2536B6703BEFCCDDB923FE871AA9" class="Crustacea" family="Potamidae" genus="Lacunipotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lacunipotamon cymatile" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cymatile">
|
||
<emphasis id="A07E99707BF8CAA30A22CD7784A1778A" italics="true" pageId="0" pageNumber="1">Lacunipotamon cymatile</emphasis>
|
||
</taxonomicName>
|
||
, inhabits the areas immediately adjacent to the spring and has been observed to prey on
|
||
<taxonomicName id="056DA6D1585A727A81C6882345689F0F" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="FBCF5BA691B2D8717BC7E2F478F3488C" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. (
|
||
<bibRefCitation id="91C6CDDB23E0BF7CC465506962748FF0" DOI="https://doi.org/10.1163/15685403-bja10067" author="Huang, C" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" pagination="1361 - 1379" refId="B21" refString="Huang, C, Shih, HT, Ahyong, ST, 2020c. The freshwater crab genus Lacunipotamon Dai, Song, He, Cao, Xu & Zhong, 1975 (Decapoda: Brachyura: Potamidae), with descriptions of two new species from southwestern China. Crustaceana 93: 1361 - 1379, DOI: https://doi.org/10.1163/15685403-bja10067" title="The freshwater crab genus Lacunipotamon Dai, Song, He, Cao, Xu & Zhong, 1975 (Decapoda: Brachyura: Potamidae), with descriptions of two new species from southwestern China." url="https://doi.org/10.1163/15685403-bja10067" volume="93" year="2020 c">Huang et al. 2020c</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4336D50512F7EC26AB1329C9DD73B075" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="BF20EE97CC35D802B637765488C61D12" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="51C7E993A8B7A10B2ED6F33B27497F98" pageId="0" pageNumber="1">Chongzuo City, Guangxi Province, China.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A7B0B26BB67C9D793D8AB14E7B837D5" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="1EFABB2AC6E4D558A20D1B276F108C47" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="617FE06CB6E808345BEE259973056CAC" pageId="0" pageNumber="1">
|
||
<taxonomicName id="C1649BA9A94CFAD575CB6CD7465DD91F" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="34BCCAB455D7789A1F9F62F1453D450E" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. can be considered a true stygobite owing to its stygomorphic features, in particular the strong reduction of the eyes, body depigmentation and slightly elongated appendages, which are consistent with its subterranean lifestyle (
|
||
<bibRefCitation id="7B4072D880AB0CA5DE87C83D32AAAE05" author="Holthuis, LB" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="0" pageNumber="1" refId="B14" refString="Holthuis, LB, 1986. Decapoda. In: Botosaneanu, L. (Ed.) Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial). Brill, Leiden, 589-615." title="Decapoda. In: Botosaneanu, L. (Ed.) Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial). Brill, Leiden, 589 - 615." year="1986">Holthuis 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="95289CFE77D6C67F8CA4F94374483096" author="Ng, PKL" journalOrPublisher="Stygologia" pageId="0" pageNumber="1" pagination="313 - 330" refId="B31" refString="Ng, PKL, Goh, R, 1987. Cavernicolous freshwater crabs (Crustacea Decapoda, Brachyura) from Sabah, Borneo. Stygologia 3: 313 - 330" title="Cavernicolous freshwater crabs (Crustacea Decapoda, Brachyura) from Sabah, Borneo." volume="3" year="1987">Ng and Goh 1987</bibRefCitation>
|
||
). Apart from
|
||
<taxonomicName id="542B03EAB21A1C135D31E7085C6A4D0D" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="8073B86D2EE5AED8CF859618E9992CA3" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov.,
|
||
<taxonomicName id="F4424524B4B898959024D5867742011D" authorityName="Huang, Shih & Ng" authorityYear="2017" class="Malacostraca" family="Potamidae" genus="Diyutamon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Diyutamon cereum" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cereum">
|
||
<emphasis id="EC3D020BA9142A11E08B5063ABDAEFD8" italics="true" pageId="0" pageNumber="1">Diyutamon cereum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="650788BD85767C30D213F05062D53DBA" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cerberusa caeca" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caeca">
|
||
<emphasis id="054F56EB9A8F331A45AEBB70E4571683" italics="true" pageId="0" pageNumber="1">Cerberusa caeca</emphasis>
|
||
</taxonomicName>
|
||
are the only other apparently blind stygomorphic potamid crabs known. We have not directly examined the eyes of
|
||
<taxonomicName id="B15271617BDFC582689B7B36F8DF759C" lsidName="C. caeca" pageId="0" pageNumber="1" rank="species" species="caeca">
|
||
<emphasis id="7999A932BE1F41D2F45A89A0220A34FF" italics="true" pageId="0" pageNumber="1">C. caeca</emphasis>
|
||
</taxonomicName>
|
||
, but those of
|
||
<taxonomicName id="B0BECFEFFF3CEB66E1BAEB44F06E1A2E" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="301F3DC78F87CB5F58628E7526F43A7F" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="44FB6B66E131FFEE943D034900A6B42F" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="B60E48FDBBE927EA40373BDABCA2B868" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
are unpigmented and the cornea is vestigial and without facets. Although we cannot exclude the possibility that the eyes of
|
||
<taxonomicName id="A81D05CAB165D85D4133FD658C7D5144" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="9CCB28396B24DD51CA178C70F1AFDEAA" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF11E118A1EBA20565F41915785C69B" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="4908B8A3E239CEECD240DAD64044AD1A" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
are capable of light detection, the absence of pigmentation or ommatidial facets indicates that the eyes are incapable of image formation. The enlarged antennules as present in
|
||
<taxonomicName id="CA4FEF0FD5E60CC951AB2926C3A7487A" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B2D34ACAFEADD23D622225F41D9ED9CF" italics="true" pageId="0" pageNumber="1">Phasmon</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. are otherwise seen in only a few cavernicolous freshwater crabs such as the gecarcinucids
|
||
<taxonomicName id="96ACB3510AEA28B1A08F4242728BEC77" authorityName="Husana, Naruse & Kase" authorityYear="2009" class="Reptilia" family="Gecarcinucidae" genus="Sundathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sundathelphusa waray" order="Squamata" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="waray">
|
||
<emphasis id="3B410E61744CAB25FC1C20C458B2E243" italics="true" pageId="0" pageNumber="1">Sundathelphusa waray</emphasis>
|
||
</taxonomicName>
|
||
Husana, Naruse & Kase, 2009, and
|
||
<taxonomicName id="9791F8986B9F5E56C2F178197CEE0EB3" lsidName="S. lobo" pageId="0" pageNumber="1" rank="species" species="lobo">
|
||
<emphasis id="A54530002FCCD97146728A5217B0851A" italics="true" pageId="0" pageNumber="1">S. lobo</emphasis>
|
||
</taxonomicName>
|
||
Husana, Naruse & Kase, 2009 (
|
||
<bibRefCitation id="75987C12C52A7589403BFAC7D258AAB7" DOI="https://doi.org/10.1651/08-3081.1" author="Husana, DEM" journalOrPublisher="Journal of Crustacean Biology" pageId="0" pageNumber="1" pagination="419 - 427" refId="B24" refString="Husana, DEM, Naruse, T, Kase, T, 2009. Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae). Journal of Crustacean Biology 29: 419 - 427, DOI: https://doi.org/10.1651/08-3081.1" title="Two new cavernicolous species of the genus Sundathelphusa from western Samar, Philippines (Decapoda: Brachyura: Parathelphusidae)." url="https://doi.org/10.1651/08-3081.1" volume="29" year="2009">Husana et al. 2009</bibRefCitation>
|
||
: figs 2B, 5B), and are likely a sensory compensation for the loss of vision (
|
||
<bibRefCitation id="D2161D66A6270D967F780EF39A441176" author="Culver, DC" journalOrPublisher="Harvard University Press, Cambridge" pageId="0" pageNumber="1" refId="B6" refString="Culver, DC, Kane, TC, Fong, DW, 1995. Adaptation and Natural Selection in Caves: the Evolution of Gammarusminus. Harvard University Press, Cambridge" title="Adaptation and Natural Selection in Caves: the Evolution of Gammarusminus." year="1995">Culver et al. 1995</bibRefCitation>
|
||
). Other than these two species, there are other stygomorphic gecarcinucids from Asia, but these can be separated from the new species by obvious family-level characters.
|
||
</paragraph>
|
||
<paragraph id="52579998BB42DB8D4A0CCD10F5FD7296" pageId="0" pageNumber="1">
|
||
Sexual dimorphism is evident in our two specimens of
|
||
<taxonomicName id="2F299E0E7229BF417E774EE6901AC858" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="E1A1E6FD37AA6223E03427AC249A6429" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
: the smaller male holotype has proportionally longer but stouter legs in comparison to the larger female. The anterior carapace of the larger female is also proportionately wider than the posterior than in the male. Although the differences in leg proportions follow the pattern of sexual dimorphism observed in other potamids (e.g.,
|
||
<bibRefCitation id="1A511C8C686AB33457E11D744F3850F4" DOI="https://doi.org/10.11646/zootaxa.4766.3.4" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="447 - 456" refId="B18" refString="Huang, C, Huang, SZ, Shen, ZX, 2020b. A new long-legged terrestrial freshwater crab, Calcipotamon puglabrum gen. nov. et sp. nov. (Crustacea: Decapoda: Potamidae), from Hainan Island, China. Zootaxa 4766: 447 - 456, DOI: https://doi.org/10.11646/zootaxa.4766.3.4" title="A new long-legged terrestrial freshwater crab, Calcipotamon puglabrum gen. nov. et sp. nov. (Crustacea: Decapoda: Potamidae), from Hainan Island, China." url="https://doi.org/10.11646/zootaxa.4766.3.4" volume="4766" year="2020 b">Huang et al. 2020b</bibRefCitation>
|
||
), whether this carapace difference is due to size, sex or general variation remains to be determined.
|
||
</paragraph>
|
||
<paragraph id="30C45341AF8A3F572630EDA434026070" pageId="0" pageNumber="1">
|
||
Taxonomically, the most striking features of
|
||
<taxonomicName id="45B0C0B619D979B75B9762B7C7EA0DF5" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4F9D89B4F6AF63BF57CE38253FB7D3EB" italics="true" pageId="0" pageNumber="1">Phasmon</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. are its very wide carapace (CW/CL=1.6; Fig.
|
||
<figureCitation id="72E1E93F4029C4EC7568832F1C061DBD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) and wide male anterior thoracic sternum (width 2.3
|
||
<normalizedToken id="A8B557C85B972E722D871BD4A2139A4E" originalValue="×">x</normalizedToken>
|
||
length; Fig.
|
||
<figureCitation id="85487ABFD223EED375A730D0FD52902B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2B</figureCitation>
|
||
). These characters combined immediately separate
|
||
<taxonomicName id="982D9D37ED7940F248C7DF97958A1C66" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="48775477C5DFEDE2064D4AD4E8023D72" italics="true" pageId="0" pageNumber="1">Phasmon</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. from all other potamid genera.
|
||
<taxonomicName id="88A87780B14BE403802081F24E17C74B" authorityName="Huang, Shih & Ng" authorityYear="2017" class="Malacostraca" family="Potamidae" genus="Diyutamon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Diyutamon cereum" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cereum">
|
||
<emphasis id="170DFEBF8E1486519D1FE8BB2CDFA3C1" italics="true" pageId="0" pageNumber="1">Diyutamon cereum</emphasis>
|
||
</taxonomicName>
|
||
occurs in Guizhou, which is relatively close to the type locality of
|
||
<taxonomicName id="A3F4B4674B8D33FB3C3383EE362B0264" lsidName="P. typhlops" pageId="0" pageNumber="1" rank="species" species="typhlops">
|
||
<emphasis id="926DDD74DDE7D3585C3B3379826DB366" italics="true" pageId="0" pageNumber="1">P. typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov.
|
||
<taxonomicName id="4FF680DB8C097A9A89D366DF3B6A6A25" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="B5EBFC0208F7ECBC03F4987FADAE843E" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. n. et sp. n. can be separated from
|
||
<taxonomicName id="74861E89F1BD7D98C1E6A0A46FEF081A" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="1B1CF29A5FF65DC9A16B597083797430" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
by its proportionally wider carapace (CW/CL=1.6 vs. 1.3-1.4 in
|
||
<taxonomicName id="D70D3909FBE01491155E596B7D85B28C" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="51B7BF270B4E11F278166564104D2741" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EBA914F1B286A5937CF5B93413D1E9EC" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 2A); granulate anterolateral carapace margins (Fig.
|
||
<figureCitation id="D6D8C0C59B8216FD7CACCB404F6AA01A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) (vs. spinose in
|
||
<taxonomicName id="D410B2FF18CE390B07A35916702027C3" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="9B3D31824026FFA4526F016E771D1B6C" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="C91CB806080C8E900F9E031AF67F2C50" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 2A); proportionally wider male anterior thoracic sternum (width 2.3
|
||
<normalizedToken id="14791E4094D685494934EA47A653758F" originalValue="×">x</normalizedToken>
|
||
length vs. width 1.7
|
||
<normalizedToken id="B9B9CEA892618872222FDDAB7C818529" originalValue="×">x</normalizedToken>
|
||
length in
|
||
<taxonomicName id="7BBDDD48E980F88A4731AD48C1685B76" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="5CE183A942D441669FE3A1916DB4DD54" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="DD2F78469937C19E447F8A19E3C06124" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 6C); proportionally wider male pleon (compare Fig.
|
||
<figureCitation id="3B67A44A0448C1A0D102E49B802B48CC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2C</figureCitation>
|
||
with
|
||
<bibRefCitation id="91B3F527A0481D17CECA09A04DDF2A3F" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 2C); male thoracic sternite 8 being fully concealed when the pleon is closed (Fig.
|
||
<figureCitation id="5B89462374BCBBA40147D5A990FBB308" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2C</figureCitation>
|
||
) (vs. partially exposed in
|
||
<taxonomicName id="A09B87C56EC0E2240F554D79B5057A63" lsidName="D. cereum" pageId="0" pageNumber="1" rank="species" species="cereum">
|
||
<emphasis id="325095CA805389F075334E4670554F8C" italics="true" pageId="0" pageNumber="1">D. cereum</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="DD5119F6FF1D5F3CAF1B5258D6430150" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 3E, F); and its relatively shorter and stouter walking legs (Fig.
|
||
<figureCitation id="F060E7E0C5B1F4D8C9C8B4D4370B0951" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) (see
|
||
<bibRefCitation id="CA2D47A1C37515B51ADE5D6FE75C6167" DOI="https://doi.org/10.11646/zootaxa.4232.1.5" author="Huang, C" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="71 - 84" refId="B22" refString="Huang, C, Shih, HT, Ng, PKL, 2017b. A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia. Zootaxa 4232: 71 - 84, DOI: https://doi.org/10.11646/zootaxa.4232.1.5" title="A new genus and new species of Potamidae (Crustacea: Decapoda: Brachyura: Potamoidea), the first stygomorphic cave crab known from China and East Asia." url="https://doi.org/10.11646/zootaxa.4232.1.5" volume="4232" year="2017 b">Huang et al. 2017b</bibRefCitation>
|
||
: fig. 2A).
|
||
</paragraph>
|
||
<paragraph id="D7F11A4E85D3FA3DE6BA7F8DCAEFCCFA" pageId="0" pageNumber="1">
|
||
<taxonomicName id="7F5D991DDE5172BF71B228191287DE43" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon typhlops" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="typhlops">
|
||
<emphasis id="8CC1D5C97A3E347E81A57A3F0853A2B7" italics="true" pageId="0" pageNumber="1">Phasmon typhlops</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. is similar to
|
||
<taxonomicName id="54F07F085C9AC6D9A44E507DE37A06D1" authorityName="Holthuis" authorityYear="1979" class="Malacostraca" family="Potamidae" genus="Cerberusa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cerberusa caeca" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="caeca">
|
||
<emphasis id="F78704E6A838A9470481CDABFD91DFA9" italics="true" pageId="0" pageNumber="1">Cerberusa caeca</emphasis>
|
||
</taxonomicName>
|
||
in general physiognomy and size. However, the new species can immediately be distinguished by its proportionally wider carapace (CW/CL=1.6 vs. 1.3-1.4 in
|
||
<taxonomicName id="BF38C3DD22B4C6DCADF53B449921BFDD" lsidName="C. caeca" pageId="0" pageNumber="1" rank="species" species="caeca">
|
||
<emphasis id="88B321FE3E8A4D382E94E90ECE85A4C7" italics="true" pageId="0" pageNumber="1">C. caeca</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="5C51383BD7CC287C8DECBBAA9B7E7584" author="Holthuis, LB" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="0" pageNumber="1" pagination="1 - 47" refId="B13" refString="Holthuis, LB, 1979. Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo. Zoologische Verhandelingen, Leiden 171: 1 - 47" title="Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo." volume="171" year="1979">Holthuis 1979</bibRefCitation>
|
||
: pl. 8); almost indiscernible postorbital cristae (Fig.
|
||
<figureCitation id="582E2D31195210EF8E490F0C2272B4E9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) (vs. low, indicated by a transverse row of granules in
|
||
<taxonomicName id="1A5220F5A5CA5CC3900F3B3652280FE3" lsidName="C. caeca" pageId="0" pageNumber="1" rank="species" species="caeca">
|
||
<emphasis id="1C806192B08B3758DA41900C5CC6E26D" italics="true" pageId="0" pageNumber="1">C. caeca</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="6353F01AEC4ADC1D2E200D44A4756E1C" author="Holthuis, LB" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="0" pageNumber="1" pagination="1 - 47" refId="B13" refString="Holthuis, LB, 1979. Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo. Zoologische Verhandelingen, Leiden 171: 1 - 47" title="Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo." volume="171" year="1979">Holthuis 1979</bibRefCitation>
|
||
: fig. 3A); proportionally wider male pleon (compare Fig.
|
||
<figureCitation id="9BA736173245B15D3ED45DF6EC156BC6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2C</figureCitation>
|
||
with
|
||
<bibRefCitation id="D3E0A2A5EAE416390F0E70EDB60F5165" author="Holthuis, LB" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="0" pageNumber="1" pagination="1 - 47" refId="B13" refString="Holthuis, LB, 1979. Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo. Zoologische Verhandelingen, Leiden 171: 1 - 47" title="Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo." volume="171" year="1979">Holthuis 1979</bibRefCitation>
|
||
: fig. 3C); and its slightly sinuous G1 (Figs
|
||
<figureCitation id="2EAA53777CFD5B6DF227459FD4A0194E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982. Left maxilliped 3 (A) left G 2, pleonal view (B) left G 1, pleonal view (C) major cheliped (D) minor cheliped (E) colour in life (F). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.58854.figure3" httpUri="https://binary.pensoft.net/fig/492917" pageId="0" pageNumber="1">3C</figureCitation>
|
||
,
|
||
<figureCitation id="6886957587EC0FBBBB9B6D9B83272EF0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phasmon typhlops gen. nov. et sp. nov., gonopods: left G 1, abdominal view (A) left G 1, sternal view (B) left G 2, mesial view (C) left G 2, pleonal view (D). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1008.58854.figure4" httpUri="https://binary.pensoft.net/fig/492918" pageId="0" pageNumber="1">4A, B</figureCitation>
|
||
) (vs. strongly bent outwards in
|
||
<taxonomicName id="7630923B48B026030638B4838162827A" lsidName="C. caeca" pageId="0" pageNumber="1" rank="species" species="caeca">
|
||
<emphasis id="24D3542AC34BF945C6D606F56BF23DEE" italics="true" pageId="0" pageNumber="1">C. caeca</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="D39F34EFB270DB7385BB6DE4AD373429" author="Holthuis, LB" journalOrPublisher="Zoologische Verhandelingen, Leiden" pageId="0" pageNumber="1" pagination="1 - 47" refId="B13" refString="Holthuis, LB, 1979. Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo. Zoologische Verhandelingen, Leiden 171: 1 - 47" title="Cavernicolous and terrestrial decapod Crustacea from northern Sarawak, Borneo." volume="171" year="1979">Holthuis 1979</bibRefCitation>
|
||
: fig. 3D).
|
||
</paragraph>
|
||
<paragraph id="8855BF9ECC995BB6F44C08498FB5CA44" pageId="0" pageNumber="1">
|
||
The G1 characteristics of
|
||
<taxonomicName id="EE941CECE3D950087B0A83F11F9A2479" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="24ACF5F0F5D084E92D5660395EA8A711" italics="true" pageId="0" pageNumber="1">Phasmon</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. are rather unremarkable and particularly similar to those of
|
||
<taxonomicName id="0BFF538DFFAF43C66099B3F5DAE65D75" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7E8EE489D1C888556548A327210208DB" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="100274ACAA4CD149C0FFD4135F91F2C7" authorityName="Huang, Shih & Ng" authorityYear="2017" class="Malacostraca" family="Potamidae" genus="Diyutamon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Diyutamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="60F68DD75B95EC2C4D34CF4EE0C688D3" italics="true" pageId="0" pageNumber="1">Diyutamon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="9B84AB3C1186DAFA2D9BD77A0BFBB534" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D1217C11E9EEABBFCA5FA6731EB6781E" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
is also found in Guangxi and includes two cavernicolous species,
|
||
<taxonomicName id="9F07C20B68155A4C379F3E0B1606456E" lsidName="C. dashiwei" pageId="0" pageNumber="1" rank="species" species="dashiwei">
|
||
<emphasis id="60DCC992BDFE33AA4F3565A18971A002" italics="true" pageId="0" pageNumber="1">C. dashiwei</emphasis>
|
||
</taxonomicName>
|
||
Ng, 2017 and
|
||
<taxonomicName id="65A5279AAD66968DD62F3F99720F6CB6" lsidName="C. clarkei" pageId="0" pageNumber="1" rank="species" species="clarkei">
|
||
<emphasis id="C477CDFFAA9B4A1463C5BB9C62261127" italics="true" pageId="0" pageNumber="1">C. clarkei</emphasis>
|
||
</taxonomicName>
|
||
Ng, 2017, of which the latter displays evidence of stygomorphism in reduced body pigmentation and well-developed, albeit proportionally smaller eyes than epigean congeners (
|
||
<bibRefCitation id="58FC5CD4DB0C298A734566C79FEA084C" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="E077B2B20FC99FC366C60258D888233B" authorityName="Huang & Ahyong & Shih" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Phasmon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phasmon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="54F1D25BE79BEF800A865BBCB0A58681" italics="true" pageId="0" pageNumber="1">Phasmon</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. is readily distinguished from
|
||
<taxonomicName id="CED7B219F350538C44121229D876961B" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61AAE9BD8298B38CC10AAA31FAF42B98" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
in: the proportionally wider carapace (CW/CL=1.6 vs. 1.3-1.4 in
|
||
<taxonomicName id="C762A1106948A308B3DEC66FA6CCF2B3" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6AA4F7DD5D55A216293709306349F230" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="F10356CC0A7D5B85A10466F4EE487C11" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
: figs 2, 6;
|
||
<bibRefCitation id="A798787D9CBFF8DD18EB34132F203860" DOI="https://doi.org/10.7717/peerj.5947" author="Zou, JX" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" refId="B51" refString="Zou, JX, Bai, J, Zhou, XM, 2018. A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e5947. https://doi.org/10.7717/peerj.5947" title="A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e 5947." url="https://doi.org/10.7717/peerj.5947" year="2018">Zou et al. 2018</bibRefCitation>
|
||
: fig. 2); the frontal margin being continuous with the supraorbital margin, forming an almost straight transverse margin in dorsal view (Fig.
|
||
<figureCitation id="69B41876E048A614E62CCAE9EACC621F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) (vs. supraorbital margin distinctly concave in dorsal view in
|
||
<taxonomicName id="3767F217038D84627E279AE23C3CA6FF" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F1393A36DC448470E53A77E63DDC65" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="60EE9C978F4EDCA93735D42423A67B90" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
: figs 2, 6;
|
||
<bibRefCitation id="0AB97A6A8203861FBDCD781B7E95CCEF" DOI="https://doi.org/10.7717/peerj.5947" author="Zou, JX" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" refId="B51" refString="Zou, JX, Bai, J, Zhou, XM, 2018. A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e5947. https://doi.org/10.7717/peerj.5947" title="A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e 5947." url="https://doi.org/10.7717/peerj.5947" year="2018">Zou et al. 2018</bibRefCitation>
|
||
: fig. 2); the vestigial, unpigmented eyes (Fig.
|
||
<figureCitation id="5830AFC2A5F51E32B89C1584BA10DDDA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2A</figureCitation>
|
||
) (vs. well-developed, pigmented eyes in
|
||
<taxonomicName id="B1359BC09AFA3C5D5D44370CADE1E349" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="35D70DBE02C3F63282C9DA9E4D72C59E" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="DEE32EFE21FCCD4D2C54414477F84BBE" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
: fig. 6); the almost indiscernible epigastric cristae and postorbital cristae (Fig.
|
||
<figureCitation id="58399C8AAEB37FEBBC329525B99E8E57" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A) female paratype (22.1 x 13.7 mm), AM P. 105524 (B). Dorsal habitus." figureDoi="10.3897/zookeys.1008.58854.figure1" httpUri="https://binary.pensoft.net/fig/492915" pageId="0" pageNumber="1">1</figureCitation>
|
||
) (vs. clearly discernible in
|
||
<taxonomicName id="F918798829039AC272810563B8133251" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EE92AD8FA433591B78DA193B395F3A74" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="C478AD1C4EC785C24FEC8D777C9020EB" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
: figs 2, 6;
|
||
<bibRefCitation id="5206B79F209012DE28E7F64A05DDAE7E" DOI="https://doi.org/10.7717/peerj.5947" author="Zou, JX" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" refId="B51" refString="Zou, JX, Bai, J, Zhou, XM, 2018. A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e5947. https://doi.org/10.7717/peerj.5947" title="A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e 5947." url="https://doi.org/10.7717/peerj.5947" year="2018">Zou et al. 2018</bibRefCitation>
|
||
: fig. 2); the proportionally wider male anterior thoracic sternum (width/length 2.3 vs. 1.6-1.7 in
|
||
<taxonomicName id="7F330233E7B73FE7E0F86619344000AD" authorityName="Dai & Naiyanetr" authorityYear="1994" class="Crustacea" family="Potamidae" genus="Chinapotamon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chinapotamon" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0FBBBB1D0B8C7E5A44AB84ED6621D8D9" italics="true" pageId="0" pageNumber="1">Chinapotamon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="BF9DAF2F8BB7A86E9FF8921188753D6C" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
|
||
: figs 3A, 7A;
|
||
<bibRefCitation id="8E800884CFA9BD9BBA1DD2C355664E4A" DOI="https://doi.org/10.7717/peerj.5947" author="Zou, JX" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" refId="B51" refString="Zou, JX, Bai, J, Zhou, XM, 2018. A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e5947. https://doi.org/10.7717/peerj.5947" title="A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e 5947." url="https://doi.org/10.7717/peerj.5947" year="2018">Zou et al. 2018</bibRefCitation>
|
||
: fig. 3A); and the proportionally wider male pleon (compare Fig.
|
||
<figureCitation id="8CA86E55C94EE4DF5A8B34B85E75681C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phasmon typhlops gen. nov. et sp. nov., male holotype (14.4 x 9.0 mm), SYSBM 001982 (A-D) female paratype (22.1 x 13.7 mm), AM P. 105524 (E, F). Cephalothorax, anterior view (A) anterior thoracic sternum (B) anterior thoracic sternum and pleon, ventral view (C) sterno-pleonal cavity with G 1 in situ, ventral view (D) pleon, ventral view (E) vulvae, ventral view (F)." figureDoi="10.3897/zookeys.1008.58854.figure2" httpUri="https://binary.pensoft.net/fig/492916" pageId="0" pageNumber="1">2C</figureCitation>
|
||
with
|
||
<bibRefCitation id="4A2EDE9F05284AD598A8362FCCF0B755" DOI="https://doi.org/10.18353/crustacea.46.0_1" author="Ng, PKL" journalOrPublisher="Crustacean Research" pageId="0" pageNumber="1" pagination="1 - 16" refId="B30" refString="Ng, PKL, 2017. Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China. Crustacean Research 46: 1 - 16, DOI: https://doi.org/10.18353/crustacea.46.0_1" title="Descriptions of two new cavernicolous species of Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Brachyura: Potamidae) from China." url="https://doi.org/10.18353/crustacea.46.0_1" volume="46" year="2017">Ng 2017</bibRefCitation>
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: figs 3B, 7B;
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<bibRefCitation id="3D7A77CDF1C0AF133A68D7FAFA202F3C" DOI="https://doi.org/10.7717/peerj.5947" author="Zou, JX" journalOrPublisher="Crustaceana" pageId="0" pageNumber="1" refId="B51" refString="Zou, JX, Bai, J, Zhou, XM, 2018. A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e5947. https://doi.org/10.7717/peerj.5947" title="A new species of karst-dwelling freshwater crab of the genus Chinapotamon Dai & Naiyanetr, 1994 (Crustacea: Decapoda: Brachyura: Potamidae), from Guizhou, southwest China. PeerJ 6: e 5947." url="https://doi.org/10.7717/peerj.5947" year="2018">Zou et al. 2018</bibRefCitation>
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: fig. 3B).
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</paragraph>
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</subSubSection>
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</treatment>
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</subSection>
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</document> |