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<mods:title id="827668FA20F5FB14F4554CA2D74EC984">Taxonomic revision of the jumping spider genus Tullgrenella Mello-Leitão, 1941 (Araneae: Salticidae: Freyina)</mods:title>
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<mods:namePart id="2923F40C8367BBCD6E19894D9150FE44">Marta, Kimberly S.</mods:namePart>
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<mods:namePart id="C59350CC90D16E80FB3BBFCFCC2EE972">Bustamante, Abel A.</mods:namePart>
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<taxonomicName id="47B24D53FFE24C286EAE5E1BFD550347" authority="Marta &amp; Hagopian" authorityName="Marta &amp; Hagopian" authorityYear="2024" box="[151,693,1631,1659]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="legalissima" status="sp. nov.">
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Marta &amp; Hagopián
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<figureCitation id="18892A55FFE24C286EAE5EFFFF1003A2" box="[151,240,1668,1695]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="44" pageNumber="45">Figs 7L</figureCitation>
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<figureCitation id="18892A55FFE24C286F8B5EFFFDE303A2" box="[434,515,1668,1695]" captionStart="FIGURE 53" captionStartId="66.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@66.[151,1435,181,1465]" captionTargetPageId="66" captionText="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675934" httpUri="https://zenodo.org/record/10675934/files/figure.png" pageId="44" pageNumber="45">53A–B</figureCitation>
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[
<geoCoordinate id="E5865017FFE24C286FB25F6FFDC70212" box="[395,551,1812,1839]" degrees="26" direction="south" minutes="55" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="23.8" value="-26.923277">26°55’23.8”S</geoCoordinate>
<geoCoordinate id="E5865017FFE24C286C0A5F6FFD3C0213" box="[563,732,1812,1838]" degrees="49" direction="west" minutes="03" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="27.6" value="-49.057667">49°03’27.6”W</geoCoordinate>
], 
<date id="F40C1010FFE24C286CCD5F6FFC880212" box="[756,872,1812,1839]" pageId="44" pageNumber="45" value="2004-05-18">
<collectingDate id="E448E9F8FFE24C286CCD5F6FFC880212" box="[756,872,1812,1839]" pageId="44" pageNumber="45" value="2004-05-18">18.V.2004</collectingDate>
</date>
– 
<date id="F40C1010FFE24C286D415F6FFC0C0212" box="[888,1004,1812,1839]" pageId="44" pageNumber="45" value="2005-02-28">
<collectingDate id="E448E9F8FFE24C286D415F6FFC0C0212" box="[888,1004,1812,1839]" pageId="44" pageNumber="45" value="2005-02-28">28.II.2005</collectingDate>
</date>
, 
<collectionCode id="E6A3AE15FFE24C286DC45F6FFBF30213" box="[1021,1043,1812,1838]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="44" pageNumber="45">R</collectionCode>
. 
<collectorName id="2D475306FFE24C286A2E5F6FFB4A0213" box="[1047,1194,1812,1838]" pageId="44" pageNumber="45">C. Francisco</collectorName>
leg.
</materialsCitation>
<materialsCitation id="30DA3C8DFFE24C286AD55F6FFDB1026F" collectingDate="2004-05-18" collectingDateMax="2005-02-28" collectingDateMin="2004-05-18" collectionCode="IBSP" collectorName="Parque Municipal das Nascentes do Ribeirao Garcia &amp; C. Francisco" country="Brazil" latitude="-26.923277" location="Nascentes do Ribeirao Garcia" longLatPrecision="1" longitude="-49.057667" municipality="Blumenau" pageId="44" pageNumber="45" specimenCode="IBSP 118217" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina" typeStatus="paratype">
<emphasis id="B2C6EAC2FFE24C286AD55F6FFA8D0213" bold="true" box="[1260,1389,1812,1838]" pageId="44" pageNumber="45">
<typeStatus id="5F098872FFE24C286AD55F6FFA880213" box="[1260,1384,1812,1838]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="96B4FD59FFE24C286B405F6FFA7C0213" box="[1401,1436,1812,1838]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
(
<specimenCode id="D0149EABFFE24C286EA65F43FEDA026F" box="[159,314,1848,1874]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="44" pageNumber="45">IBSP 118217</specimenCode>
) same data as holotype
</materialsCitation>
; 
<materialsCitation id="30DA3C8DFFE24C286C665F43FC90024B" collectingDate="2017-04-13" collectingDateMax="2017-04-16" collectingDateMin="2017-04-13" collectorName="D. Hagopian" country="Uruguay" latitude="-32.13386" location="Aduana" longLatPrecision="1" longitude="-53.732582" municipality="Paso Centurion" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Cerro Largo" typeStatus="paratype">
<specimenCount id="96B4FD59FFE24C286C665F43FD62026F" box="[607,642,1848,1874]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
(FCE-Ar 9066), 
<collectingCountry id="F8A57640FFE24C286D745F43FC36026F" box="[845,982,1848,1874]" name="Uruguay" pageId="44" pageNumber="45">URUGUAY</collectingCountry>
, 
<emphasis id="B2C6EAC2FFE24C286DDC5F42FB94026F" box="[997,1140,1849,1875]" italics="true" pageId="44" pageNumber="45">
<collectingRegion id="4276F832FFE24C286DDC5F42FB94026F" box="[997,1140,1849,1875]" country="Uruguay" name="Cerro Largo" pageId="44" pageNumber="45">Cerro Largo</collectingRegion>
</emphasis>
, 
<collectingMunicipality id="6069ACAAFFE24C286ABF5F42FAD8026F" box="[1158,1336,1848,1875]" pageId="44" pageNumber="45">Paso Centurión</collectingMunicipality>
, 
<location id="856D600BFFE24C286B7C5F43FA7C026F" LSID="urn:lsid:plazi:treatment:081B87C6FFE24C2A6EAE5E1BFCCD07CC:856D600BFFE24C286B7C5F43FA7C026F" box="[1349,1436,1848,1874]" country="Uruguay" latitude="-32.13386" longLatPrecision="1" longitude="-53.732582" municipality="Paso Centurion" name="Aduana" pageId="44" pageNumber="45" stateProvince="Cerro Largo">Aduana</location>
[
<geoCoordinate id="E5865017FFE24C286EA75F27FEDA024B" box="[158,314,1884,1910]" degrees="32" direction="south" minutes="08" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="01.9" value="-32.13386">32°08’01.9”S</geoCoordinate>
<geoCoordinate id="E5865017FFE24C286F785F27FE0A024A" box="[321,490,1884,1911]" degrees="53" direction="west" minutes="43" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="57.3" value="-53.732582">53°43’57.3”W</geoCoordinate>
], 
<date id="F40C1010FFE24C286FC45F27FD45024B" box="[509,677,1884,1910]" pageId="44" pageNumber="45" value="2017-04-13" valueMax="2017-04-16" valueMin="2017-04-13">
<collectingDate id="E448E9F8FFE24C286FC45F27FD45024B" box="[509,677,1884,1910]" pageId="44" pageNumber="45" value="2017-04-13" valueMax="2017-04-16" valueMin="2017-04-13">13–16.IV.2017</collectingDate>
</date>
, 
<collectorName id="2D475306FFE24C286C885F26FCA0024B" box="[689,832,1884,1911]" pageId="44" pageNumber="45">D. Hagopián</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="800D36D0FFE24C286EFE5FFBFBBF02DE" blockId="44.[151,1437,1776,2019]" pageId="44" pageNumber="45">
<emphasis id="B2C6EAC2FFE24C286EFE5FFBFE1802A7" bold="true" box="[199,504,1920,1946]" pageId="44" pageNumber="45">Other material examined.</emphasis>
<materialsCitation id="30DA3C8DFFE24C286FC45FFAFC6D0282" collectingDate="2006-04-24" collectionCode="MCTP" collectorName="A. A. Lise" country="Brazil" latitude="-29.480696" location="Potreiro Velho" longLatPrecision="1" longitude="-50.174446" municipality="Sao Francisco de Paula" pageId="44" pageNumber="45" specimenCode="MCTP 35720" specimenCount="1" specimenCount-female="1" stateProvince="Rio Grande do Sul">
<collectingCountry id="F8A57640FFE24C286FC45FFAFD8502A6" box="[509,613,1921,1947]" name="Brazil" pageId="44" pageNumber="45">BRAZIL</collectingCountry>
, 
<emphasis id="B2C6EAC2FFE24C286C565FFAFCDB02A7" box="[623,827,1920,1946]" italics="true" pageId="44" pageNumber="45">
<collectingRegion id="4276F832FFE24C286C565FFAFCDB02A7" box="[623,827,1920,1946]" country="Brazil" name="Rio Grande do Sul" pageId="44" pageNumber="45">
<taxonomicName id="47B24D53FFE24C286C565FFAFD1102A7" box="[623,753,1920,1946]" class="Arachnida" family="Salticidae" genus="Rio" kingdom="Animalia" order="Araneae" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="grande">Rio Grande</taxonomicName>
do Sul
</collectingRegion>
</emphasis>
, 
<collectingMunicipality id="6069ACAAFFE24C286D715FFBFBAA02A7" box="[840,1098,1920,1946]" pageId="44" pageNumber="45">São Francisco de Paula</collectingMunicipality>
, 
<location id="856D600BFFE24C286A6D5FFBFB1402A7" LSID="urn:lsid:plazi:treatment:081B87C6FFE24C2A6EAE5E1BFCCD07CC:856D600BFFE24C286A6D5FFBFB1402A7" box="[1108,1268,1920,1946]" country="Brazil" latitude="-29.480696" longLatPrecision="1" longitude="-50.174446" municipality="Sao Francisco de Paula" name="Potreiro Velho" pageId="44" pageNumber="45" stateProvince="Rio Grande do Sul">Potreiro Velho</location>
[
<geoCoordinate id="E5865017FFE24C286B395FFBFA7C02A6" box="[1280,1436,1920,1947]" degrees="29" direction="south" minutes="28" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="50.5" value="-29.480696">29°28’50.5”S</geoCoordinate>
<geoCoordinate id="E5865017FFE24C286EAE5FDFFEA00282" box="[151,320,1956,1983]" degrees="50" direction="west" minutes="10" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="28.0" value="-50.174446">50°10’28.0”W</geoCoordinate>
], 
<date id="F40C1010FFE24C286F605FDFFE380283" box="[345,472,1956,1982]" pageId="44" pageNumber="45" value="2006-04-24">
<collectingDate id="E448E9F8FFE24C286F605FDFFE380283" box="[345,472,1956,1982]" pageId="44" pageNumber="45" value="2006-04-24">24.IV.2006</collectingDate>
</date>
, 
<collectorName id="2D475306FFE24C286FD15FDEFDBC0283" box="[488,604,1956,1983]" pageId="44" pageNumber="45">A.A. Lise</collectorName>
leg., 
<specimenCount id="96B4FD59FFE24C286C9E5FDFFD2A0283" box="[679,714,1956,1982]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
(
<specimenCode id="D0149EABFFE24C286CE65FDFFC650282" box="[735,901,1956,1983]" collectionCode="MCTP" country="Brazil" httpUri="http://grbio.org/cool/rq94-ajyg" name="Museu de Ciencias" pageId="44" pageNumber="45">MCTP 35720</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="30DA3C8DFFE24C286DA45FDFFBBB02DE" collectingDate="2019-10-21" country="Uruguay" latitude="-34.528057" location="Sierra del Carape" longLatPrecision="1" longitude="-54.969166" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1" stateProvince="Maldonado">
<collectingCountry id="F8A57640FFE24C286DA45FDFFBC60283" box="[925,1062,1956,1982]" name="Uruguay" pageId="44" pageNumber="45">URUGUAY</collectingCountry>
, 
<emphasis id="B2C6EAC2FFE24C286A015FDFFB590283" box="[1080,1209,1956,1982]" italics="true" pageId="44" pageNumber="45">
<collectingRegion id="4276F832FFE24C286A015FDFFB590283" box="[1080,1209,1956,1982]" country="Uruguay" name="Maldonado" pageId="44" pageNumber="45">Maldonado</collectingRegion>
</emphasis>
, 
<location id="856D600BFFE24C286AF45FDFFA7C0283" LSID="urn:lsid:plazi:treatment:081B87C6FFE24C2A6EAE5E1BFCCD07CC:856D600BFFE24C286AF45FDFFA7C0283" box="[1229,1436,1956,1982]" country="Uruguay" latitude="-34.528057" longLatPrecision="1" longitude="-54.969166" name="Sierra del Carape" pageId="44" pageNumber="45" stateProvince="Maldonado">Sierra del Carapé</location>
[
<geoCoordinate id="E5865017FFE24C286EA75FB3FEDA02DF" box="[158,314,1992,2018]" degrees="34" direction="south" minutes="31" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="41.0" value="-34.528057">34°31’41.0”S</geoCoordinate>
<geoCoordinate id="E5865017FFE24C286F785FB3FE0A02DE" box="[321,490,1992,2019]" degrees="54" direction="west" minutes="58" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="09.0" value="-54.969166">54°58’09.0”W</geoCoordinate>
], 
<date id="F40C1010FFE24C286FC45FB3FD9602DF" box="[509,630,1992,2018]" pageId="44" pageNumber="45" value="2019-10-21">
<collectingDate id="E448E9F8FFE24C286FC45FB3FD9602DF" box="[509,630,1992,2018]" pageId="44" pageNumber="45" value="2019-10-21">21.X.2019</collectingDate>
</date>
, collected under rock, 
<specimenCount id="96B4FD59FFE24C286D4D5FB3FC7602DF" box="[884,918,1992,2018]" count="1" pageId="44" pageNumber="45" type="female">1♀</specimenCount>
(FCE-Ar 12155)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="D4CD6658FFE34C296EAE5DA9FBEB0333" ID-DOI="http://doi.org/10.5281/zenodo.10675894" ID-Zenodo-Dep="10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,250,1490,1514]" targetBox="[151,1383,199,1452]" targetPageId="45" targetType="figure">
<paragraph id="800D36D0FFE34C296EAE5DA9FBEB0333" blockId="45.[151,1437,1490,1550]" pageId="45" pageNumber="46">
<emphasis id="B2C6EAC2FFE34C296EAE5DA9FEFF00D7" bold="true" box="[151,287,1490,1514]" pageId="45" pageNumber="46">FIGURE 35.</emphasis>
Female of 
<taxonomicName id="47B24D53FFE34C296FB75DA8FD3D00D7" authority="Marta &amp; Hagopian" authorityName="Marta &amp; Hagopian" authorityYear="2024" box="[398,733,1490,1514]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="legalissima" status="sp. nov.">
<emphasis id="B2C6EAC2FFE34C296FB75DA8FDF800D7" box="[398,536,1490,1514]" italics="true" pageId="45" pageNumber="46">T. legalissima</emphasis>
Marta &amp; Hagopián
</taxonomicName>
, 
<emphasis id="B2C6EAC2FFE34C296CDF5DA9FCD700D4" bold="true" box="[742,823,1490,1514]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A9F557B9FFE34C296CDF5DA9FCD700D4" box="[742,823,1490,1514]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C8A8655BFFE34C296EFE5E47FBFA0347" pageId="45" pageNumber="46" type="etymology">
<paragraph id="800D36D0FFE34C296EFE5E47FBFA0347" blockId="45.[151,1437,1596,2019]" pageId="45" pageNumber="46">
<emphasis id="B2C6EAC2FFE34C296EFE5E47FEAE036B" bold="true" box="[199,334,1596,1622]" pageId="45" pageNumber="46">Etymology.</emphasis>
The name means “very cool” in Portuguese, a comment frequently made by the authors of this species whenever they find new species of jumping spiders. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C8A8655BFFE34C296EFE5EFFFC050212" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="800D36D0FFE34C296EFE5EFFFC050212" blockId="45.[151,1437,1596,2019]" pageId="45" pageNumber="46">
<emphasis id="B2C6EAC2FFE34C296EFE5EFFFEA203A3" bold="true" box="[199,322,1668,1694]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
<taxonomicName id="47B24D53FFE34C296F715EFFFCCE03A3" authority="Marta &amp; Hagopian" authorityName="Marta &amp; Hagopian" authorityYear="2024" box="[328,814,1668,1695]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="legalissima" status="sp. nov.">
<emphasis id="B2C6EAC2FFE34C296F715EFFFDB603A3" box="[328,598,1668,1694]" italics="true" pageId="45" pageNumber="46">Tullgrenella legalissima</emphasis>
Marta &amp; Hagopián
</taxonomicName>
<emphasis id="B2C6EAC2FFE34C296D0D5EFFFC6D03A3" bold="true" box="[820,909,1668,1694]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A9F557B9FFE34C296D0D5EFFFC6D03A3" box="[820,909,1668,1694]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="47B24D53FFE34C296A305EFEFB9B03A3" authorityName="Galiano" authorityYear="1970" box="[1033,1147,1669,1694]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="serrana">
<emphasis id="B2C6EAC2FFE34C296A305EFEFB9B03A3" box="[1033,1147,1669,1694]" italics="true" pageId="45" pageNumber="46">T. serrana</emphasis>
</taxonomicName>
, 
<taxonomicName id="47B24D53FFE34C296AB05EFEFA9503A2" authority="Marta" authorityName="Marta" authorityYear="2024" box="[1161,1397,1668,1695]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="aisenbergae" status="sp. nov.">
<emphasis id="B2C6EAC2FFE34C296AB05EFEFACB03A3" box="[1161,1323,1668,1694]" italics="true" pageId="45" pageNumber="46">T. aisenbergae</emphasis>
Marta
</taxonomicName>
<taxonomicNameLabel id="A9F557B9FFE34C296B425EFFFF2903FE" pageId="45" pageNumber="46" rank="species">
<emphasis id="B2C6EAC2FFE34C296B425EFFFF2303FF" bold="true" pageId="45" pageNumber="46">sp. nov</emphasis>
.
</taxonomicNameLabel>
, and 
<taxonomicName id="47B24D53FFE34C296F315ED2FD9D03FE" authority="Marta &amp; Bustamante" authorityName="Marta &amp; Bustamante" authorityYear="2024" box="[264,637,1704,1731]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="sciosciae" status="sp. nov.">
<emphasis id="B2C6EAC2FFE34C296F315ED2FE6B03FF" box="[264,395,1705,1730]" italics="true" pageId="45" pageNumber="46">T. sciosciae</emphasis>
Marta &amp; Bustamante
</taxonomicName>
<emphasis id="B2C6EAC2FFE34C296CBD5ED3FD3E03FF" bold="true" box="[644,734,1704,1730]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A9F557B9FFE34C296CBD5ED3FD3E03FF" box="[644,734,1704,1730]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the C-shaped initial copulatory ducts (
<figureCitation id="18892A55FFE34C296A8C5ED3FAD703FF" box="[1205,1335,1704,1730]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="45" pageNumber="46">Figs 7K–N</figureCitation>
), but can be distinguished from 
<taxonomicName id="47B24D53FFE34C296FAB5EB6FD9F03DA" authority="Marta" authorityName="Marta" authorityYear="2024" box="[402,639,1740,1767]" class="Arachnida" family="Salticidae" genus="Tullgrenella" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="aisenbergae" status="sp. nov.">
<emphasis id="B2C6EAC2FFE34C296FAB5EB6FDD603DB" box="[402,566,1740,1766]" italics="true" pageId="45" pageNumber="46">T. aisenbergae</emphasis>
Marta
</taxonomicName>
<emphasis id="B2C6EAC2FFE34C296CBF5EB7FD0003DB" bold="true" box="[646,736,1740,1766]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A9F557B9FFE34C296CBF5EB7FD0003DB" box="[646,736,1740,1766]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the sclerotized, posterior-ventral space between copulatory opening and posteriorly-opening coupling pocket (
<figureCitation id="18892A55FFE34C296CE35E8BFCD30236" box="[730,819,1776,1803]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="45" pageNumber="46">Figs 7L</figureCitation>
vs 7M, 35C–D vs 37C–D). It can also be recognized from aforementioned congeners by the ovoid spermathecae (
<figureCitation id="18892A55FFE34C296D075F6FFC780212" box="[830,920,1812,1839]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="45" pageNumber="46">Figs 7L</figureCitation>
, 
<figureCitation id="18892A55FFE34C296D9D5F6FFC380212" box="[932,984,1812,1839]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="45" pageNumber="46">35D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8A8655BFFE34C2A6EFE5F43FF380700" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="800D36D0FFE34C296EFE5F43FDFD026F" blockId="45.[151,1437,1596,2019]" box="[199,541,1848,1875]" pageId="45" pageNumber="46">
<emphasis id="B2C6EAC2FFE34C296EFE5F43FEB9026F" bold="true" box="[199,345,1848,1874]" pageId="45" pageNumber="46">Description.</emphasis>
<emphasis id="B2C6EAC2FFE34C296F595F43FE7A026F" box="[352,410,1848,1874]" italics="true" pageId="45" pageNumber="46">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="800D36D0FFE34C2A6EFE5F27FF380700" blockId="45.[151,1437,1596,2019]" lastBlockId="46.[151,1437,150,753]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="B2C6EAC2FFE34C296EFE5F27FEFB024B" box="[199,283,1884,1910]" italics="true" pageId="45" pageNumber="46">Female</emphasis>
(FCE-Ar 12155). Total length 6.10. Carapace length 2.80; width 1.95; height 1.12. Cephalic region length 1.65; thoracic region length 1.15. Ocular area length 1.11; Anterior eyes row 1.50 wide; Posterior eyes row 1.50 wide; Distance ALE–PME 0.35; PME–PLE 0.30. Ocular diameter AME 0.45; ALE 0.24; PME 0.09; PLE 0.21. Clypeus height 0.15. Chelicerae, paturon length 0.75; 0.50 wide; Fang 0.31 length. Endites length 0.50; 0.45 wide. Labium length 0.21; 0.40 wide. Sternum length 1.09; 0.70 wide. Abdomen length 3.31; 2.05 wide, height 1.75. Legs, femur length I: 1.31; II: 1.23; III: 1.21; IV: 1.45; patella I: 0.85; II: 0.81; III: 0.71; IV: 0.72; tibia I: 0.75; II: 0.65; III: 0.73; IV: 1.11; metatarsus I: 0.60; II: 0.50; III: 0.71; IV: 1.20; tarsus I: 0.45; II: 0.40; III: 0.50; IV: 0.81. Formula legs: 4132. Palp, length of femur 0.70, patella 0.39, tibia 0.30, tarsus 0.45. Legs macrosetae, femur I–II: d1–1–1, p2di; III–IV: d1–1–1, p0–1–2di, r0–1–2di. Patella I–II: 0; III–IV: p1, r1. Tibia I: v2–2–2, p1subdi; II: v1r–1–2, p1di; III: v2di p1–1–1, r1–1–1; IV: v0–1p–2di, p1–1–1, r1–1–1. Metatarsus I: v0–2–2; II: v0–2–2, p1di; III: v0–1–2, p1di, r1di, d0–2–2; IV: v0–2–2, p1di, r1di, d2–2–2. Carapace and abdomen as in genus (
<figureCitation id="18892A55FFE04C2A6B3B5914FA7004B4" box="[1282,1424,366,393]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="46" pageNumber="47">Figs 35A–B</figureCitation>
). Spinnerets: coloration as abdomen’s dorsum (
<figureCitation id="18892A55FFE04C2A6C9959E8FCD00490" box="[672,816,402,429]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="46" pageNumber="47">Figs 35A–B</figureCitation>
). Epigynum: copulatory opening posterior-medial into a fillet that widens posteriorly forming pockets near a short pCP (
<figureCitation id="18892A55FFE04C2A6DB859CCFC3C04EC" box="[897,988,439,465]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="46" pageNumber="47">Figs 7L</figureCitation>
, 
<figureCitation id="18892A55FFE04C2A6DD059CDFBDE04EC" box="[1001,1086,438,465]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="46" pageNumber="47">35C–D</figureCitation>
). Copulatory ducts arranged in one monoplanar spiral, C-shaped, with one medial-dorsal thinner twist right before reaching the spermathecae (
<figureCitation id="18892A55FFE04C2A6B5359A0FF580724" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="46" pageNumber="47">Figs 7L</figureCitation>
, 
<figureCitation id="18892A55FFE04C2A6EFF5985FF190724" box="[198,249,510,537]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="46" pageNumber="47">35D</figureCitation>
). Spermathecae ovoid, with the base of fertilization ducts placed on the anterior/dorsal surface (
<figureCitation id="18892A55FFE04C2A6B055984FA770724" box="[1340,1431,511,537]" captionStart="FIGURE 7" captionStartId="13.[151,250,1455,1479]" captionTargetBox="[164,1432,191,1420]" captionTargetId="figure-17@13.[151,1435,181,1431]" captionTargetPageId="13" captionText="FIGURE 7. Illustrations of female epigynum of Tullgrenella, dorsal view. A–E morenensis group: A T. morenensis (Tullgren, 1905) (MCN 19940), B T. guayapae Galiano, 1970 (MCN 55520), C T. brescoviti Estol, Marta &amp; Rodrigues, 2020 (MCN 24104), D T. musica (Mello-Leitão,1945) (MCN 53372), E T. corrugata Galiano, 1981 (MACN-Ar 7206). F–H quadripunctata group: F T. quadripunctata (Mello-Leitão, 1944) (MCTP 2239), G T. nadjae Estol, Marta &amp; Rodrigues, 2020 (MCN 56814), H T. lunata (Mello-Leitão, 1944) (MACN-Ar 6214). I–J melanica group: I T. melanica (Mello-Leitão, 1941) (MACN-Ar 6244), J T. yungae Galiano, 1970 (MCTP 39713). K–O serrana group: K T. serrana Galiano, 1970 (MCTP 12173), L T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228), M T. aisenbergae Marta, sp. nov. (IBSP 123129), N T. gertschi Galiano, 1981 (MACN-Ar 7206), O T. sciosciae Marta &amp; Bustamante, sp. nov. (MCTP 31135). P–Q peniaflorensis group: P T. peniaflorensis Galiano, 1970 (MACN-Ar 6242), Q T. selenita Galiano, 1970 (FNAC 36). Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675804" httpUri="https://zenodo.org/record/10675804/files/figure.png" pageId="46" pageNumber="47">Figs 7L</figureCitation>
, 
<figureCitation id="18892A55FFE04C2A6EAE5A59FF2B0700" box="[151,203,546,573]" captionStart="FIGURE 35" captionStartId="45.[151,250,1490,1514]" captionTargetBox="[151,1383,199,1452]" captionTargetId="figure-17@45.[151,1435,181,1465]" captionTargetPageId="45" captionText="FIGURE 35. Female of T. legalissima Marta &amp; Hagopián, sp. nov. (IBSP 118228).A habitus, dorsal view. B same, ventral view. C epigynum, ventral view. D same, dorsal view. Scale bars: A–B 1 mm, C–D 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10675894" httpUri="https://zenodo.org/record/10675894/files/figure.png" pageId="46" pageNumber="47">35D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8A8655BFFE04C2A6EFE5A3DFCCD07CC" pageId="46" pageNumber="47" type="distribution">
<paragraph id="800D36D0FFE04C2A6EFE5A3DFF1907B8" blockId="46.[151,1437,150,753]" pageId="46" pageNumber="47">
<emphasis id="B2C6EAC2FFE04C2A6EFE5A3DFE81075D" bold="true" box="[199,353,582,608]" pageId="46" pageNumber="47">Distribution.</emphasis>
<collectingCountry id="F8A57640FFE04C2A6F5E5A3CFE4B075C" box="[359,427,583,609]" name="Brazil" pageId="46" pageNumber="47">Brazil</collectingCountry>
(
<collectingRegion id="4276F832FFE04C2A6F835A3CFDBF075C" box="[442,607,583,609]" country="Brazil" name="Santa Catarina" pageId="46" pageNumber="47">Santa Catarina</collectingRegion>
and 
<collectingRegion id="4276F832FFE04C2A6CAD5A3CFC86075C" box="[660,870,583,609]" country="Brazil" name="Rio Grande do Sul" pageId="46" pageNumber="47">
<taxonomicName id="47B24D53FFE04C2A6CAD5A3CFCF5075C" box="[660,789,583,609]" class="Arachnida" family="Salticidae" genus="Rio" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="55" phylum="Arthropoda" rank="species" species="grande">Rio Grande</taxonomicName>
do Sul
</collectingRegion>
) and 
<collectingCountry id="F8A57640FFE04C2A6D9A5A3CFBE5075C" box="[931,1029,583,609]" name="Uruguay" pageId="46" pageNumber="47">Uruguay</collectingCountry>
(
<collectingRegion id="4276F832FFE04C2A6A2A5A3CFB7D075C" box="[1043,1181,583,609]" country="Uruguay" name="Cerro Largo" pageId="46" pageNumber="47">Cerro Largo</collectingRegion>
and 
<collectingRegion id="4276F832FFE04C2A6AEA5A3CFAB4075C" box="[1235,1364,583,609]" country="Uruguay" name="Maldonado" pageId="46" pageNumber="47">Maldonado</collectingRegion>
) (
<figureCitation id="18892A55FFE04C2A6B535A3CFF0C07B8" captionStart="FIGURE 53" captionStartId="66.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@66.[151,1435,181,1465]" captionTargetPageId="66" captionText="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675934" httpUri="https://zenodo.org/record/10675934/files/figure.png" pageId="46" pageNumber="47">Figs 53A–B</figureCitation>
).
</paragraph>
<paragraph id="800D36D0FFE04C2A6EFE5AF4FCCD07CC" blockId="46.[151,1437,150,753]" pageId="46" pageNumber="47">
<emphasis id="B2C6EAC2FFE04C2A6EFE5AF4FE690795" bold="true" box="[199,393,654,681]" pageId="46" pageNumber="47">Natural history.</emphasis>
Some individuals were collected under rocks in 
<collectingCountry id="F8A57640FFE04C2A6DF25AF4FBCF0794" box="[971,1071,655,681]" name="Uruguay" pageId="46" pageNumber="47">Uruguay</collectingCountry>
. This suggests that this species inhabits a soil habitat. It is found only in the East of the Chacoan subregion (
<figureCitation id="18892A55FFE04C2A6DE05AC8FB8507F0" box="[985,1125,690,717]" captionStart="FIGURE 53" captionStartId="66.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@66.[151,1435,181,1465]" captionTargetPageId="66" captionText="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675934" httpUri="https://zenodo.org/record/10675934/files/figure.png" pageId="46" pageNumber="47">Figs 53A–B</figureCitation>
). The variation in altitudinal occurrence of this species is 
<quantity id="474A9B35FFE04C2A6FE15AACFDDB07CC" box="[472,571,727,753]" metricMagnitude="2" metricUnit="m" metricValue="4.58" metricValueMax="9.08" metricValueMin="0.08" pageId="46" pageNumber="47" unit="m" value="458.0" valueMax="908.0" valueMin="8.0">8–908 m</quantity>
a. s. l. (
<figureCitation id="18892A55FFE04C2A6CAA5AACFCC107CC" box="[659,801,726,753]" captionStart="FIGURE 53" captionStartId="66.[151,250,1490,1514]" captionTargetBox="[151,1435,181,1465]" captionTargetId="figure-20@66.[151,1435,181,1465]" captionTargetPageId="66" captionText="FIGURE 53. Neotropical andAndean regionalization (Morrone 2014) and distributional records for species of the serrana group. A Neotropical, South America, Brazilian subregion Chacoan subregion, South America Transition Zone and Andean region. B Southern South America, focusing on species occurrence. C habitat inference, URUGUAY: Sierra de Ríos, photographed by collector Damián Hagopián." figureDoi="http://doi.org/10.5281/zenodo.10675934" httpUri="https://zenodo.org/record/10675934/files/figure.png" pageId="46" pageNumber="47">Figs 53A–B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>