294 lines
33 KiB
XML
294 lines
33 KiB
XML
<document id="FD1B33C6034D5E03B203D49B559DC073" ID-DOI="10.5281/zenodo.191476" ID-GBIF-Dataset="8603d4d1-efc8-44bb-84ae-76d3d8dd8161" ID-ISSN="1175-5326" ID-Zenodo-Dep="191476" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461128727612" checkinUser="plazi" docAuthor="Passos, Paulo, Mueses-Cisneros, Jonh Jairo, Lynch, John D. & Fernandes, Ronaldo" docDate="2009" docId="03D087ECAA477749FF11B43D101D2591" docLanguage="en" docName="zt02293p034.pdf" docOrigin="Zootaxa 2293" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atractus typhon Passos, Mueses-Cisneros, Lynch & Fernandes, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="FFE9FF94AA5E7755FF86B32815112046" masterDocTitle="Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="26" updateTime="1698597252759" updateUser="plazi">
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<mods:mods id="F3B1B04ED93F6FDE7C396E518EBAF04F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EA4827E410A9B160E1A762EC5729A768">
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<mods:title id="F8B702D0B04C51C5F74AB1827889C4EB">Pacific lowland snakes of the genus Atractus (Serpentes: Dipsadidae), with description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="BCC7968280959351BA81CFEEF5671FDA" type="personal">
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<mods:roleTerm id="CC085EF540D5C33E9EA43EE7172C83A3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C5C0E854D435D84F027E87E33F5E0C84">Passos, Paulo</mods:namePart>
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</mods:name>
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<mods:roleTerm id="04F188404245483D3590B1677CA5BD7B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D1A8E851C61A57B923CF7B59AB4F2894">Mueses-Cisneros, Jonh Jairo</mods:namePart>
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</mods:name>
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<mods:name id="605D1C8E32F12A7D11748B123818E4D6" type="personal">
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<mods:roleTerm id="85762C5DA69A7A5123C087799DDF63EE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="177033DFBEFF84E0571B46F7ECDB53FF">Lynch, John D.</mods:namePart>
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</mods:name>
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<mods:name id="94B93E08B412E7302843D8FC5ECA819A" type="personal">
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<mods:roleTerm id="41503B809545ABE21B7E90E460566C52">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1C6F8592AD54D72BA2EF37F0F63796E4">Fernandes, Ronaldo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4F5C088A3B93902BD40F0260E530C105">text</mods:typeOfResource>
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<mods:title id="AEBF709E726B517913910CDED663ECA9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="92E929A85EA138A149BD0CD08F9EDA33">
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<mods:date id="BB3268A50C3407AF526DE79BFA0F2702">2009</mods:date>
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<mods:detail id="BA48780262E9EE81ED553992BB1C3C05" type="volume">
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<mods:number id="20A8E8F664F4AC13030D9E01A946DB11">2293</mods:number>
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<mods:start id="F372597C84AF58745C6CF64E497FB41F">1</mods:start>
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<mods:end id="9CB284C8E8F2ADDD7FE8DC1AFC176F2B">34</mods:end>
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<mods:classification id="8107847198785FA22E3BCE819CCAFA4B">journal article</mods:classification>
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<mods:identifier id="F2BEC6EF03A3BA404CA4853D9052F276" type="DOI">10.5281/zenodo.191476</mods:identifier>
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<mods:identifier id="FE9F6B868C6D41171DBA82E4681AF0FB" type="GBIF-Dataset">8603d4d1-efc8-44bb-84ae-76d3d8dd8161</mods:identifier>
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<mods:identifier id="DC5EE227E0E757B6059007886C5F6C99" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="50BCF4EA69D0111D16CD46B38AF1EE08" type="Zenodo-Dep">191476</mods:identifier>
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<treatment id="03D087ECAA477749FF11B43D101D2591" ID-DOI="http://doi.org/10.5281/zenodo.6214673" ID-GBIF-Taxon="119626682" ID-Zenodo-Dep="6214673" LSID="urn:lsid:plazi:treatment:03D087ECAA477749FF11B43D101D2591" httpUri="http://treatment.plazi.org/id/03D087ECAA477749FF11B43D101D2591" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
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<subSubSection id="C3636571AA47774CFF11B43D14302714" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8BC636FAAA47774CFF11B43D14AC2769" blockId="25.[151,445,1813,1874]" box="[151,445,1813,1839]" pageId="25" pageNumber="26">
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<heading id="D08E8196AA47774CFF11B43D14AC2769" bold="true" box="[151,445,1813,1839]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="B90DEAE8AA47774CFF11B43D14AC2769" bold="true" box="[151,445,1813,1839]" pageId="25" pageNumber="26">
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<taxonomicName id="4C794D79AA47774CFF11B43D14492769" box="[151,344,1813,1839]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="typhon" status="sp. nov.">
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<emphasis id="B90DEAE8AA47774CFF11B43D14492769" bold="true" box="[151,344,1813,1839]" italics="true" pageId="25" pageNumber="26">Atractus typhon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23E5793AA47774CFED9B43D14AC2769" box="[351,445,1813,1839]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC636FAAA47774CFF11B41014302714" blockId="25.[151,445,1813,1874]" box="[151,289,1848,1874]" pageId="25" pageNumber="26">
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<figureCitation id="13422A7FAA47774CFF11B41015E62714" box="[151,247,1848,1874]" captionStart="FIGURE 17" captionStartId="26.[151,254,807,831]" captionTargetBox="[217,1360,189,765]" captionTargetId="figure@26.[204,1383,182,784]" captionTargetPageId="26" captionText="FIGURE 17. Dorsal (A) and ventral (B) views of the holotype of Atractus typhon sp. nov. (ICN 10901). SVL 293 mm, CL 79 mm." httpUri="https://zenodo.org/record/191493/files/figure.png" pageId="25" pageNumber="26">Figs. 17</figureCitation>
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,
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<figureCitation id="13422A7FAA47774CFE82B41014302714" box="[260,289,1848,1874]" captionStart="FIGURE 18" captionStartId="27.[151,254,1901,1925]" captionTargetBox="[275,1308,277,1825]" captionTargetId="figure@27.[256,1354,253,1895]" captionTargetPageId="27" captionText="FIGURE 18. Dorsal (A) and lateral (B) views of head and lateral view of body (C) of the holotype of Atractus typhon sp. nov. (ICN 10901). Scale = 10 mm." httpUri="https://zenodo.org/record/191494/files/figure.png" pageId="25" pageNumber="26">18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636571AA47774CFF11B4AD17E127AA" pageId="25" pageNumber="26" type="materials_examined">
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<paragraph id="8BC636FAAA47774CFF11B4AD17E127AA" blockId="25.[151,1436,1925,2028]" pageId="25" pageNumber="26">
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<emphasis id="B90DEAE8AA47774CFF11B4AD140327D9" bold="true" box="[151,274,1925,1951]" pageId="25" pageNumber="26">
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<typeStatus id="54C28858AA47774CFF11B4AD141F27D9" box="[151,270,1925,1951]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Adult male,
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<collectionCode id="ED68AE3FAA47774CFE2DB4AD14CF27D9" box="[427,478,1925,1951]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="25" pageNumber="26">ICN</collectionCode>
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10901, from Reserva Natural Biotopo Selva Húmeda (
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<geoCoordinate id="EE4D503DAA47774CFBE7B4AD11D627D9" box="[1121,1223,1925,1951]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="1.4166666">01º25’N</geoCoordinate>
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,
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<geoCoordinate id="EE4D503DAA47774CFB55B4AD102D27D9" box="[1235,1340,1925,1951]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="-78.28333">78º17’W</geoCoordinate>
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, ca.
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<quantity id="4C819B1FAA47774CFAF6B4AD15A32780" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="26" unit="m" value="600.0">600 m</quantity>
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), vereda Berlín, El Diviso, municipality of Barbacoas, department of Nariño,
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<collectingCountry id="F36E766AAA47774CFBE6B48411C82780" box="[1120,1241,1964,1990]" name="Colombia" pageId="25" pageNumber="26">Colombia</collectingCountry>
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, collected by B. Cépeda and J. J. Mueses-Cisneros on
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<date id="FFC7103AAA47774CFDD4B4FA17FD27AA" box="[594,748,2002,2028]" pageId="25" pageNumber="26" value="2006-07-14">14 July 2006</date>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF066672AA44774FFF11B00F14032324" httpUri="https://zenodo.org/record/191493/files/figure.png" pageId="26" pageNumber="27" targetBox="[217,1360,189,765]" targetPageId="26">
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<paragraph id="8BC636FAAA44774FFF11B00F14032324" blockId="26.[151,1436,807,866]" pageId="26" pageNumber="27">
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<emphasis id="B90DEAE8AA44774FFF11B00F14362379" bold="true" box="[151,295,807,831]" pageId="26" pageNumber="27">FIGURE 17.</emphasis>
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Dorsal (A) and ventral (B) views of the holotype of
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<taxonomicName id="4C794D79AA44774FFCE6B000111B2379" box="[864,1034,808,831]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="typhon" status="sp. nov.">
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<emphasis id="B90DEAE8AA44774FFCE6B000111B2379" box="[864,1034,808,831]" italics="true" pageId="26" pageNumber="27">Atractus typhon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23E5793AA44774FFB97B00011742379" box="[1041,1125,808,831]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
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(ICN 10901). SVL 293 mm, CL 79 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3636571AA44774FFF40B09210322511" pageId="26" pageNumber="27" type="diagnosis">
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<paragraph id="8BC636FAAA44774FFF40B09214FF24A5" blockId="26.[151,1437,954,2025]" pageId="26" pageNumber="27">
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<emphasis id="B90DEAE8AA44774FFF40B09214592392" bold="true" box="[198,328,954,980]" pageId="26" pageNumber="27">Diagnosis:</emphasis>
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<taxonomicName id="4C794D79AA44774FFEDFB093170C2392" box="[345,541,955,980]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="typhon">
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<emphasis id="B90DEAE8AA44774FFEDFB093170C2392" box="[345,541,955,980]" italics="true" pageId="26" pageNumber="27">Atractus typhon</emphasis>
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</taxonomicName>
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is distinguished from all congeners by the following combination of characters: (1) 15/15/15 smooth dorsal; (2) two postoculars; (3) long loreal; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) seven infralabials, first three contacting chinshields; (7) seven maxillary teeth; (8) three gular scale rows; (9) three preventrals; (10) 156 ventrals in the single male; (11) 58 subcaudals in male; (12) dorsal ground colour beige with broad black bands alternated in the flanks; (13) venter cream with dark brown squared blotches concentrated on lateral portion of ventrals; (14) moderate body size, with males reaching
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<quantity id="4C819B1FAA44774FFD83B78A177424FA" box="[517,613,1186,1212]" metricMagnitude="-1" metricUnit="m" metricValue="2.93" pageId="26" pageNumber="27" unit="mm" value="293.0">293 mm</quantity>
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SVL; (15) long tail (27.0% SVL); (16) hemipenis moderately bilobed, semicapitate, semicalyculate.
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</paragraph>
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<paragraph id="8BC636FAAA44774FFF40B7D810322511" blockId="26.[151,1437,954,2025]" pageId="26" pageNumber="27">
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<emphasis id="B90DEAE8AA44774FFF40B7D81467254C" bold="true" box="[198,374,1264,1290]" pageId="26" pageNumber="27">Comparisons:</emphasis>
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<taxonomicName id="4C794D79AA44774FFE07B7D81751254F" box="[385,576,1264,1289]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="typhon">
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<emphasis id="B90DEAE8AA44774FFE07B7D81751254F" box="[385,576,1264,1289]" italics="true" pageId="26" pageNumber="27">Atractus typhon</emphasis>
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</taxonomicName>
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is distinguished from all congeners, except
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<taxonomicName id="4C794D79AA44774FFBD8B7D811E8254F" box="[1118,1273,1264,1289]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="iridescens">
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<emphasis id="B90DEAE8AA44774FFBD8B7D811E8254F" box="[1118,1273,1264,1289]" italics="true" pageId="26" pageNumber="27">A. iridescens</emphasis>
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</taxonomicName>
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, by having a large lateral projection in the basal portion of hemipenis.
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<taxonomicName id="4C794D79AA44774FFCB3B63F16FE2576" box="[821,1007,1303,1328]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="typhon">
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<emphasis id="B90DEAE8AA44774FFCB3B63F16FE2576" box="[821,1007,1303,1328]" italics="true" pageId="26" pageNumber="27">Atractus typhon</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C794D79AA44774FFB0FB63F10302576" box="[1161,1313,1303,1328]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="iridescens">
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<emphasis id="B90DEAE8AA44774FFB0FB63F10302576" box="[1161,1313,1303,1328]" italics="true" pageId="26" pageNumber="27">A. iridescens</emphasis>
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</taxonomicName>
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by having 15/15/15 dorsals and 58 subcaudals in males (vs. 17/17/17 dorsals and 36–38 subcaudals in males).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3636571AA447749FF40B64C174A248F" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="8BC636FAAA44774EFF40B64C175F209E" blockId="26.[151,1437,954,2025]" lastBlockId="27.[151,1436,152,216]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<emphasis id="B90DEAE8AA44774FFF40B64C14E22538" bold="true" box="[198,499,1380,1406]" pageId="26" pageNumber="27">
|
||
Description of
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<typeStatus id="54C28858AA44774FFEF8B64C14FF2538" box="[382,494,1380,1406]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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Adult male, SVL
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||
<quantity id="4C819B1FAA44774FFD4CB64C163F2538" box="[714,814,1380,1406]" metricMagnitude="-1" metricUnit="m" metricValue="2.93" pageId="26" pageNumber="27" unit="mm" value="293.0">293 mm</quantity>
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, CL
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||
<quantity id="4C819B1FAA44774FFCE0B64C16A82538" box="[870,953,1380,1406]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="26" pageNumber="27" unit="mm" value="79.0">79 mm</quantity>
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||
(27% SVL); head length 13.0 mm (4.4% SVL); head width
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<quantity id="4C819B1FAA44774FFEF5B6A214DC25E2" box="[371,461,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="26" pageNumber="27" unit="mm" value="6.2">6.2 mm</quantity>
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||
(48% of head length); body diameter
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||
<quantity id="4C819B1FAA44774FFC17B6A216FD25E2" box="[913,1004,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="26" pageNumber="27" unit="mm" value="7.3">7.3 mm</quantity>
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||
(2.6% of SVL); interocular distance
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||
<quantity id="4C819B1FAA44774FFF11B69915EA258D" box="[151,251,1457,1483]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="27" unit="mm" value="3.9">3.9 mm</quantity>
|
||
; rostro-orbital distance
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||
<quantity id="4C819B1FAA44774FFDAAB6991780258D" box="[556,657,1457,1483]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="26" pageNumber="27" unit="mm" value="3.9">3.9 mm</quantity>
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||
; naso-rostral distance 3.0 mm; head flattened in lateral view, subtriangular in dorsal view; snout truncate in lateral view, rounded in dorsal view; cervical constriction indistinct; rostral subtriangular in frontal view,
|
||
<quantity id="4C819B1FAA44774FFD5CB6D71626265F" box="[730,823,1535,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="mm" value="1.5">1.5 mm</quantity>
|
||
wide,
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||
<quantity id="4C819B1FAA44774FFC0CB6D716F6265F" box="[906,999,1535,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="mm" value="1.2">1.2 mm</quantity>
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||
high, poorly visible in dorsal view; internasal
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||
<quantity id="4C819B1FAA44774FFE94B50D147C2679" box="[274,365,1573,1599]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="27" unit="mm" value="0.9">0.9 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFE3CB50D17042679" box="[442,533,1573,1599]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="27" unit="mm" value="0.9">0.9 mm</quantity>
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||
wide; internasal suture sinistral with respect to prefrontal suture; prefrontal
|
||
<quantity id="4C819B1FAA44774FFF11B56415E32620" box="[151,242,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="27" unit="mm" value="2.5">2.5 mm</quantity>
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long,
|
||
<quantity id="4C819B1FAA44774FFEBBB56414892620" box="[317,408,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="26" pageNumber="27" unit="mm" value="2.3">2.3 mm</quantity>
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wide; supraocular subtriangular,
|
||
<quantity id="4C819B1FAA44774FFCA2B564166E2620" box="[804,895,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="26" pageNumber="27" unit="mm" value="1.9">1.9 mm</quantity>
|
||
long, 1.0 mm wide; frontal subtriangular,
|
||
<quantity id="4C819B1FAA44774FFAF1B56415D726CA" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="26" pageNumber="27" unit="mm" value="3.1">3.1 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFE91B55A146726CA" box="[279,374,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="26" pageNumber="27" unit="mm" value="2.7">2.7 mm</quantity>
|
||
wide; parietal
|
||
<quantity id="4C819B1FAA44774FFDB2B55A178226CA" box="[564,659,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="26" pageNumber="27" unit="mm" value="5.1">5.1 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFD62B55A165226CA" box="[740,835,1650,1676]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="mm" value="2.8">2.8 mm</quantity>
|
||
wide; nasal divided; nostril almost restricted to prenasal; prenasal
|
||
<quantity id="4C819B1FAA44774FFEF5B5B114DF26F5" box="[371,462,1689,1715]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="26" pageNumber="27" unit="mm" value="0.6">0.6 mm</quantity>
|
||
high,
|
||
<quantity id="4C819B1FAA44774FFD9CB5B1176426F5" box="[538,629,1689,1715]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="mm" value="0.3">0.3 mm</quantity>
|
||
long; postnasal
|
||
<quantity id="4C819B1FAA44774FFCB1B5B1168326F5" box="[823,914,1689,1715]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="mm" value="0.4">0.4 mm</quantity>
|
||
high,
|
||
<quantity id="4C819B1FAA44774FFC58B5B1112826F5" box="[990,1081,1689,1715]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long; loreal
|
||
<quantity id="4C819B1FAA44774FFB56B5B1103A26F5" box="[1232,1323,1689,1715]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="26" pageNumber="27" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFAF1B5B115D4269C" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="mm" value="0.3">0.3 mm</quantity>
|
||
high, contacting second and third supralabials; eye diameter
|
||
<quantity id="4C819B1FAA44774FFC37B5E81100269C" box="[945,1041,1728,1754]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="27" unit="mm" value="1.6">1.6 mm</quantity>
|
||
; pupil round; two postoculars of similar size; upper postocular
|
||
<quantity id="4C819B1FAA44774FFE7EB5CF17432747" box="[504,594,1767,1793]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="mm" value="0.4">0.4 mm</quantity>
|
||
high,
|
||
<quantity id="4C819B1FAA44774FFD1AB5CF17E72747" box="[668,758,1767,1793]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="mm" value="0.4">0.4 mm</quantity>
|
||
long; temporals 1+2; first temporal
|
||
<quantity id="4C819B1FAA44774FFB18B5CF11E92747" box="[1182,1272,1767,1793]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="27" unit="mm" value="1.6">1.6 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFAC4B5CF108D2747" box="[1346,1436,1767,1793]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="27" unit="mm" value="0.9">0.9 mm</quantity>
|
||
high; upper posterior temporal elongate,
|
||
<quantity id="4C819B1FAA44774FFD0BB42517FA2761" box="[653,747,1805,1831]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="26" pageNumber="27" unit="mm" value="4.2">4.2 mm</quantity>
|
||
long, 1.0 mm wide; seven supralabials, third and fourth contacting orbit; second supralabial higher than first and with similar size of third; sixth (left side) and seventh (right side) higher and longer than remaining supralabials; symphisial subtriangular,
|
||
<quantity id="4C819B1FAA44774FFB14B47211FF2732" box="[1170,1262,1882,1908]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="27" unit="mm" value="1.7">1.7 mm</quantity>
|
||
wide,
|
||
<quantity id="4C819B1FAA44774FFAC6B472108D2732" box="[1344,1436,1882,1908]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="26" pageNumber="27" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long; seven infralabials, first three contacting chinshields; first pair of infralabials in contact behind symphisial, preventing symphisial/chinshields contact; chinshields
|
||
<quantity id="4C819B1FAA44774FFC47B480110C2784" box="[961,1053,1960,1986]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="26" pageNumber="27" unit="mm" value="2.9">2.9 mm</quantity>
|
||
long,
|
||
<quantity id="4C819B1FAA44774FFBEDB48011D62784" box="[1131,1223,1960,1986]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="27" unit="mm" value="0.9">0.9 mm</quantity>
|
||
wide; three gular scale rows; three preventrals; 156 ventrals; 58 subcaudals; 15/15/15 smooth dorsal scale rows; dorsals lacking apical pits, supra-anal tubercles, and keels; six dorsal scale rows at the level of second subcaudal; caudal spine long, conical, narrow, and acuminate.
|
||
</paragraph>
|
||
<caption id="DF066672AA45774EFF11B445171827E1" httpUri="https://zenodo.org/record/191494/files/figure.png" pageId="27" pageNumber="28" targetBox="[275,1308,277,1825]" targetPageId="27">
|
||
<paragraph id="8BC636FAAA45774EFF11B445171827E1" blockId="27.[151,1436,1901,1959]" pageId="27" pageNumber="28">
|
||
<emphasis id="B90DEAE8AA45774EFF11B445143427C3" bold="true" box="[151,293,1901,1925]" pageId="27" pageNumber="28">FIGURE 18.</emphasis>
|
||
Dorsal (A) and lateral (B) views of head and lateral view of body (C) of the holotype of
|
||
<taxonomicName id="4C794D79AA45774EFB48B446106627C3" box="[1230,1399,1902,1925]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="typhon" status="sp. nov.">
|
||
<emphasis id="B90DEAE8AA45774EFB48B446106627C3" box="[1230,1399,1902,1925]" italics="true" pageId="27" pageNumber="28">Atractus typhon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23E5793AA45774EFAFBB44515D427E1" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(ICN 10901). Scale = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC636FAAA427749FF40B3BF14942109" blockId="28.[151,1437,151,1495]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90DEAE8AA427749FF40B3BF149D20F7" bold="true" box="[198,396,151,177]" pageId="28" pageNumber="29">Maxillary arch:</emphasis>
|
||
Arched in dorsal view, with six prediastemal and one postdiastemal teeth; prediastemal teeth large, of similar size, curved posteriorly, angular in cross section, robust at base, and narrower at apices; first four teeth poorly spaced, space between 4–5th and 5–6th teeth moderately long; maxillary diastema long; postdiastemal teeth half size of the last prediastemal tooth; lateral process moderately developed, lacking posterior projection.
|
||
</paragraph>
|
||
<paragraph id="8BC636FAAA427749FF40B27317B322BF" blockId="28.[151,1437,151,1495]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90DEAE8AA427749FF40B27317972133" bold="true" box="[198,646,347,373]" pageId="28" pageNumber="29">
|
||
Colour in preservative of
|
||
<typeStatus id="54C28858AA427749FD94B27317932133" box="[530,642,347,373]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Dorsum of head uniformly black; background of head uniformly black to ventral margin of supralabials; supralabials black, except for cream area on the anterior portions of each scale; mental region mostly black with light spots covering anterior region of each scale; fifth infralabial mostly cream; belly anteriorly cream with squarish black blotches concentrated on lateral portion of ventrals; ventral blotches lacking median connection at midbody; posterior third of belly with ground colour creamish gray; ventral blotches more concentrated posteriorly, forming a barely variegated pattern; lower surface of tail black, with diffuse cream spots; dorsal ground colour reddish light brown with 39–40 alternating black bands (three to five scales long), decreasing in size posteriorly; opposite bands connected along vertebral line, and reaching paraventral region along all of body; 43 reddish brown interspaces (one to three scales long), increasing in size from about midbody; paraventral region cream between dorsal bands; tail dorsally most black, with light interspaces weakly defined.
|
||
</paragraph>
|
||
<paragraph id="8BC636FAAA427749FF40B02D174A248F" blockId="28.[151,1437,151,1495]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90DEAE8AA427749FF40B02D14E22359" bold="true" box="[198,499,773,799]" pageId="28" pageNumber="29">Hemipenis morphology</emphasis>
|
||
(everted organ
|
||
<emphasis id="B90DEAE8AA427749FD43B02E17C52359" box="[709,724,774,799]" italics="true" pageId="28" pageNumber="29">n</emphasis>
|
||
= 2): Hemipenis moderately bilobed, semicapitate, and semicalyculate; lobes distinct of and restricted to distal region of
|
||
<emphasis id="B90DEAE8AA427749FC46B004112B2303" box="[960,1082,812,837]" italics="true" pageId="28" pageNumber="29">capitulum</emphasis>
|
||
; lobes attenuated and barely centrifugally oriented; lobes and
|
||
<emphasis id="B90DEAE8AA427749FDC5B07B17D3232A" box="[579,706,851,876]" italics="true" pageId="28" pageNumber="29">capitulum</emphasis>
|
||
uniformly covered with small spinulate calyces; calyces transversally arranged, forming conspicuous calyculate flounce above lobular region; capitular groove indistinct on the sulcate and weakly distinct on the asulcate side of organ; barely defined
|
||
<emphasis id="B90DEAE8AA427749FB44B088102923FF" box="[1218,1336,928,953]" italics="true" pageId="28" pageNumber="29">capitulum</emphasis>
|
||
situated just above
|
||
<emphasis id="B90DEAE8AA427749FE93B0EF14E323A6" box="[277,498,967,992]" italics="true" pageId="28" pageNumber="29">sulcus spermaticus</emphasis>
|
||
bifurcation;
|
||
<emphasis id="B90DEAE8AA427749FD0CB0EF17EE23A6" box="[650,767,967,992]" italics="true" pageId="28" pageNumber="29">capitulum</emphasis>
|
||
slightly higher on the sulcate and smaller on the asulcate side than hemipenial body;
|
||
<emphasis id="B90DEAE8AA427749FE61B0C617DD2441" box="[487,716,1006,1031]" italics="true" pageId="28" pageNumber="29">sulcus spermaticus</emphasis>
|
||
bifurcates at middle of organ;
|
||
<emphasis id="B90DEAE8AA427749FBC2B0C610382441" box="[1092,1321,1006,1031]" italics="true" pageId="28" pageNumber="29">sulcus spermaticus</emphasis>
|
||
branches centrifugally oriented, running to the tips of lobes; margins of
|
||
<emphasis id="B90DEAE8AA427749FCFFB73C1149246B" box="[889,1112,1044,1069]" italics="true" pageId="28" pageNumber="29">sulcus spermaticus</emphasis>
|
||
stout and narrow, bordered with spinules along sulcus’ extension; hemipenial body subelliptical, broader than
|
||
<emphasis id="B90DEAE8AA427749FBFCB71311E02412" box="[1146,1265,1083,1108]" italics="true" pageId="28" pageNumber="29">capitulum</emphasis>
|
||
, covered with moderate hooked spines; large spines located at lateral portion of sulcate side of the hemipenis; basal naked pocket extends to distal portion of hemipenial body; proximal region of hemipenis with longitudinal plicae and a large lateral projection (
|
||
<figureCitation id="13422A7FAA427749FE73B787172D248F" box="[501,572,1199,1225]" captionStart="FIGURE 9" captionStartId="13.[151,255,1719,1743]" captionTargetBox="[188,1418,185,1689]" captionTargetId="figure@13.[163,1425,160,1709]" captionTargetPageId="13" captionText="FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B) detailed view of lateral expansion of the hemipenis of Atractus iridesnces, (C) sulcate and asulcate sides of the hemipenis of Atractus typhon sp. nov. (ICN 10901). Scale = 5 mm." httpUri="https://zenodo.org/record/191485/files/figure.png" pageId="28" pageNumber="29">Fig. 9</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3636571AA427749FF40B7FD117B25CC" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8BC636FAAA427749FF40B7FD117B25CC" blockId="28.[151,1437,151,1495]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90DEAE8AA427749FF40B7FD144624A9" bold="true" box="[198,343,1237,1263]" pageId="28" pageNumber="29">Etymology:</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="4C794D79AA427749FDDEB7FE17B924A9" box="[600,680,1238,1263]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="typhon">
|
||
<emphasis id="B90DEAE8AA427749FDDEB7FE17B924A9" box="[600,680,1238,1263]" italics="true" pageId="28" pageNumber="29">typhon</emphasis>
|
||
</taxonomicName>
|
||
” is derived from the name of the Greek monster
|
||
<taxonomicName id="4C794D79AA427749FB64B7FD102A24A9" box="[1250,1339,1237,1263]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="34" phylum="Chordata" rank="species" species="typhon">Typhon</taxonomicName>
|
||
(Greek: Τυφoν). According to the Greek myth,
|
||
<taxonomicName id="4C794D79AA427749FDF7B7D417DD2550" box="[625,716,1276,1302]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="34" phylum="Chordata" rank="species" species="typhon">Typhon</taxonomicName>
|
||
married Echydna and fathered most mythological monsters that populated Earth.
|
||
<taxonomicName id="4C794D79AA427749FE1DB60A14E7257A" box="[411,502,1314,1340]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="34" phylum="Chordata" rank="species" species="typhon">Typhon</taxonomicName>
|
||
was described by the Greek writer Hesiod as one of the most fearsome of all creatures, “covered by a hundred serpent heads with dark flickering tongues flashing fire from their eyes”. This word is employed herein to allude to the impressive aspect of
|
||
<taxonomicName id="4C794D79AA427749FC2FB658117225CF" box="[937,1123,1392,1417]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="typhon">
|
||
<emphasis id="B90DEAE8AA427749FC2FB658117225CF" box="[937,1123,1392,1417]" italics="true" pageId="28" pageNumber="29">Atractus typhon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3636571AA427749FF40B6BE101D2591" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8BC636FAAA427749FF40B6BE101D2591" blockId="28.[151,1437,151,1495]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90DEAE8AA427749FF40B6BE147825F6" bold="true" box="[198,361,1430,1456]" pageId="28" pageNumber="29">Distribution:</emphasis>
|
||
Known only from Barbacoas in the Pacific versant of Cordillera Occidental at the department of Nariño,
|
||
<collectingCountry id="F36E766AAA427749FE24B69517082591" box="[418,537,1469,1495]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C794D79AA427749FDA2B69617CF2591" box="[548,734,1470,1495]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="typhon">
|
||
<emphasis id="B90DEAE8AA427749FDA2B69617CF2591" box="[548,734,1470,1495]" italics="true" pageId="28" pageNumber="29">Atractus typhon</emphasis>
|
||
</taxonomicName>
|
||
inhabits rainforest at
|
||
<quantity id="4C819B1FAA427749FC5BB69511392591" box="[989,1064,1469,1495]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="29" unit="m" value="600.0">600 m</quantity>
|
||
elevations (
|
||
<figureCitation id="13422A7FAA427749FB30B69511EC2591" box="[1206,1277,1469,1495]" captionStart="FIGURE 6" captionStartId="9.[151,254,1797,1821]" captionTargetBox="[153,1428,904,1773]" captionTargetId="figure@9.[151,1436,890,1812]" captionTargetPageId="9" captionText="FIGURE 6. Geographical distribution of the Atractus paucidens species group." httpUri="https://zenodo.org/record/191482/files/figure.png" pageId="28" pageNumber="29">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |