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<mods:title id="A53BF46B1AFF0E3C676FDCAF927BADB3">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB2906161BD6BDFCFEFB8AFB70" ID-DOI="http://doi.org/10.5281/zenodo.5491526" ID-GBIF-Taxon="119590170" ID-Zenodo-Dep="5491526" LSID="urn:lsid:plazi:treatment:710D87BB2906161BD6BDFCFEFB8AFB70" httpUri="http://treatment.plazi.org/id/710D87BB2906161BD6BDFCFEFB8AFB70" lastPageId="19" lastPageNumber="320" pageId="16" pageNumber="317">
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<taxonomicName id="3EA44D2E29061618D6BDFCFEFD54FCA7" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[151,645,861,887]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paraxanthusi">
<emphasis id="CBD0EABF29061618D6BDFCFEFD54FCA7" bold="true" box="[151,645,861,887]" pageId="16" pageNumber="317">
<emphasis id="CBD0EABF29061618D6BDFCFEFE3DFCA7" bold="true" box="[151,492,861,887]" italics="true" pageId="16" pageNumber="317">Ptychophallus paraxanthusi</emphasis>
(
<bibRefCitation id="9D354B5C29061618D7D1FCFEFDACFCA7" author="Bott" box="[507,637,861,887]" pageId="16" pageNumber="317" refString="Bott, R. (1968) Fluß-Krabben aus dem ostlichen Mittel-Amerika und von den Großen Antillen (Crustacea, Decapoda). Senckenbergiana biologica, 49 (1), 39 - 49." type="journal article" year="1968">Bott, 1968</bibRefCitation>
)
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(
<figureCitation id="619F2A2829061618D6B5FCDCFEF6FC48" box="[159,295,895,920]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="16" pageNumber="317">Figs. 4453</figureCitation>
,
<figureCitation id="619F2A2829061618D718FC23FE9EFC48" box="[306,335,896,920]" captionStart="FIGURE 82" captionStartId="36.[151,250,1208,1230]" captionTargetBox="[158,1431,200,1182]" captionTargetId="figure@36.[150,1436,193,1187]" captionTargetPageId="36" captionText="FIGURE 82. Distribution map of Ptychophallus paraxanthusi (Bott, 1968)." httpUri="https://zenodo.org/record/235140/files/figure.png" pageId="16" pageNumber="317">82</figureCitation>
)
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<taxonomicName id="3EA44D2E29061618D6BDFC61FE13FC08" box="[151,450,962,984]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="colombiana">
<emphasis id="CBD0EABF29061618D6BDFC61FE13FC08" box="[151,450,962,984]" italics="true" pageId="16" pageNumber="317">Pseudothelphusa colombiana</emphasis>
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<treatmentCitation id="780510BC29061618D7F1FC61FD4CFC08" author="Rathbun" box="[475,669,962,984]" page="533" pageId="16" pageNumber="317" year="1898">
<bibRefCitation id="9D354B5C29061618D7F1FC61FDBCFC08" author="Rathbun" box="[475,621,962,984]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun 1898</bibRefCitation>
: 533
</treatmentCitation>
(part).—
<treatmentCitation id="780510BC29061618D4D2FC61FC7BFC08" author="Young" box="[760,938,962,984]" page="219" pageId="16" pageNumber="317" year="1900">
<bibRefCitation id="9D354B5C29061618D4D2FC61FCABFC08" author="Young" box="[760,890,962,984]" pageId="16" pageNumber="317" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
: 219
</treatmentCitation>
(part).—
<treatmentCitation id="780510BC29061618D22FFC61FB1BFC08" author="Rathbun" box="[1029,1226,962,984]" page="302" pageId="16" pageNumber="317" year="1905">
<bibRefCitation id="9D354B5C29061618D22FFC61FB4BFC08" author="Rathbun" box="[1029,1178,962,984]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
: 302
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(part).
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</paragraph>
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<taxonomicName id="3EA44D2E29061618D6BDFC42FE71FC27" ID-CoL="7YM5P" box="[151,416,993,1015]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="xantusi">
<emphasis id="CBD0EABF29061618D6BDFC42FE71FC27" box="[151,416,993,1015]" italics="true" pageId="16" pageNumber="317">Pseudothelphusa xantusi</emphasis>
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<treatmentCitation id="780510BC29061618D790FC42FD47FC27" author="Rathbun" box="[442,662,993,1015]" page="536" pageId="16" pageNumber="317" year="1898">
<bibRefCitation id="9D354B5C29061618D790FC42FD88FC27" author="Rathbun" box="[442,601,993,1015]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun 1898</bibRefCitation>
: 536
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(part).—
<treatmentCitation id="780510BC29061618D4D4FC42FC1AFC27" author="Young" box="[766,971,993,1015]" page="208" pageId="16" pageNumber="317" year="1900">
<bibRefCitation id="9D354B5C29061618D4D4FC42FC5FFC27" author="Young" box="[766,910,993,1015]" pageId="16" pageNumber="317" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
: 208
</treatmentCitation>
(part).—
<treatmentCitation id="780510BC29061618D219FC42FAC2FC27" author="Rathbun" box="[1075,1299,993,1015]" page="292" pageId="16" pageNumber="317" year="1905">
<bibRefCitation id="9D354B5C29061618D219FC42FB07FC27" author="Rathbun" box="[1075,1238,993,1015]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
: 292
</treatmentCitation>
(part), fig. 85b.—
<treatmentCitation id="780510BC29061618D723FBA3FE1EFBC6" author="Rathbun" box="[265,463,1024,1046]" page="501" pageId="16" pageNumber="317" year="1906">
<bibRefCitation id="9D354B5C29061618D723FBA3FE4EFBC6" author="Rathbun" box="[265,415,1024,1046]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1906) Catalogue des Potamonides des collections du Museum d'histoire naturelle de Paris, d'apres les revisions et determinations de Mlle. Mary Rathbun. Seconde et derniere partie. Bulletin du Museum d'histoire naturelle Paris, 12 (7), 500 - 504." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
: 501
</treatmentCitation>
(in list).
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</paragraph>
<paragraph id="F91B36AD29061618D6BDFBBDFCB9FBE4" blockId="16.[151,1437,962,1322]" box="[151,872,1054,1077]" pageId="16" pageNumber="317">
<treatmentCitationGroup id="D9B4118329061618D6BDFBBDFCB9FBE4" box="[151,872,1054,1077]" pageId="16" pageNumber="317">
<taxonomicName id="3EA44D2E29061618D6BDFBBDFDEAFBE5" ID-CoL="7YM5P" box="[151,571,1054,1077]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="xantusi" subGenus="Ptychophallus">
<emphasis id="CBD0EABF29061618D6BDFBBDFE95FBE4" box="[151,324,1054,1076]" italics="true" pageId="16" pageNumber="317">Pseudothelphusa</emphasis>
(
<emphasis id="CBD0EABF29061618D779FBBDFE35FBE4" box="[339,484,1054,1076]" italics="true" pageId="16" pageNumber="317">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF29061618D7D8FB83FDEAFBE5" box="[498,571,1056,1077]" italics="true" pageId="16" pageNumber="317">xantusi</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC29061618D47FFBBDFCC4FBE4" author="Smalley" box="[597,789,1054,1076]" page="11" pageId="16" pageNumber="317" year="1964">
<bibRefCitation id="9D354B5C29061618D47FFBBDFD21FBE4" author="Smalley" box="[597,752,1054,1076]" pageId="16" pageNumber="317" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley 1964b</bibRefCitation>
: 11
</treatmentCitation>
, fig. 15.
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</paragraph>
<paragraph id="F91B36AD29061618D6BDFB9EFC89FB83" blockId="16.[151,1437,962,1322]" box="[151,856,1085,1107]" pageId="16" pageNumber="317">
<treatmentCitationGroup id="D9B4118329061618D6BDFB9EFC89FB83" box="[151,856,1085,1107]" pageId="16" pageNumber="317">
<taxonomicName id="3EA44D2E29061618D6BDFB9EFCC3FB83" authority="Bott, 1968: 43" authorityName="Bott" authorityPageNumber="43" authorityYear="1968" box="[151,786,1085,1107]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paraxanthusi" subGenus="Ptychophallus">
<emphasis id="CBD0EABF29061618D6BDFB9EFE95FB83" box="[151,324,1085,1107]" italics="true" pageId="16" pageNumber="317">Pseudothelphusa</emphasis>
(
<emphasis id="CBD0EABF29061618D779FB9EFE35FB83" box="[339,484,1085,1107]" italics="true" pageId="16" pageNumber="317">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF29061618D7D8FB9EFDA9FB83" box="[498,632,1085,1107]" italics="true" pageId="16" pageNumber="317">paraxanthusi</emphasis>
<treatmentCitation id="780510BC29061618D454FB9DFCC3FB83" author="Bott" box="[638,786,1085,1107]" page="43" pageId="16" pageNumber="317" year="1968">
<bibRefCitation id="9D354B5C29061618D454FB9DFD3DFB83" author="Bott" box="[638,748,1085,1107]" pageId="16" pageNumber="317" refString="Bott, R. (1968) Fluß-Krabben aus dem ostlichen Mittel-Amerika und von den Großen Antillen (Crustacea, Decapoda). Senckenbergiana biologica, 49 (1), 39 - 49." type="journal article" year="1968">Bott, 1968</bibRefCitation>
: 43
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</taxonomicName>
, fig. 8.
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</paragraph>
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<taxonomicName id="3EA44D2E29061618D6BDFBFFFCC0FBA2" box="[151,785,1116,1138]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="subSpecies" species="columbianus" subGenus="Semiptychophallus" subSpecies="paraxanthusi">
<emphasis id="CBD0EABF29061618D6BDFBFFFEF9FBA2" box="[151,296,1116,1138]" italics="true" pageId="16" pageNumber="317">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF29061618D71CFBFFFE24FBA2" box="[310,501,1116,1138]" italics="true" pageId="16" pageNumber="317">Semiptychophallus</emphasis>
)
<emphasis id="CBD0EABF29061618D42EFBFFFCC0FBA2" box="[516,785,1116,1138]" italics="true" pageId="16" pageNumber="317">columbianus paraxanthusi</emphasis>
</taxonomicName>
[sic]—
<treatmentCitation id="780510BC29061618D573FBFFFBFCFBA2" author="Pretzmann" box="[857,1069,1116,1138]" page="21" pageId="16" pageNumber="317" year="1971">
<bibRefCitation id="9D354B5C29061618D573FBFFFBD6FBA2" author="Pretzmann" box="[857,1031,1116,1138]" pageId="16" pageNumber="317" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 21
</treatmentCitation>
.
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</paragraph>
<paragraph id="F91B36AD29061618D6BDFBD9FB2AFB40" blockId="16.[151,1437,962,1322]" box="[151,1275,1146,1168]" pageId="16" pageNumber="317">
<treatmentCitationGroup id="D9B4118329061618D6BDFBD9FB2AFB40" box="[151,1275,1146,1168]" pageId="16" pageNumber="317">
<taxonomicName id="3EA44D2E29061618D6BDFBD9FCC0FB40" box="[151,785,1146,1168]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="subSpecies" species="colombianus" subGenus="Semiptychophallus" subSpecies="paraxanthusi">
<emphasis id="CBD0EABF29061618D6BDFBD9FEF9FB40" box="[151,296,1146,1168]" italics="true" pageId="16" pageNumber="317">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF29061618D71CFBD9FE24FB40" box="[310,501,1146,1168]" italics="true" pageId="16" pageNumber="317">Semiptychophallus</emphasis>
)
<emphasis id="CBD0EABF29061618D42EFBD9FCC0FB40" box="[516,785,1146,1168]" italics="true" pageId="16" pageNumber="317">colombianus paraxanthusi</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC29061618D501FBD8FC29FB40" author="Pretzmann" box="[811,1016,1146,1168]" page="89" pageId="16" pageNumber="317" year="1972">
<bibRefCitation id="9D354B5C29061618D501FBD8FC02FB40" author="Pretzmann" box="[811,979,1146,1168]" pageId="16" pageNumber="317" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 89
</treatmentCitation>
, figs. 488491, 516518.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD29061618D6BDFB3AFDE5FB3C" blockId="16.[151,1437,962,1322]" pageId="16" pageNumber="317">
<treatmentCitationGroup id="D9B4118329061618D6BDFB3AFDE5FB3C" pageId="16" pageNumber="317">
<taxonomicName id="3EA44D2E29061618D6BDFB3AFE62FB7F" box="[151,435,1177,1199]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="paraxanthusi">
<emphasis id="CBD0EABF29061618D6BDFB3AFE62FB7F" box="[151,435,1177,1199]" italics="true" pageId="16" pageNumber="317">Ptychophallus paraxanthusi</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC29061618D7E4FB3AFD48FB7F" author="Rodriguez" box="[462,665,1177,1199]" page="82" pageId="16" pageNumber="317" year="1982">
<bibRefCitation id="9D354B5C29061618D7E4FB3AFDA5FB7F" author="Rodriguez" box="[462,628,1177,1199]" pageId="16" pageNumber="317" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
: 82
</treatmentCitation>
, figs. 49a, d, j, 50ad.—
<treatmentCitation id="780510BC29061618D5B8FB3AFB1EFB7F" author="Campos" box="[914,1231,1177,1199]" page="560" pageId="16" pageNumber="317" year="1999">
<bibRefCitation id="9D354B5C29061618D5B8FB3AFB71FB7F" author="Campos" box="[914,1184,1177,1199]" pageId="16" pageNumber="317" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC29061618D305FB3AFE6BFB1E" author="Rodriguez" page="424" pageId="16" pageNumber="317" year="2000">
<bibRefCitation id="9D354B5C29061618D305FB3AFE59FB1E" author="Rodriguez" pageId="16" pageNumber="317" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 424
</treatmentCitation>
.—
<treatmentCitation id="780510BC29061618D7F2FB1BFD79FB1E" author="Ng" box="[472,680,1208,1230]" page="177" pageId="16" pageNumber="317" year="2008">
<bibRefCitation id="9D354B5C29061618D7F2FB1BFDA4FB1E" author="Ng" box="[472,629,1208,1230]" pageId="16" pageNumber="317" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF29061618D42BFB1AFDE6FB1E" box="[513,567,1208,1230]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2008
</bibRefCitation>
: 177
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC29061618D530FB1BFB68FB1E" author="Villalobos" box="[794,1209,1208,1230]" page="297" pageId="16" pageNumber="317" year="2008">
<bibRefCitation id="9D354B5C29061618D530FB1BFB56FB1E" author="Villalobos" box="[794,1159,1208,1230]" pageId="16" pageNumber="317" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC29061618D306FB1BFEFCFB3C" author="Lara" page="689" pageId="16" pageNumber="317" year="2013">
<bibRefCitation id="9D354B5C29061618D306FB1BFF2CFB3C" author="Lara" pageId="16" pageNumber="317" refString="Lara, L. R., Wehrtmann, I. S., Magalhaes, C. &amp; Mantelatto, F. L. (2013) Species diversity and distribution of freshwater crabs (Decapoda: Pseudothelphusidae) inhabiting the basin of the rio Grande de Terraba, Pacific slope of Costa Rica. Latin American Journal of Aquatic Research, 41 (4), 685 - 695. http: // dx. doi. org / 10.3856 / vol 41 - issue 4 - fulltext- 5" type="journal article" year="2013">
Lara
<emphasis id="CBD0EABF29061618D34CFB1AFA4CFB1E" box="[1382,1437,1208,1230]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2013
</bibRefCitation>
: 689
</treatmentCitation>
(table), 691, fig. 3 (map).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD29061618D6BDFB56FF24FAFA" blockId="16.[151,1437,962,1322]" pageId="16" pageNumber="317">
<treatmentCitationGroup id="D9B4118329061618D6BDFB56FF24FAFA" pageId="16" pageNumber="317">
<taxonomicName id="3EA44D2E29061618D6BDFB56FDB8FADB" ID-CoL="6WN76" authority="Rodriguez, 2001: 440" authorityName="Rodriguez" authorityPageNumber="440" authorityYear="2001" box="[151,617,1269,1291]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF29061618D6BDFB56FE55FADB" box="[151,388,1269,1291]" italics="true" pageId="16" pageNumber="317">Ptychophallus osaensis</emphasis>
<treatmentCitation id="780510BC29061618D7A0FB56FDB8FADB" author="Rodriguez" box="[394,617,1269,1291]" page="440" pageId="16" pageNumber="317" year="2001">
<bibRefCitation id="9D354B5C29061618D7A0FB56FDE7FADB" author="Rodriguez" box="[394,566,1269,1291]" pageId="16" pageNumber="317" refString="Rodriguez, G. (2001) New species and records of pseudothelphusid crabs (Crustacea: Brachyura) from Central America in the Museum of Natural History of Tulane University. Proceedings of the Biological Society of Washington, 114 (2), 435 - 443." type="journal article" year="2001">Rodríguez, 2001</bibRefCitation>
: 440
</treatmentCitation>
</taxonomicName>
, fig. 3;
<treatmentCitation id="780510BC29061618D49CFB55FCABFADB" author="Ng" box="[694,890,1269,1291]" page="177" pageId="16" pageNumber="317" year="2008">
<bibRefCitation id="9D354B5C29061618D49CFB55FC9BFADB" author="Ng" box="[694,842,1269,1291]" pageId="16" pageNumber="317" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF29061618D4F1FB55FCDFFADB" box="[731,782,1269,1291]" italics="true" pageId="16" pageNumber="317">et al.</emphasis>
2008
</bibRefCitation>
: 177
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC29061618D5C2FB56FAA8FADB" author="Villalobos" box="[1000,1401,1269,1291]" page="297" pageId="16" pageNumber="317" year="2008">
<bibRefCitation id="9D354B5C29061618D5C2FB56FA9BFADB" author="Villalobos" box="[1000,1354,1269,1291]" pageId="16" pageNumber="317" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629061619D6BDFAF2FDC5FA10" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="materials_examined">
<paragraph id="F91B36AD29061618D6BDFAF2FA4DF821" blockId="16.[151,1437,1361,2034]" pageId="16" pageNumber="317">
<emphasis id="CBD0EABF29061618D6BDFAF2FEAAFABA" bold="true" box="[151,379,1361,1386]" pageId="16" pageNumber="317">Material examined</emphasis>
.
<collectingCountry id="81B3763D29061618D7A1FAF1FDE5FABA" box="[395,564,1362,1386]" name="Costa Rica" pageId="16" pageNumber="317">COSTA RICA</collectingCountry>
. Locality not specified:
<specimenCount id="EFA2FD2429061618D562FAF1FC72FABA" box="[840,931,1361,1386]" pageId="16" pageNumber="317" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429061618D59AFAF1FBC7FABA" box="[944,1046,1361,1386]" pageId="16" pageNumber="317" type="female">1 female</specimenCount>
, UCR-MZ 678, Los Reyes Creek, Iglesia Los Reyes,
<date id="8D1A106D29061618D75FFAD6FE26FA5E" box="[373,503,1397,1422]" pageId="16" pageNumber="317" value="1972-07-22">22.vii.1972</date>
, V. Juárez leg.—San José Province, Pacific drainage:
<specimenCount id="EFA2FD2429061618D247FAD5FB6FFA5E" box="[1133,1214,1397,1422]" pageId="16" pageNumber="317" type="male">1 male</specimenCount>
, UCR-MZ 646, La Palma, La Bonita Creek,
<quantity id="3E5C9B4829061618D798FA3AFDD0FA62" box="[434,513,1433,1458]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="16" pageNumber="317" unit="km" value="6.5">6.5 km</quantity>
from San Isidro,
<date id="8D1A106D29061618D4EFFA3AFCEBFA62" box="[709,826,1433,1458]" pageId="16" pageNumber="317" value="1971-12-05">5.xii.1971</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D5C0FA3AFB95FA62" box="[1002,1092,1433,1458]" pageId="16" pageNumber="317" type="male">6 males</specimenCount>
, UCR-MZ 673, Pérez Zeledón Canton, Convento River,
<quantity id="3E5C9B4829061618D79CFA1DFE2FFA06" box="[438,510,1469,1494]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="317" unit="km" value="38.0">38 km</quantity>
E San Isidro de General,
<date id="8D1A106D29061618D508FA1EFC5FFA06" box="[802,910,1469,1494]" pageId="16" pageNumber="317" value="1972-01-31">31.i.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D268FA1DFB79FA06" box="[1090,1192,1469,1494]" pageId="16" pageNumber="317" type="male">11 males</specimenCount>
,
<specimenCount id="EFA2FD2429061618D29EFA1EFAF5FA06" box="[1204,1316,1469,1494]" pageId="16" pageNumber="317" type="female">6 females</specimenCount>
, UCR-MZ 640, Pérez Zeledón Canton, Cajón River, on Panamerican Highway,
<quantity id="3E5C9B4829061618D5B9FA41FC0BFA2A" box="[915,986,1505,1530]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="317" unit="km" value="24.0">24 km</quantity>
from San Isidro del General,
<date id="8D1A106D29061618D30EFA42FA46FA2A" box="[1316,1431,1505,1530]" pageId="16" pageNumber="317" value="1971-02-28">28.ii.1971</date>
, V. Juárez, C. Villalobos &amp; R.A. Cruz leg.;
<specimenCount id="EFA2FD2429061618D4A8F9A5FD03F9CE" box="[642,722,1541,1566]" pageId="16" pageNumber="317" type="male">1 male</specimenCount>
, UCR-MZ 657, same data;
<specimenCount id="EFA2FD2429061618D23AF9A5FBBAF9CE" box="[1040,1131,1541,1566]" pageId="16" pageNumber="317" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429061618D252F9A5FB38F9CE" box="[1144,1257,1541,1566]" pageId="16" pageNumber="317" type="female">2 females</specimenCount>
, UCR-MZ 689, Pérez Zeledón Canton, San Pablo, Platanares River,
<date id="8D1A106D29061618D4C8F98AFCB5F992" box="[738,868,1577,1602]" pageId="16" pageNumber="317" value="1972-07-15">15.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D23FF989FBBFF992" box="[1045,1134,1577,1602]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
, 1 immat. male,
<specimenCount id="EFA2FD2429061618D303F989FA49F992" box="[1321,1432,1577,1602]" pageId="16" pageNumber="317" type="female">2 females</specimenCount>
, UCR-MZ 697, Las Bonitas Creek, San Rafael de Platanares,
<date id="8D1A106D29061618D57EF9EEFC07F9B6" box="[852,982,1613,1638]" pageId="16" pageNumber="317" value="1972-07-15">15.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D2A7F9EDFB39F9B6" box="[1165,1256,1613,1638]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
, UCR-MZ 672, Platanares River,
<quantity id="3E5C9B4829061618D775F9D2FE4BF959" box="[351,410,1649,1674]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="317" unit="m" value="75.0">75 m</quantity>
N from bridge, Paso de Pejivalle [= Pejibaye] to Achotales,
<date id="8D1A106D29061618D27EF9D2FB07F95A" box="[1108,1238,1649,1674]" pageId="16" pageNumber="317" value="1972-07-22">22.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D3A4F9D1FF01F97E" pageId="16" pageNumber="317" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429061618D6F7F935FE92F97E" box="[221,323,1685,1710]" pageId="16" pageNumber="317" type="female">1 female</specimenCount>
, UCR-MZ 674, Ignacio
<collectingCountry id="81B3763D29061618D476F935FD44F97D" box="[604,661,1686,1709]" name="Malta" pageId="16" pageNumber="317">Mata</collectingCountry>
Creek,
<quantity id="3E5C9B4829061618D4DBF935FCFDF97E" box="[753,812,1685,1710]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="317" unit="km" value="2.0">2 km</quantity>
N from Pejivalle [= Pejibaye], direction to San Isidro,
<date id="8D1A106D29061618D6BDF91AFEC8F902" box="[151,281,1721,1746]" pageId="16" pageNumber="317" value="1972-07-22">22.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D7E1F919FDF5F902" box="[459,548,1721,1746]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429061618D41AF919FD45F902" box="[560,660,1721,1746]" pageId="16" pageNumber="317" type="female">1 female</specimenCount>
, UCR-MZ 692, Pérez Zeledón Canton, creek
<quantity id="3E5C9B4829061618D2B2F919FB0EF901" box="[1176,1247,1722,1746]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="317" unit="m" value="100.0">100 m</quantity>
N from plaza de Pejivalle [= Pejibaye],
<date id="8D1A106D29061618D7B2F97EFDCBF926" box="[408,538,1757,1782]" pageId="16" pageNumber="317" value="1972-07-22">22.vii.1972</date>
, V. Juárez leg.—
<taxonomicName id="3EA44D2E29061618D4FFF97EFC82F926" box="[725,851,1757,1782]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
Province, Pacific drainage:
<specimenCount id="EFA2FD2429061618D2A5F97DFB38F926" box="[1167,1257,1757,1782]" pageId="16" pageNumber="317" type="male">4 males</specimenCount>
, 1 immat. male, 1 immat. female, UCR-M 694, Ceibo River,
<quantity id="3E5C9B4829061618D4A3F8A2FD01F8CA" box="[649,720,1793,1818]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="317" unit="km" value="35.0">35 km</quantity>
E San Isidro,
<date id="8D1A106D29061618D547F8A2FC0DF8CA" box="[877,988,1793,1818]" pageId="16" pageNumber="317" value="1972-02-19">19.ii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D2A1F8A2FB35F8CA" box="[1163,1252,1793,1818]" pageId="16" pageNumber="317" type="male">6 males</specimenCount>
,
<specimenCount id="EFA2FD2429061618D2C5F8A1FA82F8CA" box="[1263,1363,1793,1818]" pageId="16" pageNumber="317" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829061618D377F8A1FA4DF8CA" box="[1373,1436,1794,1818]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="16" pageNumber="317">INPA</collectionCode>
1854, Cola de Gallo River, tributary of Cabagra River,
<geoCoordinate id="9C90506A29061618D4D4F885FC7BF8EE" box="[766,938,1830,1854]" direction="north" orientation="latitude" pageId="16" pageNumber="317" precision="1" value="9.02493">9°0129.748”N</geoCoordinate>
<geoCoordinate id="9C90506A29061618D59BF886FBA0F8EE" box="[945,1137,1829,1854]" direction="west" orientation="longitude" pageId="16" pageNumber="317" precision="1" value="-83.1738">83°1025.680”W</geoCoordinate>
,
<date id="8D1A106D29061618D256F885FB3CF8EE" box="[1148,1261,1830,1854]" pageId="16" pageNumber="317" value="2007-05-17">17.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429061618D6BDF8E9FF23F8B2" box="[151,242,1865,1890]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
, UCR-MZ 676, Sabanilla River, Paso de Sabanillas,
<date id="8D1A106D29061618D57CF8EAFC03F8B2" box="[854,978,1865,1890]" pageId="16" pageNumber="317" value="1972-03-12">12.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D2A6F8E9FB36F8B2" box="[1164,1255,1865,1890]" pageId="16" pageNumber="317" type="male">3 males</specimenCount>
, UCR-MZ 677, Escuadra River,
<quantity id="3E5C9B4829061618D765F8CDFE4FF856" box="[335,414,1901,1926]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="16" pageNumber="317" unit="km" value="2.1">2.1 km</quantity>
S Paso Real,
<date id="8D1A106D29061618D412F8CEFD77F856" box="[568,678,1901,1926]" pageId="16" pageNumber="317" value="1972-03-05">5.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D57DF8CDFC74F856" box="[855,933,1901,1926]" pageId="16" pageNumber="317" type="male">1 male</specimenCount>
, UCR-MZ 679, Curré River,
<quantity id="3E5C9B4829061618D2C4F8CEFA9BF856" box="[1262,1354,1901,1926]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="317" unit="km" value="10.5">10.5 km</quantity>
S Paso Real,
<date id="8D1A106D29061618D6F1F832FE98F87A" box="[219,329,1937,1962]" pageId="16" pageNumber="317" value="1972-03-03">3.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D42CF831FDB3F87A" box="[518,610,1937,1962]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
, UCR-MZ 691, Puerto Nuevo River,
<quantity id="3E5C9B4829061618D232F832FBB3F87A" box="[1048,1122,1937,1962]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="317" unit="km" value="16.0">16 km</quantity>
S Paso Real,
<date id="8D1A106D29061618D322F832FAA7F87A" box="[1288,1398,1937,1962]" pageId="16" pageNumber="317" value="1972-03-05">5.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D70EF815FE50F81E" box="[292,385,1973,1998]" pageId="16" pageNumber="317" type="male">2 males</specimenCount>
, UCR-MZ 675, Disciplina River,
<quantity id="3E5C9B4829061618D525F815FC88F81E" box="[783,857,1973,1998]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="317" unit="km" value="20.0">20 km</quantity>
S Paso Real,
<date id="8D1A106D29061618D22BF816FBBEF81E" box="[1025,1135,1973,1998]" pageId="16" pageNumber="317" value="1972-03-05">5.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429061618D304F815FAAEF81E" box="[1326,1407,1973,1998]" pageId="16" pageNumber="317" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429061618D3A4F815FF37F822" pageId="16" pageNumber="317" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829061618D6DFF879FEE5F822" box="[245,308,2010,2034]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="16" pageNumber="317">INPA</collectionCode>
1855, Veraguas Creek, tributary General River,
<geoCoordinate id="9C90506A29061618D558F87AFBCCF822" box="[882,1053,2009,2034]" direction="north" orientation="latitude" pageId="16" pageNumber="317" precision="1" value="9.057222">09°0326.00&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29061618D202F879FB08F822" box="[1064,1241,2010,2034]" direction="west" orientation="longitude" pageId="16" pageNumber="317" precision="1" value="-83.47973">83°2847.02&quot;W</geoCoordinate>
,
<date id="8D1A106D29061618D2C0F87AFA8BF822" box="[1258,1370,2009,2034]" pageId="16" pageNumber="317" value="2007-05-15">15.v.2007</date>
, L.R.
</paragraph>
<paragraph id="F91B36AD29071619D6BDFF3BFD39FAE0" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Lara leg.;
<specimenCount id="EFA2FD2429071619D726FF3BFE89FF60" box="[268,344,151,176]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
(cw 26.0, cl 16.3)
<specimenCount id="EFA2FD2429071619D418FF3BFD44FF60" box="[562,661,151,176]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
(cw 21.4, cl 13.5)
<specimenCount id="EFA2FD2429071619D545FF3BFC63FF60" box="[879,946,151,176]" pageId="17" pageNumber="318" type="generic">1 juv.</specimenCount>
,
<collectionCode id="9FB5AE6829071619D594FF3BFBDEFF60" LSID="urn:lsid:biocol.org:col:34871" box="[958,1039,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
20331, Boruca, undated, H. Pittier leg.; right G1,
<typeStatus id="261F880F29071619D76FFF1FFE7BFF05" box="[325,426,188,213]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
, MNHN-B 5328, Boruca, undated, H. Pittier leg.;
<specimenCount id="EFA2FD2429071619D5D1FF1FFB9CFF05" box="[1019,1101,188,213]" pageId="17" pageNumber="318" type="male">4 male</specimenCount>
, UCR-MZ 683, La Guácima Creek, on road to San Vito,
<date id="8D1A106D29071619D7CCFF7CFD8FFF28" box="[486,606,223,248]" pageId="17" pageNumber="318" value="1972-03-12">12.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D509FF7CFC53FF28" box="[803,898,223,248]" pageId="17" pageNumber="318" type="male">5 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D5B8FF43FBD7FF28" box="[914,1030,223,248]" pageId="17" pageNumber="318" type="female">2 females</specimenCount>
, UCR-MZ 2650-01, Cacao Creek, tributary Fresco River, Grande de Térraba River basin,
<date id="8D1A106D29071619D524FEA7FCAFFECC" box="[782,894,260,284]" pageId="17" pageNumber="318" value="2007-05-22">22.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D210FEA7FBA2FECD" box="[1082,1139,260,285]" pageId="17" pageNumber="318" type="generic">2 ad.</specimenCount>
males,
<specimenCount id="EFA2FD2429071619D2E1FEA7FADDFECD" box="[1227,1292,260,285]" pageId="17" pageNumber="318" type="generic">1 juv.</specimenCount>
male, UCR- MZ 3217-05, road to Ciudad Neily-San Vito, below bridge of Agua Buena,
<geoCoordinate id="9C90506A29071619D5CBFE84FB51FE90" box="[993,1152,295,320]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="8.712625">8°4245.45”N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D2ACFE84FAE9FE90" box="[1158,1336,295,320]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-82.90926">82°5433.33”W</geoCoordinate>
,
<quantity id="3E5C9B4829071619D369FE8BFA46FEEF" box="[1347,1431,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.117" pageId="17" pageNumber="318" unit="m" value="1117.0">1117 m</quantity>
,
<date id="8D1A106D29071619D6BDFEEFFED2FEB5" box="[151,259,332,357]" pageId="17" pageNumber="318" value="2014-01-27">27.i.2014</date>
, I.S. Wehrtmann, C. Magalhães &amp; E. Souza-Carvalho leg.;
<specimenCount id="EFA2FD2429071619D585FEEFFBD6FEB5" box="[943,1031,332,357]" pageId="17" pageNumber="318" type="male">6 males</specimenCount>
<specimenCount id="EFA2FD2429071619D23AFEEFFB50FEB5" box="[1040,1153,332,357]" pageId="17" pageNumber="318" type="female">2 females</specimenCount>
, UCR-MZ 696, Catarata River,
<quantity id="3E5C9B4829071619D6CEFED3FECFFE58" box="[228,286,367,392]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="318" unit="km" value="7.0">7 km</quantity>
S of bridge over El Brujo, Panamerican Highway,
<date id="8D1A106D29071619D542FECCFC0DFE58" box="[872,988,367,392]" pageId="17" pageNumber="318" value="1972-02-19">19.ii.1972</date>
, V. Juárez leg.;
<quantity id="3E5C9B4829071619D2B9FECCFB13FE57" box="[1171,1218,367,392]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="318" unit="m" value="5.0">5 m</quantity>
, UCR-MZ 681, La Vieja River,
<quantity id="3E5C9B4829071619D70FFE37FEA2FE7D" box="[293,371,404,429]" metricMagnitude="3" metricUnit="m" metricValue="7.8" pageId="17" pageNumber="318" unit="km" value="7.8">7.8 km</quantity>
S from Paso Real,
<date id="8D1A106D29071619D466FE37FD68FE7D" box="[588,697,404,429]" pageId="17" pageNumber="318" value="1972-03-05">5.iii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D541FE37FC6BFE7D" box="[875,954,404,429]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
, UCR-MZ 2660-01, Catarata River,
<quantity id="3E5C9B4829071619D37FFE37FA4DFE7D" box="[1365,1436,404,429]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="318" unit="km" value="10.0">10 km</quantity>
NW intersection of Interamericana highway, road to San Vito,
<date id="8D1A106D29071619D549FE14FC0CFE00" box="[867,989,439,464]" pageId="17" pageNumber="318" value="1986-06-14">14.vi.1986</date>
, Smalley, Campo, Kersting &amp; Kersting leg.;
<specimenCount id="EFA2FD2429071619D6F6FE7FFEEEFE25" box="[220,319,476,501]" pageId="17" pageNumber="318" type="male">5 males</specimenCount>
immat.,
<specimenCount id="EFA2FD2429071619D794FE7FFD95FE25" box="[446,580,476,501]" pageId="17" pageNumber="318" type="female">11 females</specimenCount>
immat.,
<specimenCount id="EFA2FD2429071619D4E8FE7FFCC0FE25" box="[706,785,476,501]" pageId="17" pageNumber="318" type="generic">2 juv.</specimenCount>
,
<collectionCode id="9FB5AE6829071619D503FE7FFCB9FE24" box="[809,872,476,500]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1856, Limón River (arriba),
<geoCoordinate id="9C90506A29071619D2DAFE7FFA4DFE25" box="[1264,1436,476,501]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="8.82409">8°4926.724”N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D6BDFE5CFE86FDC8" box="[151,343,511,536]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-83.028206">83°0141.520”W</geoCoordinate>
,
<date id="8D1A106D29071619D749FDA3FE14FDC8" box="[355,453,512,536]" pageId="17" pageNumber="318" value="2007-05-08">8.v.2007</date>
, R.L. Lara leg.;
<specimenCount id="EFA2FD2429071619D457FDA3FD1AFDC8" box="[637,715,511,536]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
, UCR-MZ 2652-01, Batambal Creek (Desfogue), Palmar Norte,
<date id="8D1A106D29071619D6BDFD87FED2FDED" box="[151,259,548,573]" pageId="17" pageNumber="318" value="2007-09-08">8.ix.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D7FDFD87FDFCFDED" box="[471,557,548,573]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429071619D415FD87FD7AFDED" box="[575,683,548,573]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
, UCR-MZ 2653-01, Escuadra Creek, Cataratas, Buenos Aires,
<date id="8D1A106D29071619D6BDFDE4FED6FDB0" box="[151,263,583,608]" pageId="17" pageNumber="318" value="2007-05-15">15.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D797FDEBFDC7FDB0" box="[445,534,583,608]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D40BFDEBFD54FDB0" box="[545,645,583,608]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
, UCR-MZ,2658-01, El Silencio Creek, Palmar Norte,
<date id="8D1A106D29071619D2CFFDE4FA8EFDB0" box="[1253,1375,583,608]" pageId="17" pageNumber="318" value="2007-06-27">27.vi.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D721FDCFFEB4FD55" box="[267,357,620,645]" pageId="17" pageNumber="318" type="male">3 males</specimenCount>
, UCR-MZ 687, Honda Creek, Palmar Norte,
<date id="8D1A106D29071619D542FDCFFC0DFD55" box="[872,988,620,645]" pageId="17" pageNumber="318" value="1972-07-07">7.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D2BBFDCFFB3BFD55" box="[1169,1258,620,645]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
, UCR-MZ 682, Zapote River, Palmar Norte,
<date id="8D1A106D29071619D7F1FD2CFD9EFD78" box="[475,591,655,680]" pageId="17" pageNumber="318" value="1972-07-07">7.vii.1972</date>
, V. Juárez leg.; 5 right and 2 left G1,
<collectionCode id="9FB5AE6829071619D5DBFD33FBE1FD78" box="[1009,1072,656,680]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1657, Camaronal River, Grande de Térraba River basin,
<geoCoordinate id="9C90506A29071619D785FD17FD8AFD1D" box="[431,603,692,717]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="8.96445">08°5752.02&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D44EFD17FCC7FD1C" box="[612,790,692,716]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-83.4706">83°2814.14&quot;W</geoCoordinate>
,
<date id="8D1A106D29071619D50EFD17FC45FD1C" box="[804,916,692,716]" pageId="17" pageNumber="318" value="2007-05-30">30.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D27CFD17FB60FD1D" box="[1110,1201,692,717]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
, UCR-MZ 690, Osa Canton, creek
<quantity id="3E5C9B4829071619D717FD7BFE57FD3F" box="[317,390,728,752]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="318" unit="m" value="100.0">100 m</quantity>
N from Zapote River,
<date id="8D1A106D29071619D4A4FD74FCD0FD20" box="[654,769,727,752]" pageId="17" pageNumber="318" value="1972-07-07">7.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D590FD7BFBD8FD20" box="[954,1033,727,752]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429071619D23DFD7BFBADFD20" box="[1047,1148,727,752]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
, UCR-MZ 684, Zapotico Creek, Palmar Norte,
<date id="8D1A106D29071619D7BBFD5FFE2DFCC5" box="[401,508,764,789]" pageId="17" pageNumber="318" value="1972-06-07">7.vi.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D49EFD5FFCD2FCC5" box="[692,771,764,789]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
, UCR-MZ 695, Salama Viejo River,
<quantity id="3E5C9B4829071619D286FD5FFB24FCC5" box="[1196,1269,764,789]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="318" unit="km" value="30.0">30 km</quantity>
S from bridge over Grande de Térraba River, Palmar Norte,
<date id="8D1A106D29071619D482FCBCFCCDFCE8" box="[680,796,799,824]" pageId="17" pageNumber="318" value="1972-07-05">5.vii.1972</date>
, V. Juárez leg.;
<specimenCount id="EFA2FD2429071619D5FDFC83FBF6FCE8" box="[983,1063,799,824]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829071619D21FFC83FBA5FCE8" box="[1077,1140,800,824]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1655, Grande de Térraba River basin,
<date id="8D1A106D29071619D706FCE7FE5FFC8C" box="[300,398,836,860]" pageId="17" pageNumber="318" value="2007-05-09">9.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D478FCE7FD7EFC8D" box="[594,687,836,861]" pageId="17" pageNumber="318" type="male">4 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D497FCE7FCF4FC8D" box="[701,805,836,861]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829071619D519FCE7FCA3FC8C" box="[819,882,836,860]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1852, Térraba Creek, Grande de Térraba River basin,
<geoCoordinate id="9C90506A29071619D6C9FCC4FE5EFC50" box="[227,399,871,896]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="9.06582">9°0356.952”N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D7B2FCCBFD89FC50" box="[408,600,872,896]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-83.291336">83°1728.824”W</geoCoordinate>
,
<date id="8D1A106D29071619D44DFCCBFD18FC50" box="[615,713,872,896]" pageId="17" pageNumber="318" value="2007-05-09">9.v.2007</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D5A1FCCBFC0DFC50" box="[907,988,871,896]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829071619D5C0FCCBFBF8FC50" box="[1002,1065,872,896]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1833, near Sierpe, Estero Seco,
<geoCoordinate id="9C90506A29071619D6BDFC2FFEF9FC74" box="[151,296,908,932]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="8.786366">08°47.182N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D705FC2FFE16FC75" box="[303,455,908,933]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-82.927765">82°55.666W</geoCoordinate>
,
<date id="8D1A106D29071619D7F9FC2FFD9CFC75" box="[467,589,908,933]" pageId="17" pageNumber="318" value="2009-11-16">16.xi.2009</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429071619D529FC2FFC8DFC75" box="[771,860,908,933]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
,
<typeStatus id="261F880F29071619D542FC2FFC1BFC75" box="[872,970,908,933]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
(cw 47.9, cl 28.5) and
<typeStatus id="261F880F29071619D2E6FC2EFAFDFC75" box="[1228,1324,909,933]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
(cw 31.9, cl 19.9) of
<taxonomicName id="3EA44D2E29071619D73EFC0CFD31FC18" authority="Rodriguez, 2001" authorityName="Rodriguez" authorityYear="2001" box="[276,736,943,968]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF29071619D73EFC0CFDCBFC18" box="[276,538,943,968]" italics="true" pageId="17" pageNumber="318">Ptychophallus osaensis</emphasis>
<bibRefCitation id="9D354B5C29071619D408FC0CFD31FC18" author="Rodriguez" box="[546,736,943,968]" pageId="17" pageNumber="318" refString="Rodriguez, G. (2001) New species and records of pseudothelphusid crabs (Crustacea: Brachyura) from Central America in the Museum of Natural History of Tulane University. Proceedings of the Biological Society of Washington, 114 (2), 435 - 443." type="journal article" year="2001">Rodríguez, 2001</bibRefCitation>
</taxonomicName>
,
<collectionCode id="9FB5AE6829071619D4C6FC13FCEEFC17" box="[748,831,944,967]" httpUri="http://grbio.org/cool/1h73-pg9e" name="Tianjin Museum of Natural History" pageId="17" pageNumber="318">TMNH</collectionCode>
6271, Rincón de Osa,1971, leg. unknown;
<specimenCount id="EFA2FD2429071619D303FC13FA53FC18" box="[1321,1410,943,968]" pageId="17" pageNumber="318" type="male">8 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D3A4FC13FF37FC3D" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
,
<specimenCount id="EFA2FD2429071619D6D8FC77FEE2FC3D" box="[242,307,980,1005]" pageId="17" pageNumber="318" type="generic">4 juv.</specimenCount>
,
<collectionCode id="9FB5AE6829071619D715FC76FE40FC3C" box="[319,401,981,1004]" httpUri="http://grbio.org/cool/1h73-pg9e" name="Tianjin Museum of Natural History" pageId="17" pageNumber="318">TMNH</collectionCode>
6251, Rincón de Osa,
<date id="8D1A106D29071619D4BEFC77FD22FC3D" box="[660,755,980,1005]" pageId="17" pageNumber="318" value="1972-07">vii.1972</date>
, R. Zeledón leg.;
<specimenCount id="EFA2FD2429071619D596FC77FBC7FC3D" box="[956,1046,980,1005]" pageId="17" pageNumber="318" type="male">3 males</specimenCount>
, 1 immat. male, 2 immat. females, UCR-MZ 680, Rincón de Osa, creek
<quantity id="3E5C9B4829071619D46BFC5BFDADFBC0" box="[577,636,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="318" unit="km" value="2.0">2 km</quantity>
W from Barracas OTS,
<date id="8D1A106D29071619D5BEFC5BFC27FBC0" box="[916,1014,1016,1040]" pageId="17" pageNumber="318" value="1971-05-08">8.v.1971</date>
, C. Villalobos &amp; R. Barraotes leg.;
<specimenCount id="EFA2FD2429071619D3A4FC5BFF0AFBE5" pageId="17" pageNumber="318" type="male">8 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D6CCFBBFFEB3FBE5" box="[230,354,1052,1077]" pageId="17" pageNumber="318" type="female">10 females</specimenCount>
, UCR-MZ 653, Rincón de Osa, creek behind Estación OTS,
<date id="8D1A106D29071619D220FBBFFBBDFBE4" box="[1034,1132,1052,1076]" pageId="17" pageNumber="318" value="1971-05-08">8.v.1971</date>
, C. Villalobos leg.;
<specimenCount id="EFA2FD2429071619D360FBBFFA49FBE5" box="[1354,1432,1052,1077]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429071619D6BDFBE3FF07FB88" box="[151,214,1087,1112]" pageId="17" pageNumber="318" type="generic">7 juv.</specimenCount>
females, UCR-MZ 693, Rincón de Osa, creek behind Estación OTS,
<date id="8D1A106D29071619D5CCFBE3FB98FB88" box="[998,1097,1088,1112]" pageId="17" pageNumber="318" value="1971-05-07">7.v.1971</date>
, C. Villalobos leg.;
<specimenCount id="EFA2FD2429071619D303FBE3FA53FB88" box="[1321,1410,1087,1112]" pageId="17" pageNumber="318" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D3A4FBE3FF20FBAD" pageId="17" pageNumber="318" type="female">2 females</specimenCount>
, UCR-MZ 724, Rincón de Osa, creek behind Estación OTS,
<date id="8D1A106D29071619D5E6FBC7FBFEFBAC" box="[972,1071,1124,1148]" pageId="17" pageNumber="318" value="1971-05-09">9.v.1971</date>
, C. Villalobos leg.;
<specimenCount id="EFA2FD2429071619D30AFBC7FAAFFBAD" box="[1312,1406,1124,1149]" pageId="17" pageNumber="318" type="male">4 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D3A4FBC7FF37FB70" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
, UCR-MZ 724, Rincón de Osa, creek behind Estación OTS,
<date id="8D1A106D29071619D5E2FB2BFBFAFB70" box="[968,1067,1160,1184]" pageId="17" pageNumber="318" value="1971-05-09">9.v.1971</date>
, C. Villalobos leg.;
<specimenCount id="EFA2FD2429071619D335FB2BFAACFB70" box="[1311,1405,1159,1184]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D3A4FB24FF20FB15" pageId="17" pageNumber="318" type="female">6 females</specimenCount>
,
<specimenCount id="EFA2FD2429071619D6D4FB0FFE93FB15" box="[254,322,1196,1221]" pageId="17" pageNumber="318" type="generic">1 juv.</specimenCount>
, UCR-MZ 749, Rincón de Osa, Agua Buena Creek, behind Estación Experimental,
<date id="8D1A106D29071619D320FB0FFAA7FB15" box="[1290,1398,1196,1221]" pageId="17" pageNumber="318" value="1972-06-02">2.vi.1972</date>
, E. Solis &amp; G. Rojas leg.;
<specimenCount id="EFA2FD2429071619D7BAFB73FE38FB38" box="[400,489,1231,1256]" pageId="17" pageNumber="318" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D7DFFB73FD89FB38" box="[501,600,1231,1256]" pageId="17" pageNumber="318" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829071619D44EFB73FD72FB38" box="[612,675,1232,1256]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="17" pageNumber="318">INPA</collectionCode>
1853, same data;
<specimenCount id="EFA2FD2429071619D547FB73FC7DFB38" box="[877,940,1231,1256]" pageId="17" pageNumber="318" type="generic">1 juv.</specimenCount>
male, UCR-MZ 3213-01, peninsula de Osa, road to Rincón a Drake, creek after Rancho Quemado,
<geoCoordinate id="9C90506A29071619D507FB57FC1AFADD" box="[813,971,1268,1293]" direction="north" orientation="latitude" pageId="17" pageNumber="318" precision="1" value="8.690628">8°4126.26”N</geoCoordinate>
<geoCoordinate id="9C90506A29071619D5FDFB57FB58FADD" box="[983,1161,1268,1293]" direction="west" orientation="longitude" pageId="17" pageNumber="318" precision="1" value="-83.57677">83°3436.36”W</geoCoordinate>
,
<quantity id="3E5C9B4829071619D2B0FB57FB39FADC" box="[1178,1256,1268,1292]" metricMagnitude="2" metricUnit="m" metricValue="1.91" pageId="17" pageNumber="318" unit="m" value="191.0">191 m</quantity>
,
<date id="8D1A106D29071619D2D3FB57FAB4FADD" box="[1273,1381,1268,1293]" pageId="17" pageNumber="318" value="2104-01-26">26.i.2104</date>
, I.S. Wehrtmann, C. Magalhães &amp; V. Souza-Carvalho leg.
</paragraph>
<paragraph id="F91B36AD29071619D6EDFA9FFAA5FA85" blockId="17.[151,1437,151,2013]" box="[199,1396,1340,1365]" pageId="17" pageNumber="318">
Locality not specified:
<specimenCount id="EFA2FD2429071619D7E0FA9FFDC4FA85" box="[458,533,1340,1365]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
(cw 43.0, cl 24.1),
<collectionCode id="9FB5AE6829071619D4DAFA9FFC90FA84" LSID="urn:lsid:biocol.org:col:34871" box="[752,833,1340,1364]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
20035, “
<collectingCountry id="81B3763D29071619D58CFA9FFBD1FA85" box="[934,1024,1340,1365]" name="Mexico" pageId="17" pageNumber="318">Mexico</collectingCountry>
,
<quantity id="3E5C9B4829071619D226FA9FFB87FA84" box="[1036,1110,1340,1364]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="318" unit="m" value="300.0">300 m</quantity>
”, undated, leg. unknown.
</paragraph>
<paragraph id="F91B36AD29071619D6EDFAC3FDC5FA10" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
<collectingCountry id="81B3763D29071619D6EDFAC3FE93FAA8" box="[199,322,1376,1400]" name="Panama" pageId="17" pageNumber="318">PANAMA</collectingCountry>
. Chiriquí Province, Pacific drainage:
<specimenCount id="EFA2FD2429071619D4C5FAC3FCEDFAA8" box="[751,828,1375,1400]" pageId="17" pageNumber="318" type="male">1 male</specimenCount>
(cw 30.8, cl 19.0),
<collectionCode id="9FB5AE6829071619D20BFAC3FBA3FAA8" LSID="urn:lsid:biocol.org:col:34871" box="[1057,1138,1376,1400]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
1089231, Cerro Punta,
<date id="8D1A106D29071619D3AFFAC3FED4FA4D" pageId="17" pageNumber="318" value="1963-03-01" valueMax="1963-03-03" valueMin="1963-03-01">1- 3.iii.1963</date>
, E.J. Morris leg.;
<specimenCount id="EFA2FD2429071619D7FBFA27FDFAFA4D" box="[465,555,1412,1437]" pageId="17" pageNumber="318" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429071619D41DFA27FD79FA4D" box="[567,680,1412,1437]" pageId="17" pageNumber="318" type="female">5 females</specimenCount>
,
<specimenCount id="EFA2FD2429071619D49EFA27FD27FA4D" box="[692,758,1412,1437]" pageId="17" pageNumber="318" type="generic">4 juv.</specimenCount>
,
<collectionCode id="9FB5AE6829071619D528FA27FC82FA4C" LSID="urn:lsid:biocol.org:col:34871" box="[770,851,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="318">USNM</collectionCode>
1089234, vicinities La Concepción, probably Mula River,
<date id="8D1A106D29071619D6C8FA04FEE6FA10" box="[226,311,1447,1472]" pageId="17" pageNumber="318" value="1964-04">iv.1964</date>
, leg. local children.
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629071619D6EDFA6FFD05F89D" pageId="17" pageNumber="318" type="description">
<paragraph id="F91B36AD29071619D6EDFA6FFD05F89D" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
<emphasis id="CBD0EABF29071619D6EDFA6FFE35FA35" bold="true" box="[199,484,1484,1509]" pageId="17" pageNumber="318">Description of gonopod</emphasis>
. Straight in mesiocaudal view. Marginal suture on mesial side, straight; marginal process broadly rounded, not produced beyond distal border of apex. Mesial process usually broadly rounded to subretangular, transversally projected (not downturned). Lateral process lengthy, bilobed, approximately 60% of stem length (from proximal opening to caudal border of apex), with shallow depression in middle; lateral border with usually shallow median incision; proximal lobe broader, wider than distal one, as wide as or slightly wider than lateral end of apex (mesiocaudal view); distal lobe weakly to moderately developed, obliquously directed towards caudal side, usually narrower than lateral end of apex in mesiocaudal view. Distocaudal ridge rather long (nearly reaching level of median depression of lateral process), narrow, separated from distal border of lateral process by shallow depression. Apex peduncule bent approximately 5060° towards laterocephalic side; oblong, rather wide in distal view; caudal border of apex slightly convex, with notch near lateral end. Field of apical spines well developed, facing towards cephalolateral side.
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629071619D6EDF8F4FE6FF845" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="F91B36AD29071619D6EDF8F4FE6FF845" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
<emphasis id="CBD0EABF29071619D6EDF8F4FDD9F8A0" bold="true" box="[199,520,1879,1904]" pageId="17" pageNumber="318">
<typeStatus id="261F880F29071619D6EDF8F4FEE5F8A0" box="[199,308,1879,1904]" pageId="17" pageNumber="318" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F29071619D75CF8F4FE78F8A0" box="[374,425,1879,1904]" pageId="17" pageNumber="318">type</typeStatus>
locality
</emphasis>
. Male, MNHN-B 5328 (only G1 examined).
<collectingCountry id="81B3763D29071619D237F8FBFB4CF8A0" box="[1053,1181,1879,1904]" name="Costa Rica" pageId="17" pageNumber="318">Costa Rica</collectingCountry>
,
<taxonomicName id="3EA44D2E29071619D286F8FAFAF6F8A0" box="[1196,1319,1881,1904]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
Province, Boruca (Pacific drainage).
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629071619D6EDF803FD8FF80C" pageId="17" pageNumber="318" type="distribution">
<paragraph id="F91B36AD29071619D6EDF803FD8FF80C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
<emphasis id="CBD0EABF29071619D6EDF803FE88F869" bold="true" box="[199,345,1952,1977]" pageId="17" pageNumber="318">Distribution</emphasis>
. Pacific drainage of eastern
<collectingCountry id="81B3763D29071619D484F803FCFDF869" box="[686,812,1952,1977]" name="Costa Rica" pageId="17" pageNumber="318">Costa Rica</collectingCountry>
(provinces of San José and
<taxonomicName id="3EA44D2E29071619D2ABF803FB2FF868" box="[1153,1278,1952,1976]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
) and western
<collectingCountry id="81B3763D29071619D6BDF866FF21F80C" box="[151,240,1989,2012]" name="Panama" pageId="17" pageNumber="318">Panama</collectingCountry>
(province of Chiriquí) (
<figureCitation id="619F2A2829071619D7D7F867FD80F80C" box="[509,593,1988,2013]" captionStart="FIGURE 82" captionStartId="36.[151,250,1208,1230]" captionTargetBox="[158,1431,200,1182]" captionTargetId="figure@36.[150,1436,193,1187]" captionTargetPageId="36" captionText="FIGURE 82. Distribution map of Ptychophallus paraxanthusi (Bott, 1968)." httpUri="https://zenodo.org/record/235140/files/figure.png" pageId="17" pageNumber="318">Fig. 82</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="ADDB66252904161AD6BDF932FA57F8AE" httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="18" pageNumber="319" targetBox="[163,1427,199,1649]" targetPageId="18">
<paragraph id="F91B36AD2904161AD6BDF932FA57F8AE" blockId="18.[151,1436,1681,1918]" pageId="18" pageNumber="319">
<emphasis id="CBD0EABF2904161AD6BDF932FE9EF976" bold="true" box="[151,335,1681,1704]" pageId="18" pageNumber="319">FIGURES 4460</emphasis>
.
<taxonomicName id="3EA44D2E2904161AD774F932FCD6F977" authority="Bott, 1968" authorityName="Bott" authorityYear="1968" box="[350,775,1681,1703]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="paraxanthusi">
<emphasis id="CBD0EABF2904161AD774F932FDADF977" box="[350,636,1681,1703]" italics="true" pageId="18" pageNumber="319">Ptychophallus paraxanthusi</emphasis>
(Bott, 1968)
</taxonomicName>
: 44, left first gonopod, holotype, MNHN B-5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of
<taxonomicName id="3EA44D2E2904161AD213F913FF1DF934" authority="Rodriguez, 2001" authorityName="Rodriguez" authorityYear="2001" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF2904161AD213F913FAF7F916" box="[1081,1318,1712,1734]" italics="true" pageId="18" pageNumber="319">Ptychophallus osaensis</emphasis>
Rodríguez, 2001
</taxonomicName>
, TMNH 6271, caudo-mesial view; 4748, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 5152, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855.
<taxonomicName id="3EA44D2E2904161AD414F8AFFC28F8F2" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[574,1017,1804,1826]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF2904161AD414F8AFFC94F8F2" box="[574,837,1804,1826]" italics="true" pageId="18" pageNumber="319">Ptychophallus tumimanus</emphasis>
(Rathbun, 1898)
</taxonomicName>
: 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of
<taxonomicName id="3EA44D2E2904161AD321F888FD72F88F" authority="Pretzmann, 1978" authorityName="Pretzmann" authorityYear="1978" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="subSpecies" species="tumimanus" subGenus="Ptychophallus" subSpecies="ingae">
<emphasis id="CBD0EABF2904161AD321F888FA4DF891" box="[1291,1436,1835,1857]" italics="true" pageId="18" pageNumber="319">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF2904161AD6B5F8EAFEE1F88F" box="[159,304,1865,1887]" italics="true" pageId="18" pageNumber="319">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF2904161AD715F8E9FE3CF88F" box="[319,493,1866,1887]" italics="true" pageId="18" pageNumber="319">tumimanus ingae</emphasis>
Pretzmann, 1978
</taxonomicName>
, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 5960, same data, two males from INPA 1835. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="B1BE65262905161BD6EDFF34FB8AFB70" pageId="19" pageNumber="320" type="discussion">
<paragraph id="F91B36AD2905161BD6EDFF34FED1FD54" blockId="19.[151,1437,151,1184]" pageId="19" pageNumber="320">
<emphasis id="CBD0EABF2905161BD6EDFF34FEE5FF60" bold="true" box="[199,308,151,176]" pageId="19" pageNumber="320">Remarks</emphasis>
. The
<typeStatus id="261F880F2905161BD75EFF34FE30FF60" box="[372,481,151,176]" pageId="19" pageNumber="320" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD42DFF3AFD79FF60" box="[519,680,153,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="paraxantusi">
<emphasis id="CBD0EABF2905161BD42DFF3AFD79FF60" box="[519,680,153,176]" italics="true" pageId="19" pageNumber="320">P. paraxantusi</emphasis>
</taxonomicName>
and
<taxonomicName id="3EA44D2E2905161BD4F4FF3AFC88FF60" box="[734,857,153,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF2905161BD4F4FF3AFC88FF60" box="[734,857,153,176]" italics="true" pageId="19" pageNumber="320">P. osaensis</emphasis>
</taxonomicName>
were collected from localities (Boruca and Rincón, respectively) approximately
<quantity id="3E5C9B482905161BD7F6FF1FFDF4FF05" box="[476,549,188,213]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="320" unit="km" value="40.0">40 km</quantity>
apart in southern
<taxonomicName id="3EA44D2E2905161BD4D2FF1EFCA2FF04" box="[760,883,189,212]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
Province,
<collectingCountry id="81B3763D2905161BD5DAFF1FFBA1FF05" box="[1008,1136,188,213]" name="Costa Rica" pageId="19" pageNumber="320">Costa Rica</collectingCountry>
. The slight morphological differences seen in their gonopods are not consistent to treat them as separate taxa and therefore they are hereby synonymyzed. Both lobes of the lateral process are less developed in the G1 of the
<typeStatus id="261F880F2905161BD240FEA7FB1DFECD" box="[1130,1228,260,285]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD2DDFEA6FA4DFECC" box="[1271,1436,261,284]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="paraxantusi">
<emphasis id="CBD0EABF2905161BD2DDFEA6FA4DFECC" box="[1271,1436,261,284]" italics="true" pageId="19" pageNumber="320">P. paraxantusi</emphasis>
</taxonomicName>
(
<figureCitation id="619F2A282905161BD6B5FE84FEF9FE90" box="[159,296,295,320]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 44, 45</figureCitation>
) than those of the G1of the
<typeStatus id="261F880F2905161BD441FE84FD1CFE90" box="[619,717,295,320]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD4DFFE8AFCA0FE90" box="[757,881,297,320]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF2905161BD4DFFE8AFCA0FE90" box="[757,881,297,320]" italics="true" pageId="19" pageNumber="320">P. osaensis</emphasis>
</taxonomicName>
(
<figureCitation id="619F2A282905161BD5A8FE84FC04FE90" box="[898,981,295,320]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 46</figureCitation>
) and bears some minute setae along the lateral border, besides the distal lobe being somewhat irregular in the former. The lobes are slightly more prominent in the G1 of the
<typeStatus id="261F880F2905161BD772FECCFE6BFE58" box="[344,442,367,392]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD7C9FED2FDB1FE58" box="[483,608,369,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF2905161BD7C9FED2FDB1FE58" box="[483,608,369,392]" italics="true" pageId="19" pageNumber="320">P. osaensis</emphasis>
</taxonomicName>
, with distal lobe knob-like and bearing a smooth border (
<figureCitation id="619F2A282905161BD2D2FECCFA9EFE58" box="[1272,1359,367,392]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 46</figureCitation>
). Such differences, however, could be attributed to the size of the specimens (
<typeStatus id="261F880F2905161BD59EFE37FBC9FE7D" box="[948,1048,404,429]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD215FE36FB31FE7C" box="[1087,1248,405,428]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="paraxantusi">
<emphasis id="CBD0EABF2905161BD215FE36FB31FE7C" box="[1087,1248,405,428]" italics="true" pageId="19" pageNumber="320">P. paraxantusi</emphasis>
</taxonomicName>
is subadult with cw
<quantity id="3E5C9B482905161BD694FE14FEC1FE1F" box="[190,272,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="320" unit="mm" value="26.0">26 mm</quantity>
;
<typeStatus id="261F880F2905161BD731FE14FEACFE00" box="[283,381,439,464]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E2905161BD788FE1AFDCAFE00" box="[418,539,441,464]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="CBD0EABF2905161BD788FE1AFDCAFE00" box="[418,539,441,464]" italics="true" pageId="19" pageNumber="320">P. osaensis</emphasis>
</taxonomicName>
is a fully grown adult with cw
<quantity id="3E5C9B482905161BD55DFE1BFC0DFE00" box="[887,988,440,464]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" pageId="19" pageNumber="320" unit="mm" value="47.9">47.9 mm</quantity>
) as well as to intraspecific variability as intermediate conditions have been observed in the numerous specimens examined from the region. In both topotypic males in INPA 1853 the gonopods have a distal lobe that is narrow and with smooth border, but with a wide proximal lobe (
<figureCitation id="619F2A282905161BD7ACFD87FDC3FDEC" box="[390,530,548,573]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 47, 48</figureCitation>
), whereas the male collected approximately midway from the
<typeStatus id="261F880F2905161BD2F4FD86FADFFDED" box="[1246,1294,549,573]" pageId="19" pageNumber="320">type</typeStatus>
localities of both species (Camaronal River, INPA 1657) has the distal lobe bearing an irregular border and wide proximal lobe (
<figureCitation id="619F2A282905161BD6B5FDCFFF25FD54" box="[159,244,620,645]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 49</figureCitation>
).
</paragraph>
<paragraph id="F91B36AD2905161BD6EDFD2CFB8FFC8D" blockId="19.[151,1437,151,1184]" pageId="19" pageNumber="320">
Although the proximal lobe is usually larger than the distal lobe, some variability may occur in their degree of development (see
<figureCitation id="619F2A282905161BD74CFD17FE20FD1D" box="[358,497,692,717]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 4753</figureCitation>
). In some specimens the lobes can have nearly the same size (as in some males in INPA 1854, 1855;
<figureCitation id="619F2A282905161BD75AFD74FE2BFD20" box="[368,506,727,752]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 5253</figureCitation>
), and in a few specimens the distal lobe can be even slightly larger (e.g. male in INPA 1854;
<figureCitation id="619F2A282905161BD701FD5FFE53FCC5" box="[299,386,764,789]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 51</figureCitation>
). The mesial process can also be slightly variable, with rounded (e.g. TU 6271;
<figureCitation id="619F2A282905161BD313FD5FFA5EFCC5" box="[1337,1423,764,789]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 46</figureCitation>
), subtriangular (e.g. MNHN-B 5328, INPA 1657, 1855, 1854;
<figureCitation id="619F2A282905161BD568FCBCFC23FCE8" box="[834,1010,799,824]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 44, 49, 52</figureCitation>
, respectively), or rarely subretangular shape (e.g. INPA 1853;
<figureCitation id="619F2A282905161BD789FCE7FE22FC8C" box="[419,499,836,861]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Fig. 47</figureCitation>
) shape. These conditions are illustrated in
<figureCitation id="619F2A282905161BD5F8FCE7FB8BFC8D" box="[978,1114,836,861]" captionStart="FIGURES 44 60" captionStartId="18.[151,264,1681,1704]" captionTargetBox="[163,1427,199,1649]" captionTargetId="figure@18.[160,1427,193,1660]" captionTargetPageId="18" captionText="FIGURES 44 60. Ptychophallus paraxanthusi (Bott, 1968): 44, left first gonopod, holotype, MNHN B- 5328, caudo-mesial view; 45, same data, latero-cephalic view of distal part; 46, right first gonopod, holotype of Ptychophallus osaensis Rodríguez, 2001, TMNH 6271, caudo-mesial view; 47 48, outline of right first gonopod, two males from INPA 1853, caudo-mesial view; 49, same data, INPA 1657; 50, same data, INPA 1855; 51 52, outline of left first gonopod, two males from INPA 1854, caudomesial view; 53, same data, INPA 1855. Ptychophallus tumimanus (Rathbun, 1898): 54, left first gonopod, holotype, USNM 19484, latero-cephalic view of distal part; 55, same data, caudo-mesial view; 56, same data, holotype of Ptychophallus (Ptychophallus) tumimanus ingae Pretzmann, 1978, NHMW 4009, caudo-mesial view; 57, outline of left first gonopod, INPA 1832, caudo-mesial view; 58, same data, USNM 230099; 59 60, same data, two males from INPA 1835. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235132/files/figure.png" pageId="19" pageNumber="320">Figs. 4453</figureCitation>
.
</paragraph>
<paragraph id="F91B36AD2905161BD6EDFCC4FC25FBC0" blockId="19.[151,1437,151,1184]" pageId="19" pageNumber="320">
The specimens from USNM 20331 (Boruca,
<collectingCountry id="81B3763D2905161BD4C7FCCBFCA1FC50" box="[749,880,871,896]" name="Costa Rica" pageId="19" pageNumber="320">Costa Rica</collectingCountry>
) were identified by
<bibRefCitation id="9D354B5C2905161BD25AFCC4FAF3FC50" author="Rathbun" box="[1136,1314,871,896]" pageId="19" pageNumber="320" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898</bibRefCitation>
;
<bibRefCitation id="9D354B5C2905161BD31FFCC4FABFFC50" author="Rathbun" box="[1333,1390,871,896]" pageId="19" pageNumber="320" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">1905</bibRefCitation>
) as
<taxonomicName id="3EA44D2E2905161BD6BDFC2FFE7FFC74" box="[151,430,908,933]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="xantusi">
<emphasis id="CBD0EABF2905161BD6BDFC2FFE7FFC74" box="[151,430,908,933]" italics="true" pageId="19" pageNumber="320">Pseudothelphusa xantusi</emphasis>
</taxonomicName>
, a species listed as
<emphasis id="CBD0EABF2905161BD4A0FC2EFCF4FC75" box="[650,805,908,933]" italics="true" pageId="19" pageNumber="320">incertae sedis</emphasis>
by
<bibRefCitation id="9D354B5C2905161BD565FC2FFBC9FC74" author="Rodriguez" box="[847,1048,908,933]" pageId="19" pageNumber="320" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez (1982)</bibRefCitation>
.
<bibRefCitation id="9D354B5C2905161BD20EFC2FFB05FC75" author="Rathbun" box="[1060,1236,908,933]" pageId="19" pageNumber="320" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun (1905)</bibRefCitation>
stated that the lot had three males and one young male, which was sent to MNHN. The USNM specimens consist of one male, one female, and one young specimen; the male has both G1 missing, but a small label stating “app. fig.” may indicate that it was the one used by
<bibRefCitation id="9D354B5C2905161BD7EDFC54FCC4FBC0" author="Rathbun" box="[455,789,1015,1040]" pageId="19" pageNumber="320" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905" yearSuffix="b">Rathbun (1905: 292, fig. 85b)</bibRefCitation>
to illustrate the G1.
</paragraph>
<paragraph id="F91B36AD2905161BD6EDFBBFFB8AFB70" blockId="19.[151,1437,151,1184]" pageId="19" pageNumber="320">
The lot USNM 20035 with a male identified as
<taxonomicName id="3EA44D2E2905161BD4DAFBBFFBECFBE5" box="[752,1085,1052,1077]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="colombiana">
<emphasis id="CBD0EABF2905161BD4DAFBBFFBECFBE5" box="[752,1085,1052,1077]" italics="true" pageId="19" pageNumber="320">Pseudothelphusa colombiana</emphasis>
</taxonomicName>
by
<bibRefCitation id="9D354B5C2905161BD240FBBFFA89FBE5" author="Rathbun" box="[1130,1368,1052,1077]" pageId="19" pageNumber="320" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898: 533)</bibRefCitation>
has a label stating “
<collectingCountry id="81B3763D2905161BD718FB9CFE5DFB88" box="[306,396,1087,1112]" name="Mexico" pageId="19" pageNumber="320">Mexico</collectingCountry>
,
<quantity id="3E5C9B482905161BD7B3FBE3FE32FB87" box="[409,483,1088,1112]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="320" unit="m" value="300.0">300 m</quantity>
, British Museum, Acc. 31482”, and a second with “
<taxonomicName id="3EA44D2E2905161BD21EFB9CFE74FBAD" authority="Rathbun, Det. Smalley, Jun 1966" authorityName="Rathbun, Det. Smalley, Jun" authorityYear="1966" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="CBD0EABF2905161BD21EFB9CFAFFFB88" box="[1076,1326,1087,1112]" italics="true" pageId="19" pageNumber="320">Ptychophallus tristani</emphasis>
Rathbun, Det. Smalley, Jun 1966
</taxonomicName>
. Strongly suspect an incorrect label”. The specimen was examined by the senior author, who identified it as
<taxonomicName id="3EA44D2E2905161BD75FFB2AFDF4FB70" box="[373,549,1159,1184]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="paraxanthusi">
<emphasis id="CBD0EABF2905161BD75FFB2AFDF4FB70" box="[373,549,1159,1184]" italics="true" pageId="19" pageNumber="320">P. paraxanthusi</emphasis>
</taxonomicName>
; therefore the Mexican locality is highly doubtful.
</paragraph>
</subSubSection>
</treatment>
</document>