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<mods:title id="84234B0A2A2E93DE491C5CE57FDC29C7">New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)</mods:title>
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<treatment id="2F5F550FFFB4FFBFD86C65D9656F02C4" ID-DOI="http://doi.org/10.5281/zenodo.5489142" ID-GBIF-Taxon="217236042" ID-Zenodo-Dep="5489142" LSID="urn:lsid:plazi:treatment:2F5F550FFFB4FFBFD86C65D9656F02C4" httpUri="http://treatment.plazi.org/id/2F5F550FFFB4FFBFD86C65D9656F02C4" lastPageNumber="646" pageId="3" pageNumber="646">
<subSubSection id="EFECB792FFB4FFBFD86C65D964A301D7" box="[257,683,457,481]" pageId="3" pageNumber="646" type="nomenclature">
<paragraph id="A749E419FFB4FFBFD86C65D964A301D7" blockId="3.[257,683,457,481]" box="[257,683,457,481]" pageId="3" pageNumber="646">
<heading id="FC015375FFB4FFBFD86C65D964A301D7" box="[257,683,457,481]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="646" reason="5">
<taxonomicName id="60F69F9AFFB4FFBFD86C65D964A301D7" ID-CoL="7NJRM" authority="BOUSFIELD, 1977" authorityName="Bousfield" authorityYear="1977" box="[257,683,457,481]" class="Malacostraca" family="Mesogammaridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="family">
MESOGAMMARIDAE
<bibRefCitation id="C36799E8FFB4FFBFD88E65D964A301D7" author="Bousfield EL" box="[483,683,457,481]" pageId="3" pageNumber="646" pagination="282 - 316" refId="ref12929" refString="Bousfield EL. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Supplement 4: 282 - 316." type="journal article" year="1977">BOUSFIELD, 1977</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EFECB792FFB4FFBFD9CF65EA656F02C4" pageId="3" pageNumber="646" type="diagnosis">
<paragraph id="A749E419FFB4FFBFD9CF65EA648102F1" blockId="3.[162,779,505,711]" pageId="3" pageNumber="646">
<emphasis id="9582380BFFB4FFBFD9CF65EA671A0239" box="[162,274,506,527]" italics="true" pageId="3" pageNumber="646">Diagnosis</emphasis>
: Mandible: molar tritulative. Maxilla 1: outer plate with ten or 11 robust serrate setae apically. Maxilliped: outer plate reaching end of palp article 1. Pereopod 7: basis not expanded. Gills: coxal gills on pereopods 26. Uropod 1: peduncle with or without basofacial armature. Uropod 3: inner ramus present; outer ramus elongate, uni or bi-articulate.
</paragraph>
<paragraph id="A749E419FFB4FFBFD9CF66F064BB02C3" blockId="3.[162,691,735,757]" box="[162,691,735,757]" pageId="3" pageNumber="646">
<typeStatus id="784D5ABBFFB4FFBFD9CF66F066D202C3" box="[162,218,736,757]" pageId="3" pageNumber="646">Type</typeStatus>
genus:
<taxonomicName id="60F69F9AFFB4FFBFD85966F064A702C3" authority="Tzvetkova, 1965" authorityName="Tzvetkova" authorityYear="1965" box="[308,687,735,757]" class="Malacostraca" family="Mesogammaridae" genus="Mesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD85966F067E002C3" box="[308,488,736,757]" italics="true" pageId="3" pageNumber="646">Mesogammarus</emphasis>
<bibRefCitation id="C36799E8FFB4FFBFD89C66CF64A702C3" author="Tzvetkova NL" box="[497,687,735,757]" pageId="3" pageNumber="646" pagination="1631 - 1636" refId="ref13669" refString="Tzvetkova NL. 1965. A new genus of gammarids (Amphipoda, Gammaridae) from the intertidal zone of the Japan Sea. Zoologicheskie Zhurnal 44: 1631 - 1636. [In Russia]." type="journal article" year="1965">Tzvetkova, 1965</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="A749E419FFB4FFBFD9CF671D671C03ED" blockId="3.[162,778,781,1509]" pageId="3" pageNumber="646">
<emphasis id="9582380BFFB4FFBFD9CF671D670F0314" box="[162,263,781,802]" italics="true" pageId="3" pageNumber="646">Remarks</emphasis>
: The family was erected by
<bibRefCitation id="C36799E8FFB4FFBFDB22671D65020315" author="Bousfield EL" box="[591,778,781,803]" pageId="3" pageNumber="646" pagination="282 - 316" refId="ref12929" refString="Bousfield EL. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Supplement 4: 282 - 316." type="journal article" year="1977">Bousfield (1977)</bibRefCitation>
to accommodate the monotypic genus
<taxonomicName id="60F69F9AFFB4FFBFDB3B673D67600357" authority="Tzvetkova, 1965" authorityName="Tzvetkova" authorityYear="1965" class="Malacostraca" family="Mesogammaridae" genus="Mesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDB3B673D65020374" box="[598,778,813,834]" italics="true" pageId="3" pageNumber="646">Mesogammarus</emphasis>
<bibRefCitation id="C36799E8FFB4FFBFD9CF675B67600357" author="Tzvetkova NL" box="[162,360,843,865]" pageId="3" pageNumber="646" pagination="1631 - 1636" refId="ref13669" refString="Tzvetkova NL. 1965. A new genus of gammarids (Amphipoda, Gammaridae) from the intertidal zone of the Japan Sea. Zoologicheskie Zhurnal 44: 1631 - 1636. [In Russia]." type="journal article" year="1965">Tzvetkova, 1965</bibRefCitation>
</taxonomicName>
. Subsequently, another monotypic genus,
<taxonomicName id="60F69F9AFFB4FFBFD86F677A64CB0348" authority="Bousfield, 1979" authorityName="Bousfield" authorityYear="1979" box="[258,707,873,895]" class="Malacostraca" family="Mesogammaridae" genus="Paramesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD86F677A67E30349" box="[258,491,874,895]" italics="true" pageId="3" pageNumber="646">Paramesogammarus</emphasis>
<bibRefCitation id="C36799E8FFB4FFBFDB6F677964CB0348" author="Bousfield EL" box="[514,707,873,895]" pageId="3" pageNumber="646" pagination="343 - 390" refId="ref12955" refString="Bousfield EL. 1979. A revised classification and phylogeny of amphipod crustaceans. Transactions of the Royal Society of Canada, Series IV XVI: 343 - 390." type="journal article" year="1979">Bousfield, 1979</bibRefCitation>
</taxonomicName>
, was added to the family by
<bibRefCitation id="C36799E8FFB4FFBFD8D06798647403A8" author="Bousfield EL" box="[445,636,904,926]" pageId="3" pageNumber="646" pagination="343 - 390" refId="ref12955" refString="Bousfield EL. 1979. A revised classification and phylogeny of amphipod crustaceans. Transactions of the Royal Society of Canada, Series IV XVI: 343 - 390." type="journal article" year="1979">Bousfield (1979)</bibRefCitation>
.
<bibRefCitation id="C36799E8FFB4FFBFDBE76798676C038B" author="Barnard JL &amp; Barnard KH" pageId="3" pageNumber="646" refId="ref12824" refString="Barnard JL, Barnard KH. 1983. Freshwater Amphipofa of the world, I. Evolutionary patterns and II. Handbook and bibliography. Mt. Vernon, VA: Hayfield Associates." type="book" year="1983">Barnard &amp; Barnard (1983)</bibRefCitation>
emended the original familial diagnosis.
</paragraph>
<paragraph id="A749E419FFB4FFBFD9D667F467F60460" blockId="3.[162,778,781,1509]" pageId="3" pageNumber="646">
Our view of what constitutes the important diagnostics of this family is given above. Based on this new diagnosis, we assign
<taxonomicName id="60F69F9AFFB4FFBFD8C5603164330400" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[424,571,1057,1078]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD8C5603164330400" box="[424,571,1057,1078]" italics="true" pageId="3" pageNumber="646">Eoniphargus</emphasis>
</taxonomicName>
and
<taxonomicName id="60F69F9AFFB4FFBFDBE9603165010400" authority="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari, 2007" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" box="[644,777,1057,1078]" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9582380BFFB4FFBFDBE9603165010400" box="[644,777,1057,1078]" italics="true" pageId="3" pageNumber="646">Octopupilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8EB18570FFB4FFBFD9CF6051670E0460" box="[162,262,1089,1110]" pageId="3" pageNumber="646" rank="genus">gen. nov.</taxonomicNameLabel>
to
<taxonomicName id="60F69F9AFFB4FFBFD846605067F20460" authorityName="Bousfield" authorityYear="1977" box="[299,506,1088,1110]" class="Malacostraca" family="Mesogammaridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="family">Mesogammaridae</taxonomicName>
.
</paragraph>
<paragraph id="A749E419FFB4FFBFD9D6604F63AB00CE" blockId="3.[162,778,781,1509]" lastBlockId="3.[826,1443,195,585]" pageId="3" pageNumber="646">
A potential relationship of
<taxonomicName id="60F69F9AFFB4FFBFDB75604F64A30442" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[536,683,1119,1140]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDB75604F64A30442" box="[536,683,1119,1140]" italics="true" pageId="3" pageNumber="646">Eoniphargus</emphasis>
</taxonomicName>
to the monotypic genus
<taxonomicName id="60F69F9AFFB4FFBFD805606D64D304A5" authority="Straskraba, 1967" authorityName="Straskraba" authorityYear="1967" box="[360,731,1149,1172]" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD805606D641904A4" box="[360,529,1149,1170]" italics="true" pageId="3" pageNumber="646">Indoniphargus</emphasis>
Straškraba, 1967
</taxonomicName>
has been suggested by several authors (e.g. Straškraba, 1967;
<bibRefCitation id="C36799E8FFB4FFBFD98A60AB679104E6" author="Bousfield EL" box="[231,409,1211,1233]" pageId="3" pageNumber="646" pagination="282 - 316" refId="ref12929" refString="Bousfield EL. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Supplement 4: 282 - 316." type="journal article" year="1977">Bousfield, 1977</bibRefCitation>
;
<bibRefCitation id="C36799E8FFB4FFBFD8C860AB646204E7" author="Stock JH &amp; Jo YW" box="[421,618,1211,1233]" pageId="3" pageNumber="646" pagination="624 - 632" refId="ref13513" refString="Stock JH, Jo YW. 1990. The Japanese amphipod genus Eoniphargus, rediscovered in a south Korean cave. Proceedings of the Biological Society of Washington 103: 624 - 632." type="journal article" year="1990">Stock &amp; Jo, 1990</bibRefCitation>
). By contrast,
<bibRefCitation id="C36799E8FFB4FFBFD9CF60C967E604C6" author="Barnard JL &amp; Barnard KH" box="[162,494,1241,1264]" pageId="3" pageNumber="646" refId="ref12824" refString="Barnard JL, Barnard KH. 1983. Freshwater Amphipofa of the world, I. Evolutionary patterns and II. Handbook and bibliography. Mt. Vernon, VA: Hayfield Associates." type="book" year="1983">Barnard &amp; Barnard (1983)</bibRefCitation>
classified
<taxonomicName id="60F69F9AFFB4FFBFDB1A60C9650204D8" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[631,778,1241,1262]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDB1A60C9650204D8" box="[631,778,1241,1262]" italics="true" pageId="3" pageNumber="646">Eoniphargus</emphasis>
</taxonomicName>
near the gammarids, but
<taxonomicName id="60F69F9AFFB4FFBFD8AF60E86463053B" authorityName="Straskraba" authorityYear="1967" box="[450,619,1272,1293]" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD8AF60E86463053B" box="[450,619,1272,1293]" italics="true" pageId="3" pageNumber="646">Indoniphargus</emphasis>
</taxonomicName>
near the melitids. In our opinion,
<taxonomicName id="60F69F9AFFB4FFBFD8FA61076448051A" authorityName="Straskraba" authorityYear="1967" box="[407,576,1303,1324]" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFD8FA61076448051A" box="[407,576,1303,1324]" italics="true" pageId="3" pageNumber="646">Indoniphargus</emphasis>
</taxonomicName>
appear to belong to
<taxonomicName id="60F69F9AFFB4FFBFD9AD61256786057D" authorityName="Bousfield" authorityYear="1977" box="[192,398,1333,1355]" class="Malacostraca" family="Mesogammaridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="family">Mesogammaridae</taxonomicName>
. The presence or not of the coxal gill 7 is a significant high-level taxonomic character (
<bibRefCitation id="C36799E8FFB4FFBFD9C46163676D05BE" author="Bousfield EL" box="[169,357,1395,1417]" pageId="3" pageNumber="646" pagination="282 - 316" refId="ref12929" refString="Bousfield EL. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Supplement 4: 282 - 316." type="journal article" year="1977">Bousfield, 1977</bibRefCitation>
,
<bibRefCitation id="C36799E8FFB4FFBFD817616367BC05BF" author="Bousfield EL" box="[378,436,1395,1417]" pageId="3" pageNumber="646" refId="ref12985" refString="Bousfield EL. 1983. An updated phyletic classification and palaeohistory of the Amphipoda. In: Schram FR, ed. Crustacean Phylogeny. San Diego: Museum of Natural History." type="book" year="1983">1983</bibRefCitation>
). But this character is not given in
<bibRefCitation id="C36799E8FFB4FFBFD875618167DE059E" author="Chilton C" box="[280,470,1425,1448]" pageId="3" pageNumber="646" pagination="195 - 196" refId="ref13080" refString="Chilton C. 1923. A blind amphipod from a mine in Bengal. Records of the Indian Museum 25: 195 - 196." type="journal article" year="1923">Chiltons (1923)</bibRefCitation>
original description and
<bibRefCitation id="C36799E8FFB4FFBFD9CF61A0678405F0" author="Stephensen K" box="[162,396,1456,1478]" pageId="3" pageNumber="646" pagination="13 - 19" refId="ref13486" refString="Stephensen K. 1931. Neoniphargus indicus (Chilton), an Indian fresh-water amphipod. Records of the Indian Museum 33: 13 - 19." type="journal article" year="1931">Stephensens (1931)</bibRefCitation>
redescription of
<taxonomicName id="60F69F9AFFB4FFBFDB0D61A066FE05D2" baseAuthorityName="Chilton" baseAuthorityYear="1923" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="9582380BFFB4FFBFDB0D61A066FE05D2" italics="true" pageId="3" pageNumber="646">Indoniphargus indicus</emphasis>
</taxonomicName>
. The decision of whether
<taxonomicName id="60F69F9AFFB4FFBFDB0D61DF650105D2" authorityName="Straskraba" authorityYear="1967" box="[608,777,1487,1508]" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDB0D61DF650105D2" box="[608,777,1487,1508]" italics="true" pageId="3" pageNumber="646">Indoniphargus</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="60F69F9AFFB4FFBFDAA564D3629B00EF" authorityName="Bousfield" authorityYear="1977" box="[968,1171,195,217]" class="Malacostraca" family="Mesogammaridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="family">Mesogammaridae</taxonomicName>
should be postponed until the coxal gill feature of
<taxonomicName id="60F69F9AFFB4FFBFDDED64F362E300C1" baseAuthorityName="Chilton" baseAuthorityYear="1923" box="[1152,1259,226,248]" class="Malacostraca" family="Mesogammaridae" genus="Indoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="9582380BFFB4FFBFDDED64F362E300C1" box="[1152,1259,226,248]" italics="true" pageId="3" pageNumber="646">I. indicus</emphasis>
</taxonomicName>
can be clarified.
</paragraph>
<paragraph id="A749E419FFB4FFBFDA3E651165BB027F" blockId="3.[826,1443,195,585]" pageId="3" pageNumber="646">
The genera of
<taxonomicName id="60F69F9AFFB4FFBFDD60651162D00121" authorityName="Bousfield" authorityYear="1977" box="[1037,1240,257,279]" class="Malacostraca" family="Mesogammaridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="family">Mesogammaridae</taxonomicName>
may be further grouped into related complexes. Thus,
<taxonomicName id="60F69F9AFFB4FFBFDD83653063AA0103" authorityName="Tzvetkova" authorityYear="1965" box="[1262,1442,288,309]" class="Malacostraca" family="Mesogammaridae" genus="Mesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDD83653063AA0103" box="[1262,1442,288,309]" italics="true" pageId="3" pageNumber="646">Mesogammarus</emphasis>
</taxonomicName>
and
<taxonomicName id="60F69F9AFFB4FFBFDA03652F625F0162" authorityName="Bousfield" authorityYear="1979" box="[878,1111,319,340]" class="Malacostraca" family="Mesogammaridae" genus="Paramesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDA03652F625F0162" box="[878,1111,319,340]" italics="true" pageId="3" pageNumber="646">Paramesogammarus</emphasis>
</taxonomicName>
, a marine group, posess the well-developed eyes, the stout propodus of the gnathopods, the smooth palmar margin of the gnathopods and the unstalked coxal gills.
<taxonomicName id="60F69F9AFFB4FFBFDDE0658A63280199" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[1165,1312,410,431]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDDE0658A63280199" box="[1165,1312,410,431]" italics="true" pageId="3" pageNumber="646">Eoniphargus</emphasis>
</taxonomicName>
and
<taxonomicName id="60F69F9AFFB4FFBFDC35658A657601F8" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDC35658A657601F8" italics="true" pageId="3" pageNumber="646">Octopupilla</emphasis>
</taxonomicName>
, a second related group, inhabit subterranean waters and possess the reduced eyes, the feeble propodus of the gnathopods, the palmar margin with small triangular protuberances and the pedunculate coxal gills.
</paragraph>
<paragraph id="A749E419FFB4FFBFDA576672656F02C4" blockId="3.[826,1442,609,754]" pageId="3" pageNumber="646">
<emphasis id="9582380BFFB4FFBFDA57667265C00241" box="[826,968,610,631]" italics="true" pageId="3" pageNumber="646">Composition</emphasis>
: As emended above, the family now includes the following genera:
<taxonomicName id="60F69F9AFFB4FFBFDD83669162030283" authority="Tzvetkova, 1965" authorityName="Tzvetkova" authorityYear="1965" class="Malacostraca" family="Mesogammaridae" genus="Mesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDD83669163AA02A0" box="[1262,1442,641,662]" italics="true" pageId="3" pageNumber="646">Mesogammarus</emphasis>
<bibRefCitation id="C36799E8FFB4FFBFDA57668F62030283" author="Tzvetkova NL" box="[826,1035,671,693]" pageId="3" pageNumber="646" pagination="1631 - 1636" refId="ref13669" refString="Tzvetkova NL. 1965. A new genus of gammarids (Amphipoda, Gammaridae) from the intertidal zone of the Japan Sea. Zoologicheskie Zhurnal 44: 1631 - 1636. [In Russia]." type="journal article" year="1965">Tzvetkova, 1965</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="60F69F9AFFB4FFBFDD46668F657B02E5" authority="Bousfield, 1979" authorityName="Bousfield" authorityYear="1979" class="Malacostraca" family="Mesogammaridae" genus="Paramesogammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDD46668F631C0282" box="[1067,1300,671,692]" italics="true" pageId="3" pageNumber="646">Paramesogammarus</emphasis>
<bibRefCitation id="C36799E8FFB4FFBFDC5C668F657B02E5" author="Bousfield EL" pageId="3" pageNumber="646" pagination="343 - 390" refId="ref12955" refString="Bousfield EL. 1979. A revised classification and phylogeny of amphipod crustaceans. Transactions of the Royal Society of Canada, Series IV XVI: 343 - 390." type="journal article" year="1979">Bousfield, 1979</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="60F69F9AFFB4FFBFDA1266AD62A302E5" authority="(Ueno, 1955)" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[895,1195,701,723]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus">
<emphasis id="9582380BFFB4FFBFDA1266AD621A02E4" box="[895,1042,701,722]" italics="true" pageId="3" pageNumber="646">Eoniphargus</emphasis>
(
<bibRefCitation id="C36799E8FFB4FFBFDD4C66AD62AB02E5" author="Ueno M" box="[1057,1187,701,723]" pageId="3" pageNumber="646" pagination="146 - 152" refId="ref13734" refString="Ueno M. 1955. Occurrence of a freshwater gammarid (Amphipoda) of the Niphargus group in Japan. Bulletin of the Biogeographical Society of Japan 16: 146 - 152." type="journal article" year="1955">Uéno, 1955</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="60F69F9AFFB4FFBFDD8866AD636202E4" authority="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari, 2007" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" box="[1253,1386,701,722]" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="646" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9582380BFFB4FFBFDD8866AD636202E4" box="[1253,1386,701,722]" italics="true" pageId="3" pageNumber="646">Octopupilla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8EB18570FFB4FFBFDC1F66AE656F02C4" pageId="3" pageNumber="646" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>