224 lines
30 KiB
XML
224 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1188.89881" ID-Pensoft-Pub="1313-2970-1188-27" ID-Pensoft-UUID="E54C848A6ECE5CE1813ACC3F48516D8A" ID-ZooBank="0009E8965A8F4522BE77B5A25995122F" ModsDocID="1313-2970-1188-27" checkinTime="1704334804984" checkinUser="pensoft" docAuthor="Song, Zhi-Shun, O'Brien, Lois B., Malenovsky, Igor, Deckert, Juergen & Bartlett, Charles R." docDate="2024" docId="474620BB79E75B9BB9F53153619B3DD7" docLanguage="en" docName="ZooKeys 1188: 27-45" docOrigin="ZooKeys 1188" docPubDate="2024-01-03" docSource="http://dx.doi.org/10.3897/zookeys.1188.89881" docTitle="Trigava obrieni Song, Malenovsky & Deckert 2024, sp. nov." docType="treatment" docUuid="87633FF5-777E-4A66-9F5E-536B76E82C08" docUuidSource="ZooBank" docVersion="1" id="E54C848A6ECE5CE1813ACC3F48516D8A" lastPageNumber="27" masterDocId="E54C848A6ECE5CE1813ACC3F48516D8A" masterDocTitle="Revision of the Neotropical genus Trigava O'Brien, 1999 (Hemiptera, Fulgoromorpha, Dictyopharidae, Nersiini), with descriptions of two new species from Peru and Brazil" masterLastPageNumber="45" masterPageNumber="27" pageNumber="27" updateTime="1704334804984" updateUser="pensoft">
|
||
<mods:mods id="BC79A4017E09B7F089CF39C76E7EA1EC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8FA82A8840AF1F69BDC2961CE6CEF170">
|
||
<mods:title id="5B0ADAA8FBC6DC1D0825547706CAED8E">Revision of the Neotropical genus Trigava O'Brien, 1999 (Hemiptera, Fulgoromorpha, Dictyopharidae, Nersiini), with descriptions of two new species from Peru and Brazil</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="AC353B44A3B3344FBFEC947497739E87" type="personal">
|
||
<mods:role id="E585950DB923C5055138781EE7DB1156">
|
||
<mods:roleTerm id="153CBAC67CD7897D92734C9CA776B7C3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C44EB5CD673896D9C6BD92A2CD372B82">Song, Zhi-Shun</mods:namePart>
|
||
<mods:nameIdentifier id="412A59B45D56E1F0851BEE2CBD48CFF3" type="ORCID">https://orcid.org/0000-0002-5449-4646</mods:nameIdentifier>
|
||
<mods:affiliation id="63117531EF356A907809AC529558E754">Institute of Insect Resources and Biodiversity, School of Life Sciences, Chemistry & Chemical Engineering, Jiangsu Second Normal University, Nanjing, 210013 China</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="02D031B9C5899497ED7FDD3BC5BEB199" type="personal">
|
||
<mods:role id="6DEE368047444CAA7B79F77FE3FA141F">
|
||
<mods:roleTerm id="E9964EBB699BD38291959977F14F1AFE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F2E8A6558F58B004AC5B445226835DAA">O'Brien, Lois B.</mods:namePart>
|
||
<mods:affiliation id="4A652B8256AD55A2033ADBF2B3F3F220">Department of Entomology, University of Arizona, Forbes 410, PO Box 210036, Tucson, AZ 85721 - 0036, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="009D46D836DB81C2F65B4DD4CB9722E6" type="personal">
|
||
<mods:role id="AA67E6D4D953301B35C200A2B8A28D76">
|
||
<mods:roleTerm id="9B3BCEFB2C3D4B4F97759522BEC9A7FD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="847A36024676EC2ED68D1097F97BEE7D">Malenovsky, Igor</mods:namePart>
|
||
<mods:nameIdentifier id="20BD019E57730F6B016EBFE1797420AA" type="ORCID">https://orcid.org/0000-0001-8840-2263</mods:nameIdentifier>
|
||
<mods:affiliation id="A27459EE07A626B1319A01DF6D8F0B29">Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlarska 2, CZ- 611 37, Brno, Czech Republic & Department of Entomology, Moravian Museum, Zelny trh 6, CZ- 659 37, Brno, Czech Republic</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="756FE1ACE04BCF5443C5AEDF39B64FF4" type="personal">
|
||
<mods:role id="83229B808BBD4623AB280228FD4F831D">
|
||
<mods:roleTerm id="91AE683201AFE501273BC1E73093BA2C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="86101494FCF714511753E10AAE9F5DBF">Deckert, Juergen</mods:namePart>
|
||
<mods:nameIdentifier id="49F5215D6AC59F6963CCFDCEB836A64E" type="ORCID">https://orcid.org/0000-0003-4211-4463</mods:nameIdentifier>
|
||
<mods:affiliation id="D616CA55DD65488578B6EF113987AE1A">Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity, Invalidenstrasse 43, 10115, Berlin, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="6A25557A6EFBCDEE9A3909B0E35B6BB0" type="personal">
|
||
<mods:role id="4B9F5E15EAC64F5DEF17E67376943997">
|
||
<mods:roleTerm id="7D18A2DD85EB27CBFA123862F6D9C8FF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BFC6D993EAC3DD23DF0EB27B57D17E8C">Bartlett, Charles R.</mods:namePart>
|
||
<mods:nameIdentifier id="D9772DC9DC31BCA2BD40131B9E54419C" type="ORCID">https://orcid.org/0000-0001-9428-7337</mods:nameIdentifier>
|
||
<mods:affiliation id="89245BB4409117C59BB72AAB5D6CB79B">Department of Entomology and Wildlife Ecology, University of Delaware, 250 Townsend Hall, Newark, DE 19716 - 2160, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="CCCFCB3EF61BDB1BE2191A4AED12BE44" type="email">bartlett@udel.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1B6DAAB46CF6655B1B641691B6FBB8E8">text</mods:typeOfResource>
|
||
<mods:relatedItem id="FDF1CBC9A53E1A90CCF8B1A87BEFEE10" type="host">
|
||
<mods:titleInfo id="02D2F0CBD9B32C4080570F4D8266C420">
|
||
<mods:title id="5F924B3ED1F474D9BA506C79357C24C5">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B4202840915111CA7E7C13CED973B058">
|
||
<mods:date id="4C2A2A68D9BB9D222A4F2EFFA141B37F">2024</mods:date>
|
||
<mods:detail id="EAA6FBF9C44E8486E727E4C59EFE02BC" type="pubDate">
|
||
<mods:number id="A42717B1837700DCAF00DA2CBF1CF8AA">2024-01-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="8CE221783CB99F11E3DA7D453CC7D516" type="volume">
|
||
<mods:number id="07514BBCCA991FBAC5EA767AC3CDA44D">1188</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="50EE714D2125B088A90E3D1FC2F4A724" unit="page">
|
||
<mods:start id="827BDEEE022A27CDA64312FE47A1EDB4">27</mods:start>
|
||
<mods:end id="99B54300A80AF785A9FA99EC1A2977C7">45</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="4A7FDCF410BC02A82C94CDB1B840B762">
|
||
<mods:url id="45E5D16B9C825CB196656BCE394DC301">http://dx.doi.org/10.3897/zookeys.1188.89881</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5D6E7B35C63B40A24CF4B9E71EA5DE5C">journal article</mods:classification>
|
||
<mods:identifier id="69ADAC8AD57439F0AB792AE48FF041D0" type="DOI">http://dx.doi.org/10.3897/zookeys.1188.89881</mods:identifier>
|
||
<mods:identifier id="230F173D8CA28F24B4B59A028A3431CA" type="Pensoft-Pub">1313-2970-1188-27</mods:identifier>
|
||
<mods:identifier id="2C3D7DBD00177B18209EB274165388CE" type="ZooBank">0009E8965A8F4522BE77B5A25995122F</mods:identifier>
|
||
<mods:identifier id="65A5E21BF49EA71484A2A14EB1F4CFE9" type="Pensoft-UUID">E54C848A6ECE5CE1813ACC3F48516D8A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="474620BB79E75B9BB9F53153619B3DD7" LSID="urn:lsid:zoobank.org:act:87633FF5-777E-4A66-9F5E-536B76E82C08" httpUri="http://treatment.plazi.org/id/474620BB79E75B9BB9F53153619B3DD7" lastPageNumber="27" pageId="0" pageNumber="27">
|
||
<subSubSection id="99A11A495D461B0436F6946526844B36" pageId="0" pageNumber="27" type="nomenclature">
|
||
<paragraph id="E4B90E67B8BDE411F1FA98F966B960C4" pageId="0" pageNumber="27">
|
||
<taxonomicName id="492F20F3955234D7CC07FD6D7AC47098" LSID="https://zoobank.org/87633FF5-777E-4A66-9F5E-536B76E82C08" authority="Song, Malenovský & Deckert" authorityName="Song, Malenovsky & Deckert" authorityYear="2024" class="Insecta" family="Dictyopharidae" genus="Trigava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigava obrieni" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="obrieni" status="sp. nov.">
|
||
Trigava obrieni Song,
|
||
<normalizedToken id="ED5D54473C5F1D00F33FBD1296C4DBE9" originalValue="Malenovský">Malenovsky</normalizedToken>
|
||
& Deckert
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="148FD779E2EB253FCAC1C40735AC3154" pageId="0" pageNumber="27">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="252D4BBB0E9407C93CAF4F18522309A1" pageId="0" pageNumber="27" type="description">
|
||
<paragraph id="98947866D8BD7048CE4C94DA5F82552A" pageId="0" pageNumber="27">
|
||
<figureCitation id="007D87E4F74CFCD5D7D37389F6C03A10" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Trigava species, dorsal view A T. brachycephala (Melichar), lectotype, male B T. brachycephala (Melichar), male C T. obrieni sp. nov., holotype, male D T. obrieni sp. nov., paratype, female E T. peruensis sp. nov., holotype, male F T. recurva (Melichar), lectotype, male." figureDoi="10.3897/zookeys.1188.89881.figure1" httpUri="https://binary.pensoft.net/fig/961173" pageId="0" pageNumber="27">Figs 1C, D</figureCitation>
|
||
<figureCitation id="A234456D06F240BBCC83A4CB5E36E648" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Trigava species, lateral view A T. brachycephala (Melichar), lectotype, male B T. brachycephala (Melichar), male C T. obrieni sp. nov., holotype, male D T. obrieni sp. nov., paratype, female E T. peruensis sp. nov., holotype, male F T. recurva (Melichar), lectotype, male G right tegmen of T. brachycephala H right tegmen of T. recurva." figureDoi="10.3897/zookeys.1188.89881.figure2" httpUri="https://binary.pensoft.net/fig/961174" pageId="0" pageNumber="27">, 2C, D</figureCitation>
|
||
<figureCitation id="D7DAE51BE695A72280C309D59ACA011B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">, 4A-J</figureCitation>
|
||
<figureCitation id="C07FCFC381AF44C3D8E18C5889BEFB4A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Trigava obrieni sp. nov., paratype, female A terminalia, dorsal view B same, lateral view C same, ventral view D gonapophysis VIII, lateral view E gonapophysis IX, ventral view F gonapophysis IX, lateral view G gonoplac, lateral view H gonoplac, dorsal view. Abbreviations: ACL, anterior connective lamina of gonapophysis VIII; Gp I, first lobe (lateral lobe) of gonoplac; Gp II, second lobe (posterior lobe) of gonoplac; GxL, endogonocoxal lobe; GxP, endogonocoxal process; Gx VIII, gonocoxae VIII; PCL, posterior connective lamina; sx, segment X." figureDoi="10.3897/zookeys.1188.89881.figure5" httpUri="https://binary.pensoft.net/fig/961177" pageId="0" pageNumber="27">, 5A-H</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13DF6FAB80C7153F94C3657B71944245" pageId="0" pageNumber="27" type="type locality">
|
||
<paragraph id="A81AD5ADB99376C92B5112D758771944" pageId="0" pageNumber="27">Type locality.</paragraph>
|
||
<paragraph id="9398A66F02DF206F979D7BB4D431963E" pageId="0" pageNumber="27">
|
||
Brazil,
|
||
<normalizedToken id="F53D5617C0F0EA3C6299C1DEEE68DD21" originalValue="Rondônia">Rondonia</normalizedToken>
|
||
State, 62 km SW Ariquemes, Fazenda, Rancho Grande.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6CA5D5292DA2B4B9DA953B9115D7383" pageId="0" pageNumber="27" type="type material">
|
||
<paragraph id="6FD94BE08E91E1ED26E2758B13D1F60C" pageId="0" pageNumber="27">Type material.</paragraph>
|
||
<paragraph id="B876FDECB9AE4CBF78CC051CBE645FBC" pageId="0" pageNumber="27">
|
||
<emphasis id="F45AD3AD45CADD323268A69704944433" bold="true" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
|
||
♂,
|
||
<emphasis id="4EEDB5B482EF81AAA942EA4B2D6DB090" bold="true" pageId="0" pageNumber="27">Brazil</emphasis>
|
||
:
|
||
<normalizedToken id="4B06987943CCE02DD666B04F616AA940" originalValue="Rondônia">Rondonia</normalizedToken>
|
||
, 62 km, SW Ariquemes, Fzda, Rancho Grande, 19-XI-1994, C.W.
|
||
<normalizedToken id="7664572534A20C96628A8613F9926301" originalValue="O’Brien">O'Brien</normalizedToken>
|
||
& L.B.
|
||
<normalizedToken id="1A8FA4BA345B19B0C151A60B55921315" originalValue="O’Brien">O'Brien</normalizedToken>
|
||
leg. (LBOB; dry-mounted, pinned).
|
||
<emphasis id="F5DF73FBC21BA87734B3D6B13E4E9D52" bold="true" pageId="0" pageNumber="27">
|
||
<emphasis id="C5E3A346E6E6C8E27C0734DAE5F90BED" italics="true" pageId="0" pageNumber="27">Paratypes</emphasis>
|
||
: Brazil
|
||
</emphasis>
|
||
: 1♂, 1♀, same data as holotype but 18-XI-1994 (LBOB); 1♂, same data as holotype but 4-16-XI-1997, J.E. Eger leg. (LBOB).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="60990F449FCD00A5F35336747E8373FE" pageId="0" pageNumber="27" type="diagnosis">
|
||
<paragraph id="868CDBC235DEF623159064B0EB69603E" pageId="0" pageNumber="27">Diagnosis.</paragraph>
|
||
<paragraph id="0D9257A08B15121D201F3C0969857D05" pageId="0" pageNumber="27">
|
||
<taxonomicName id="D5BB8F0C9D60CBCEEF102D7388C7D35D" authorityName="Song, Malenovsky & Deckert" authorityYear="2024" class="Insecta" family="Dictyopharidae" genus="Trigava" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigava obrieni" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="obrieni">
|
||
<emphasis id="5CE51D853BE1BC8CAB0AA17C356762A3" italics="true" pageId="0" pageNumber="27">Trigava obrieni</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName id="92181DD8773749055E5B6BD1ED6B609A" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. brachycephala" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="brachycephala">
|
||
<emphasis id="09CA45225E1829FD9E8E105BDF99259D" italics="true" pageId="0" pageNumber="27">T. brachycephala</emphasis>
|
||
</taxonomicName>
|
||
in most characters, but can be separated from the latter by the longer head curved upward about 60° in front of eyes (in
|
||
<taxonomicName id="95FD4575597A6B5A83E8065F2860B763" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. brachycephala" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="brachycephala">
|
||
<emphasis id="146D94E2012DA3EE9420CB6B6EB33AFC" italics="true" pageId="0" pageNumber="27">T. brachycephala</emphasis>
|
||
</taxonomicName>
|
||
, the cephalic process is distinctly shorter and curved upward more than 90° in front of eyes) and the ventral lobes of the aedeagus without a tooth at the base (with a minute tooth at base in
|
||
<taxonomicName id="0F8B7B41DEAFFEC3325C8B96BA7F6782" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. brachycephala" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="brachycephala">
|
||
<emphasis id="C0E5522BFA94C32A76A1E4F038F05F84" italics="true" pageId="0" pageNumber="27">T. brachycephala</emphasis>
|
||
</taxonomicName>
|
||
). This new species also may be differentiated from
|
||
<taxonomicName id="1EB461068890C5858E7138101D853EE9" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. recurva" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="recurva">
|
||
<emphasis id="DD4134A3FA28E8BC46237EBDD057A167" italics="true" pageId="0" pageNumber="27">T. recurva</emphasis>
|
||
</taxonomicName>
|
||
(Melichar) by the gonostyles with the dorsal process short and the hook-like process situated submedially and curved basad (dorsal process distinctly elongate, hook-like process situated more basally and curved apicad in
|
||
<taxonomicName id="C23E7AD709DC92A28A19F00598FE60BE" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. recurva" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="recurva">
|
||
<emphasis id="584CE2C7F3BF5D1618D23667336B080D" italics="true" pageId="0" pageNumber="27">T. recurva</emphasis>
|
||
</taxonomicName>
|
||
), and the ventral lobes of the aedeagus without long spines (with long spines in
|
||
<taxonomicName id="6DC6D9E7E12AE13F989B7DC17838A960" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T. recurva" order="Hemiptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="recurva">
|
||
<emphasis id="614FC388B9B64C3C34D036833EDDE8E1" italics="true" pageId="0" pageNumber="27">T. recurva</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="90D3A4E048891EEFC3186CE1E042691A" pageId="0" pageNumber="27" type="description">
|
||
<paragraph id="C8C13C60C4B869111BBE951F2139DF77" pageId="0" pageNumber="27">Description.</paragraph>
|
||
<paragraph id="979762EE5F869DB2C5C8C21CD5DE7416" pageId="0" pageNumber="27">
|
||
<emphasis id="5FDDE6EB7402924D8F35B759900A1F91" bold="true" italics="true" pageId="0" pageNumber="27">Measurements</emphasis>
|
||
(in mm; 3♂, 1♀). Body length from apex of head to tip of tegmina: ♂ 10.8-11.2, ♀ 12.5; head length (includes: apex of cephalic process to constricted and curved part + from curved part to base of eyes): ♂♀ (0.4-0.5)+(1.0-1.1); head width including eyes: ♂ 1.4-1.5, ♀ 1.6; tegmen length: ♂ 8.5-8.9, ♀ 9.8.
|
||
</paragraph>
|
||
<paragraph id="168022465F412FC409E11C068CAD31F7" pageId="0" pageNumber="27">
|
||
<emphasis id="6823814548A6280D1E5B9949B8227DCF" bold="true" pageId="0" pageNumber="27">
|
||
<emphasis id="66D766790DE5BED67F02A90A3E571546" italics="true" pageId="0" pageNumber="27">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head stramineous green, lateral and intermediate carinae of frons in front of eyes black to blackish brown, lateral areas in front of eyes green. Pronotum and mesonotum stramineous green, upper lateral carinae of pronotum green. Tegmina and hindwings with membrane hyaline, costal margin black to dark brown, veins green to greenish yellow, pterostigmal area more or less greenish ochraceous. Legs yellowish brown, base, apex and apical spines of tibiae fuscous. Abdomen dorsally and ventrally greenish ochraceous.
|
||
</paragraph>
|
||
<paragraph id="074C5F2121AAE3AAA9084C1866BA3145" pageId="0" pageNumber="27">
|
||
<emphasis id="C72D5748DDCC97694D2437E09ED88333" bold="true" pageId="0" pageNumber="27">
|
||
<emphasis id="592D80CF24A74B60E02CA29327D59E59" italics="true" pageId="0" pageNumber="27">Structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head with cephalic process relatively long, in lateral view (Fig.
|
||
<figureCitation id="3DAE61EE5C21AB2D4E37E7C9FAAA9920" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4B</figureCitation>
|
||
), curved upward about 60° in front of eyes. Vertex (Fig.
|
||
<figureCitation id="7A6558A68D9E0F5A68DE8EB7D19AD8B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4A</figureCitation>
|
||
) broad, with ratio of length at midline to width between eyes (1.5-1.6):1. Frons (Fig.
|
||
<figureCitation id="B1D3719FCE7EE9DC5B87A015E18C00DD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4C</figureCitation>
|
||
) flat, relatively broad, with ratio of length at midline to maximum width 2.4:1.
|
||
</paragraph>
|
||
<paragraph id="7185EA5E34F5B8D92AD50748B244B0F9" pageId="0" pageNumber="27">
|
||
<emphasis id="AC6E17EA854C6F6C06A0EF79B0A30E9B" bold="true" pageId="0" pageNumber="27">
|
||
<emphasis id="FCCAB43473E0CF3D001DAA978A651D1E" italics="true" pageId="0" pageNumber="27">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Pygofer in lateral view (Fig.
|
||
<figureCitation id="775DC5E6D06FE130880DB5831F168919" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4E</figureCitation>
|
||
) with ratio of ventral to dorsal width about 1.7:1; posterior margin produced into a small, apically obtuse process near middle. Gonostyles (Fig.
|
||
<figureCitation id="90056F672F8A9681F326D9D8D622DDAC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4F</figureCitation>
|
||
) large and broad, dorsal process short, acute apically, more or less incurved and directed dorsoanteriad; hook-like process placed submedially, horizontal with dorsal process, curved basad (Fig.
|
||
<figureCitation id="CDE2F33AAF339EF925004315763A1102" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4G</figureCitation>
|
||
). Aedeagus (Fig.
|
||
<figureCitation id="D3A125074407AC356C429A0E2E81F483" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4H-J</figureCitation>
|
||
) slender and elongate, endosomal processes curved dorsoanteriad; phallobase sclerotized and pigmented at lateral sides, membranous and slightly inflated dorsally and ventrally: dorsal lobes V-shaped at apex, curved and directed posterolaterad; a pair of lateral lobes large and elongate, thumb-like, apex with a long spine, directed posteriad; ventral lobe small, butterfly-shaped, base expanded laterad, apex produced in a pair of thumb-like moderate lobes, without spine, directed laterad. Segment X (Fig.
|
||
<figureCitation id="8E994FF4947AAD1A8CDDDEFD5E345C80" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Trigava obrieni sp. nov., holotype, male A head and thorax, dorsal view B same, lateral view C same, ventral view D pygofer and segment X, dorsal view E pygofer, gonostyles, aedeagus and segment X, right lateral view F pygofer and gonostyles, ventral view G left gonostyle, lateral view H aedeagus, dorsal view I aedeagus, lateral view J aedeagus, ventral view. Abbreviations: as, anal style; dl, dorsal lobes of phallotheca; dmp, dorsal margin of pygofer; dpg, dorsal process of gonostyle; ep, endosomal processes; gs, gonostyle; hpg, hook-like process of gonostyle; ll, lateral lobes of phallotheca; pg, pygofer; pt, phallotheca; sx, segment X; vl, ventral lobes of phallotheca; vmp, ventral margin of pygofer." figureDoi="10.3897/zookeys.1188.89881.figure4" httpUri="https://binary.pensoft.net/fig/961176" pageId="0" pageNumber="27">4D, E</figureCitation>
|
||
), in dorsal view, with lateral margins more or less convex near middle, with ratio of length to width near middle about 2.0:1.
|
||
</paragraph>
|
||
<paragraph id="35D05418C80107E1EF81C78BCBB3C271" pageId="0" pageNumber="27">
|
||
<emphasis id="20195E36D8317C70EFABDBA3682EBC00" bold="true" pageId="0" pageNumber="27">
|
||
<emphasis id="5EDB2607E23323E04BF169037160E070" italics="true" pageId="0" pageNumber="27">Female genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in generic description (Fig.
|
||
<figureCitation id="793710B8AF0DF557CE65AAC0BF039D2F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Trigava obrieni sp. nov., paratype, female A terminalia, dorsal view B same, lateral view C same, ventral view D gonapophysis VIII, lateral view E gonapophysis IX, ventral view F gonapophysis IX, lateral view G gonoplac, lateral view H gonoplac, dorsal view. Abbreviations: ACL, anterior connective lamina of gonapophysis VIII; Gp I, first lobe (lateral lobe) of gonoplac; Gp II, second lobe (posterior lobe) of gonoplac; GxL, endogonocoxal lobe; GxP, endogonocoxal process; Gx VIII, gonocoxae VIII; PCL, posterior connective lamina; sx, segment X." figureDoi="10.3897/zookeys.1188.89881.figure5" httpUri="https://binary.pensoft.net/fig/961177" pageId="0" pageNumber="27">5A-H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9997EE7629A453B15D432F384E000996" pageId="0" pageNumber="27" type="etymology">
|
||
<paragraph id="FDE84B1AC17F6A8EC6319C35A7790125" pageId="0" pageNumber="27">Etymology.</paragraph>
|
||
<paragraph id="80FB735BC9A08CBBC0D56E2F9630AAC3" pageId="0" pageNumber="27">
|
||
The new species is named after the late Dr Charlie W.
|
||
<normalizedToken id="BBA9A564D804374DD3017A48D3905E25" originalValue="O’Brien">O'Brien</normalizedToken>
|
||
, former professor at Florida Agricultural and Mechanical University, USA, one of the
|
||
<normalizedToken id="4AC05D3C292601B6A388E648BBD03D30" originalValue="world’s">world's</normalizedToken>
|
||
top experts in weevils, collector of the type specimens and husband of Dr Lois B.
|
||
<normalizedToken id="D5E5AEC351F40B3C69DE51712CE181BB" originalValue="O’Brien">O'Brien</normalizedToken>
|
||
, in recognition of their kindest help and support to the first author when he visited USA in 2017. The species name is to be treated as a noun in genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1D06111BAFE591BD77C34EEB8F61ACD" pageId="0" pageNumber="27" type="distribution">
|
||
<paragraph id="83DBBA73E28226811F1E1D05E50DECFC" pageId="0" pageNumber="27">Distribution.</paragraph>
|
||
<paragraph id="016BF15F6A851A18E9F94ADEF7C18A74" pageId="0" pageNumber="27">Northwestern Brazil.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |