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<document id="BD8DCD432018EECB9C47F89EC85C909C" ID-CLB-Dataset="298742" ID-DOI="10.11646/zootaxa.5471.4.4" ID-GBIF-Dataset="cf49a9e9-837f-4e80-b5a1-1ac3ee1e6170" ID-ISSN="1175-5326" ID-Zenodo-Dep="12190387" ID-ZooBank="7486CBBE-0329-40C2-94AC-680D8703F56F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718894049516" checkinUser="plazi" docAuthor="Kim, Dooyoung &amp; Suh, Sang Jae" docDate="2024" docId="135E87AA5215FFEE2997FA9FFA724ABE" docLanguage="en" docName="zootaxa.5471.4.4.pdf" docOrigin="Zootaxa 5471 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Micronialoe (Micronialoe) Park, Kwon &amp; Lafer 1996" docType="key" docVersion="7" lastPageNumber="461" masterDocId="EF67FFD2521FFFE42900FFC0FFEE4C42" masterDocTitle="Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)" masterLastPageNumber="464" masterPageNumber="451" pageNumber="461" updateTime="1719233996494" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="24829ACDFEEBE03FC0F00FE27DAF5734">Review of the ground beetle subgenus Micronialoe from Korea, with a description of a new species (Coleoptera: Carabidae: Pterostichus)</mods:title>
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<mods:namePart id="C2CCE721188885CA56036581BDE67C38">Kim, Dooyoung</mods:namePart>
<mods:affiliation id="BB57BBC359AFED91C2759438DCBDD20D">Department of Biomedical Convergence Science and Technology, School of Industrial Technology Advances, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<mods:namePart id="A462E10D656BE9D9F4BE11C4830E6953">Suh, Sang Jae</mods:namePart>
<mods:affiliation id="E68423724A20C6E1189917BD74F3319E">Department of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea &amp; Institute of Plant Medicine, Kyungpook National University, Daegu, Republic of Korea</mods:affiliation>
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<treatment id="135E87AA5215FFEE2997FA9FFA724ABE" ID-DOI="http://doi.org/10.5281/zenodo.12211195" ID-GBIF-Taxon="233318010" ID-Zenodo-Dep="12211195" LSID="urn:lsid:plazi:treatment:135E87AA5215FFEE2997FA9FFA724ABE" httpUri="http://treatment.plazi.org/id/135E87AA5215FFEE2997FA9FFA724ABE" lastPageNumber="461" pageId="10" pageNumber="461">
<subSubSection id="D3ED65375215FFEE2997FA9FFD34493B" box="[151,730,1374,1401]" pageId="10" pageNumber="461" type="nomenclature">
<paragraph id="9B4836BC5215FFEE2997FA9FFD34493B" blockId="10.[151,730,1374,1401]" box="[151,730,1374,1401]" pageId="10" pageNumber="461">
<heading id="C00081D05215FFEE2997FA9FFD34493B" bold="true" box="[151,730,1374,1401]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
<emphasis id="A983EAAE5215FFEE2997FA9FFD34493B" bold="true" box="[151,730,1374,1401]" pageId="10" pageNumber="461">
Key to the species of the subgenus
<taxonomicName id="5CF74D3F5215FFEE2B46FA9EFD34493B" ID-CoL="8MGX7" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[582,730,1374,1401]" class="Insecta" family="Carabidae" genus="Micronialoe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="A983EAAE5215FFEE2B46FA9EFD34493B" bold="true" box="[582,730,1374,1401]" italics="true" pageId="10" pageNumber="461">Micronialoe</emphasis>
</taxonomicName>
</emphasis>
</heading>
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</subSubSection>
<subSubSection id="D3ED65375215FFEE2997FA64FA724ABE" pageId="10" pageNumber="461" type="key">
<keyStep id="20032E195215FFEE2997FA64FA724A53" pageId="10" pageNumber="461">
<paragraph id="9B4836BC5215FFEE2997FA64FA724995" blockId="10.[151,1437,1444,1788]" pageId="10" pageNumber="461">
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1. Pronotum with rudimentary or without outer basal fovea (
<figureCitation id="03CC2A395215FFEE2A16FA64FC8D49FB" box="[790,867,1444,1466]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–H; 0.2 mm I–J. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Fig. 8C</figureCitation>
); median lobe of aedeagus rather slender, apical orifice on dorsal side (
<figureCitation id="03CC2A395215FFEE2856FA01FE504994" box="[342,446,1473,1495]" captionStart="FIGURE 8" captionStartId="9.[151,250,1464,1488]" captionTargetBox="[161,1436,190,1431]" captionTargetId="figure-18@9.[151,1436,181,1440]" captionTargetPageId="9" captionText="FIGURE 8. Male habitus and genitalia of Pterostichus (Micronialoe) chogyesanus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsal view; (G) median lobe of aedeagus, ventral view; (H) median lobe of aedeagus, left lateral view; (I) left paramere; (J) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–H; 0.2 mm I–J. Abbreviation: CS: C-shaped sclerite." figureDoi="http://doi.org/10.5281/zenodo.12190405" httpUri="https://zenodo.org/record/12190405/files/figure.png" pageId="10" pageNumber="461">Figs 8E–H</figureCitation>
)...........................................
<taxonomicName id="5CF74D3F5215FFEE2AD4FA02FA724995" ID-CoL="4Q3X5" authority="Park, Kwon &amp; Lafer, 1996" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[980,1436,1473,1495]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="chogyesanus" subGenus="Micronialoe">
<emphasis id="A983EAAE5215FFEE2AD4FA02FC094995" box="[980,999,1474,1495]" italics="true" pageId="10" pageNumber="461">P.</emphasis>
(
<emphasis id="A983EAAE5215FFEE2AF3FA02FBE04995" box="[1011,1038,1474,1495]" italics="true" pageId="10" pageNumber="461">M.</emphasis>
)
<emphasis id="A983EAAE5215FFEE2D1BFA01FB784994" box="[1051,1174,1473,1494]" italics="true" pageId="10" pageNumber="461">chogyesanus</emphasis>
<bibRefCitation id="FF664B4D5215FFEE2D9CFA01FA724995" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[1180,1436,1473,1495]" pageId="10" pageNumber="461" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="9B4836BC5215FFEE2997FA1FFA724A53" blockId="10.[151,1437,1444,1788]" pageId="10" pageNumber="461">
<keyLead id="200695895215FFEE2997FA1FFA724A53" pageId="10" pageNumber="461">
- Pronotum with outer basal fovea more or less distinct (
<figureCitation id="03CC2A395215FFEE2BF3FA1EFCAC49B1" box="[755,834,1502,1524]" captionStart="FIGURE 2" captionStartId="2.[151,250,1973,1997]" captionTargetBox="[155,1436,1248,1941]" captionTargetId="figure-445@2.[151,1436,1240,1950]" captionTargetPageId="2" captionText="FIGURE 2. Pterostichus (Micronialoe) spinulosus sp. nov., male. (A) habitus, dorsal view; (B) ditto, ventral view; C head and pronotum, dorsal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12190391" httpUri="https://zenodo.org/record/12190391/files/figure.png" pageId="10" pageNumber="461">Figs 2C</figureCitation>
,
<figureCitation id="03CC2A395215FFEE2A4DFA1EFC8249B1" box="[845,876,1502,1523]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–I; 0.2 mm J–K. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="10" pageNumber="461">7C</figureCitation>
); median lobe of aedeagus stout, apical orifice bent to left (
<figureCitation id="03CC2A395215FFEE29E9FA3CFEBA4A53" box="[233,340,1531,1554]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male A–H, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm A–E, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="10" pageNumber="461">Figs 5A–E</figureCitation>
,
<figureCitation id="03CC2A395215FFEE285DFA3CFE634A53" box="[349,397,1532,1553]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–I; 0.2 mm J–K. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="10" pageNumber="461">7E–I</figureCitation>
).................................................................................... 2
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<keyStep id="20032E195215FFEE2997F9D9FA724AC4" pageId="10" pageNumber="461">
<paragraph id="9B4836BC5215FFEE2997F9D9FA724A09" blockId="10.[151,1437,1444,1788]" pageId="10" pageNumber="461">
<keyLead id="200695895215FFEE2997F9D9FA724A09" pageId="10" pageNumber="461">
2. Median lobe of aedeagus truncate at apex in dorsal view, left lateral margin below apical orifice almost linear, internal sac with three sclerotized areas (
<figureCitation id="03CC2A395215FFEE28C4F9F6FDC34A0E" box="[452,557,1590,1612]" captionStart="FIGURE 5" captionStartId="5.[151,250,1877,1901]" captionTargetBox="[161,1426,850,1841]" captionTargetId="figure-42@5.[151,1436,840,1854]" captionTargetPageId="5" captionText="FIGURE 5. Male and female genitalia of Pterostichus (Micronialoe) spinulosus sp. nov., male A–H, female (I). (A) median lobe of aedeagus, right lateral view; (B) ditto, dorsolateral view; (C) ditto, dorsal view; (D) ditto, ventral view; (E) ditto, left lateral view; (F) left paramere; (G) right paramere; (H) male sternite IX; (I) female genitalia (laterotergite, basal stylomere, and apical stylomere), ventral view. Scale bars: 0.3 mm A–E, H; 0.2 mm F, G, 0.1 mm I. Abbreviations: SP: spiny patch; VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190399" httpUri="https://zenodo.org/record/12190399/files/figure.png" pageId="10" pageNumber="461">Figs 5A–E</figureCitation>
).........................................
<taxonomicName id="5CF74D3F5215FFEE2D28F9F7FA724A09" authority="Kim &amp; Suh" authorityName="Kim &amp; Suh" authorityYear="2024" box="[1064,1436,1590,1612]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="spinulosus" status="sp. nov." subGenus="Micronialoe">
<emphasis id="A983EAAE5215FFEE2D28F9F7FBD54A0E" box="[1064,1083,1591,1612]" italics="true" pageId="10" pageNumber="461">P.</emphasis>
(
<emphasis id="A983EAAE5215FFEE2D47F9F7FB8D4A0E" box="[1095,1123,1591,1612]" italics="true" pageId="10" pageNumber="461">M.</emphasis>
)
<emphasis id="A983EAAE5215FFEE2D6FF9F6FB3A4A09" box="[1135,1236,1590,1611]" italics="true" pageId="10" pageNumber="461">spinulosus</emphasis>
Kim &amp; Suh
<emphasis id="A983EAAE5215FFEE2C50F9F6FA724A09" bold="true" box="[1360,1436,1590,1611]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="B2B057D55215FFEE2C50F9F6FA724A09" box="[1360,1436,1590,1611]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="9B4836BC5215FFEE2997F994FA724AC4" blockId="10.[151,1437,1444,1788]" pageId="10" pageNumber="461">
<keyLead id="200695895215FFEE2997F994FA724AC4" pageId="10" pageNumber="461">
- Median lobe of aedeagus rounded at apex in dorsal view, left lateral margin below apical orifice convex, internal sac with two sclerotized areas (
<figureCitation id="03CC2A395215FFEE288EF9B1FE014AC4" box="[398,495,1649,1671]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–I; 0.2 mm J–K. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="10" pageNumber="461">Figs 7E–I</figureCitation>
) (sclerotized parts for
<taxonomicName id="5CF74D3F5215FFEE2BC8F9B2FCA44AC4" ID-CoL="4Q3X6" authorityName="Morita" authorityYear="2005" box="[712,842,1649,1671]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE5215FFEE2BC8F9B2FD354AC5" box="[712,731,1650,1671]" italics="true" pageId="10" pageNumber="461">P.</emphasis>
(
<emphasis id="A983EAAE5215FFEE2BE7F9B2FCEC4AC5" box="[743,770,1650,1671]" italics="true" pageId="10" pageNumber="461">M.</emphasis>
)
<emphasis id="A983EAAE5215FFEE2A0FF9B1FCA44AC4" box="[783,842,1649,1670]" italics="true" pageId="10" pageNumber="461">kaniei</emphasis>
</taxonomicName>
is unknown)...................................... 3
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<keyLead id="200695895215FFEE2997F94EFA724A83" pageId="10" pageNumber="461">
3. Pronotum with vague outer basal fovea; smaller species (6.1–7.0 mm); distributed in Is. Tsushima,
<collectingCountry id="E3E0762C5215FFEE2D99F94FFB3E4AE6" box="[1177,1232,1679,1700]" name="Japan" pageId="10" pageNumber="461">Japan</collectingCountry>
..............................................................................................
<taxonomicName id="5CF74D3F5215FFEE2D97F96CFA724A83" ID-CoL="4Q3X6" authority="Morita, 2005" authorityName="Morita" authorityYear="2005" box="[1175,1436,1707,1730]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="kaniei" subGenus="Micronialoe">
<emphasis id="A983EAAE5215FFEE2D97F96CFB444A83" box="[1175,1194,1708,1729]" italics="true" pageId="10" pageNumber="461">P.</emphasis>
(
<emphasis id="A983EAAE5215FFEE2DB6F96CFB3F4A83" box="[1206,1233,1708,1729]" italics="true" pageId="10" pageNumber="461">M.</emphasis>
)
<emphasis id="A983EAAE5215FFEE2DDEF96CFAF74A83" box="[1246,1305,1708,1729]" italics="true" pageId="10" pageNumber="461">kaniei</emphasis>
<bibRefCitation id="FF664B4D5215FFEE2C1FF96CFA724A83" author="Morita, S." box="[1311,1436,1707,1730]" pageId="10" pageNumber="461" pagination="9 - 12" refId="ref7740" refString="Morita, S. (2005) A New Pterostichus (Coleoptera: Carabidae) from the Islands of Tsushima, West Japan. Entomological Review of Japan, 60 (1), 9 - 12." type="journal article" year="2005">Morita, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9B4836BC5215FFEE2997F909FA724ABE" blockId="10.[151,1437,1444,1788]" pageId="10" pageNumber="461">
<keyLead id="200695895215FFEE2997F909FA724ABE" pageId="10" pageNumber="461">
- Pronotum with distinct outer basal fovea (
<figureCitation id="03CC2A395215FFEE2B75F909FD2F4A9C" box="[629,705,1737,1759]" captionStart="FIGURE 7" captionStartId="8.[151,250,1520,1544]" captionTargetBox="[160,1426,282,1487]" captionTargetId="figure-74@8.[151,1436,273,1496]" captionTargetPageId="8" captionText="FIGURE 7. Male habitus and genitalia of Pterostichus (Micronialoe) bifoveolatus Park, Kwon &amp; Lafer. (A) habitus, dorsal view; (B) habitus, ventral view; (C) head and pronotum, dorsal view; (D) sternite VII; (E) median lobe of aedeagus, right lateral view; (F) median lobe of aedeagus, dorsolateral view; (G) median lobe of aedeagus, dorsal view; (H) median lobe of aedeagus, ventral view; (I) median lobe of aedeagus, left lateral view; (J) left paramere; (K) right paramere. Scale bars: 1 mm A–C; 0.5 mm D; 0.3 mm E–I; 0.2 mm J–K. Abbreviations: VS: ventral sclerite; CS: C-shaped sclerite; C: ventral carina." figureDoi="http://doi.org/10.5281/zenodo.12190403" httpUri="https://zenodo.org/record/12190403/files/figure.png" pageId="10" pageNumber="461">Fig. 7C</figureCitation>
); larger species (&gt;7.0 mm); distributed in Mt. Jirisan,
<collectingCountry id="E3E0762C5215FFEE2DC7F909FAEC4A9C" box="[1223,1282,1737,1758]" name="South Korea" pageId="10" pageNumber="461">Korea</collectingCountry>
...........................................................................
<taxonomicName id="5CF74D3F5215FFEE2AD9F927FA724ABE" ID-CoL="4Q3X4" authority="Park, Kwon &amp; Lafer, 1996" authorityName="Park, Kwon &amp; Lafer" authorityYear="1996" box="[985,1436,1766,1788]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="bifoveolatus" subGenus="Micronialoe">
<emphasis id="A983EAAE5215FFEE2AD9F927FC024ABE" box="[985,1004,1767,1788]" italics="true" pageId="10" pageNumber="461">P.</emphasis>
(
<emphasis id="A983EAAE5215FFEE2AF9F927FBFA4ABE" box="[1017,1044,1767,1788]" italics="true" pageId="10" pageNumber="461">M.</emphasis>
)
<emphasis id="A983EAAE5215FFEE2D21F926FB784AB9" box="[1057,1174,1766,1787]" italics="true" pageId="10" pageNumber="461">bifoveolatus</emphasis>
<bibRefCitation id="FF664B4D5215FFEE2D9CF926FA724ABE" author="Park, J. K. &amp; Kwon, Y. J. &amp; Lafer, G. S." box="[1180,1436,1766,1788]" pageId="10" pageNumber="461" pagination="72 - 77" refId="ref7812" refString="Park, J. K., Kwon, Y. J. &amp; Lafer, G. S. (1996) Classification of the genus Pterostichus Bonelli from Korea (Coleoptera: Harpalidae) I. Micronialoe subgen. nov. Korean Journal of Entomology, 26, 72 - 77." type="journal article" year="1996">Park, Kwon &amp; Lafer, 1996</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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