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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.747.23468" ID-GBIF-Dataset="a2142825-4b3e-4c0b-9113-059e7d717997" ID-PMC="PMC5904522" ID-Pensoft-Pub="1313-2970-747-41" ID-PubMed="29674902" ID-ZBK="A2A19CC3B61A4B64AD85583D736DFDF0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-747-41" ModsDocOrigin="ZooKeys 747" ModsDocTitle="Tubucaalcocki, a new pseudocryptic species of fiddler crab from the Indian Ocean, sister to the southeastern African T.urvillei (H. Milne Edwards, 1852) (Crustacea, Decapoda, Brachyura, Ocypodidae)" checkinTime="1522392109229" checkinUser="pensoft" docAuthor="Shih, Hsi-Te, Chan, Benny K. K. &amp; Ng, Peter K. L." docDate="2018" docId="3F42EA9869698888E0FE81C7ACB59A17" docLanguage="en" docName="ZooKeys 747: 41-62" docOrigin="ZooKeys 747" docSource="http://dx.doi.org/10.3897/zookeys.747.23468" docTitle="Tubuca alcocki Shih, Chan &amp; Ng, 2018, sp. n." docType="treatment" docUuid="0912FA92-20A2-424F-82C0-A337A20A4494" docUuidSource="ZooBank" docVersion="5" lastPageNumber="50" masterDocId="C57F386BFFC4FFA51B35407ED34A776B" masterDocTitle="Tubucaalcocki, a new pseudocryptic species of fiddler crab from the Indian Ocean, sister to the southeastern African T. urvillei (H. Milne Edwards, 1852) (Crustacea, Decapoda, Brachyura, Ocypodidae)" masterLastPageNumber="62" masterPageNumber="41" pageNumber="45" updateTime="1668165645721" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Tubucaalcocki, a new pseudocryptic species of fiddler crab from the Indian Ocean, sister to the southeastern African T. urvillei (H. Milne Edwards, 1852) (Crustacea, Decapoda, Brachyura, Ocypodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shih, Hsi-Te</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chan, Benny K. K.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>747</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>62</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.747.23468</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.747.23468</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-747-41</mods:identifier>
<mods:identifier type="ZBK">A2A19CC3B61A4B64AD85583D736DFDF0</mods:identifier>
<mods:identifier type="ZooBank">A2A19CC3B61A4B64AD85583D736DFDF0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="142670094" LSID="urn:lsid:zoobank.org:act:0912FA92-20A2-424F-82C0-A337A20A4494" httpUri="http://treatment.plazi.org/id/3F42EA9869698888E0FE81C7ACB59A17" lastPageId="9" lastPageNumber="50" pageId="4" pageNumber="45">
<subSubSection pageId="4" pageNumber="45" type="nomenclature">
<paragraph pageId="4" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/0912FA92-20A2-424F-82C0-A337A20A4494" class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="45">sp. n.</taxonomicNameLabel>
Figures 3, 4A, C, 5
<normalizedToken originalValue="A–D">A-D</normalizedToken>
, 6, 7A, C, E, G
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="47" pageId="5" pageNumber="46" type="reference_group">
<paragraph pageId="5" pageNumber="46">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus Dussumieri" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="Dussumieri">
<pageBreakToken pageId="5" pageNumber="46" start="start">Gelasimus</pageBreakToken>
Dussumieri
</taxonomicName>
H. Milne Edwards, 1852: 148, pl. 4(12) [part; Malabar, India];
<bibRefCitation author="Kingsley, JS" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="13" pageNumber="54" pagination="135 - 155" title="Carcinological notes, no. II. - Revision of the Gelasimi." volume="1880" year="1880">Kingsley 1880</bibRefCitation>
: 145 [part; list];
<bibRefCitation author="Chandy, M" journalOrPublisher="Journal of the Bombay Natural History Society" pageId="12" pageNumber="53" pagination="401 - 402" title="New records of Brachyuran decapods from the Gulf of Kutch." volume="70" year="1973">Chandy 1973</bibRefCitation>
: 402 [Gulf of Kutch, W India] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus dussumieri" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="dussumieri">Gelasimus dussumieri</taxonomicName>
H. Milne Edwards, 1852 sensu stricto).
</paragraph>
<paragraph pageId="5" pageNumber="46">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus acutus" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="acutus">Gelasimus acutus</taxonomicName>
-
<bibRefCitation author="Alcock, A" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="11" pageNumber="52" pagination="279 - 456" title="Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea." volume="69" year="1900">Alcock 1900</bibRefCitation>
: 360-361 [Sunderbunds, Mergui; Andamans; Karachi] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus acutus" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="acutus">Gelasimus acutus</taxonomicName>
Stimpson, 1858).
</paragraph>
<paragraph pageId="5" pageNumber="46">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus Urvillei" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="Urvillei">Gelasimus Urvillei</taxonomicName>
-
<bibRefCitation author="Alcock, A" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="11" pageNumber="52" pagination="279 - 456" title="Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea." volume="69" year="1900">Alcock 1900</bibRefCitation>
: 362-363 [Nicobars; Madras; Karachi] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus urvillei" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="urvillei">Gelasimus urvillei</taxonomicName>
H. Milne Edwards, 1852).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca angustifrons" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="angustifrons">
<pageBreakToken pageId="6" pageNumber="47" start="start">Uca</pageBreakToken>
angustifrons
</taxonomicName>
-
<bibRefCitation author="Lundoer, S" journalOrPublisher="Phuket Marine Biological Center Research Bulletin" pageId="14" pageNumber="55" pagination="3 - 11" title="A checklist of the marine Brachyura in the reference collection at PMBC, Thailand." volume="4" year="1974">Lundoer 1974</bibRefCitation>
: 8 [Phuket, SW Thailand];
<bibRefCitation author="Ng, PKL" journalOrPublisher="Phuket Marine Biological Center Research Bulletin" pageId="14" pageNumber="55" pagination="369 - 384" title="A checklist of the brachyuran crabs of Phuket and western Thailand." volume="23" year="2002">Ng and Davie 2002</bibRefCitation>
: 378 [list; Phuket, SW Thailand] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Gelasimus signatus var. angustifrons" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="variety" species="signatus" variety="angustifrons">Gelasimus signatus var. angustifrons</taxonomicName>
De Man, 1892 =
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca bellator" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="bellator">Tubuca bellator</taxonomicName>
(White, 1847)).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Deltuca)" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Deltuca">Uca (Deltuca)</taxonomicName>
[coarctata]
<taxonomicName family="Ocypodidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="urvillei" order="Decapoda" pageId="6" pageNumber="47" rank="species" species="urvillei">urvillei</taxonomicName>
-
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
: 35, 58-61, figs 8B, 9E, pl. 9C, D [part, Pakistan to southern India];
<bibRefCitation author="Frith, DW" journalOrPublisher="Crustaceana" pageId="12" pageNumber="53" pagination="100 - 102" title="Range extensions of fiddler crabs (Decapoda, Brachyura, Ocypodidae) in the north-western Malay Peninsula area." url="https://doi.org/10.1163/156854077X00944" volume="32" year="1977 a">Frith and Frith 1977a</bibRefCitation>
: 100-101 [Phuket, SW Thailand] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Gelasimus urvillei</taxonomicName>
H. Milne Edwards, 1852).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Uca urvillei</taxonomicName>
-
<bibRefCitation author="Frith, DW" journalOrPublisher="Phuket Marine Biological Center Research Bulletin" pageId="12" pageNumber="53" pagination="1 - 37" title="Zonation of macrofauna on a mangrove shore, Phuket Island." volume="10" year="1976">Frith et al. 1976</bibRefCitation>
: 14, 19, 23-24, 28 [Phuket, SW Thailand];
<bibRefCitation author="Tirmizi, NM" journalOrPublisher="Publication of the Centre of Excellence in Marine Biology, University of Karachi, Karachi, Pakistan" pageId="15" pageNumber="56" title="Marine fauna of Pakistan: 5, Crustacea: Brachyura, Brachyrhyncha, Part 1. (Xanthidae, Goneplacidae, Pinnotheridae, Ocypodidae, Grapsidae)." year="1996">Tirmizi and Ghani 1996</bibRefCitation>
: 103-105, fig. 39 [Pakistan];
<bibRefCitation pageId="6" pageNumber="47">Jaroensutasinee et al. 2003</bibRefCitation>
: 1-3 [W Thailand];
<bibRefCitation author="Jaroensutasinee, M" journalOrPublisher="Crustaceana" pageId="13" pageNumber="54" pagination="533 - 551" title="Morphology, density, and sex ratio of fiddler crabs from southern Thailand (Decapoda, Brachyura, Ocypodidae)." url="https://doi.org/10.1163/1568540041718000" volume="77" year="2004">Jaroensutasinee and Jaroensutasinee 2004</bibRefCitation>
: 534, 538, 540-548 [W Thailand];
<bibRefCitation author="Naiyanetr, P" journalOrPublisher="Office of Natural Resources and Environmental Policy and Planning, Bangkok, Thailand" pageId="14" pageNumber="55" title="Checklist of crustacean fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda)." year="2007">Naiyanetr 2007</bibRefCitation>
: 133 [list; Thailand];
<bibRefCitation author="Saher, NU" journalOrPublisher="PhD Thesis, University of Karachi, Karachi" pageId="14" pageNumber="55" title="Population dynamics and biology of fiddler crabs in the mangrove areas of Karachi coast. Karachi, Pakistan, University of Karachi." year="2008">Saher 2008</bibRefCitation>
: 21-22, fig. 2.2, pl. 2.1 [Pakistan];
<bibRefCitation pageId="6" pageNumber="47">Dev Roy and Nandi 2012</bibRefCitation>
: 218 [Nicobar, India];
<bibRefCitation author="Hossain, MAR" journalOrPublisher="IUCN, International Union for Conservation of Nature, Bangladesh Country Office, Dhaka" pageId="13" pageNumber="54" title="Red List of Bangladesh, volume 6: Crustaceans." year="2015">Hossain 2015</bibRefCitation>
: 203, 1 unnumbered fig. [Bangladesh];
<bibRefCitation author="Odhano, S" journalOrPublisher="Biharean Biologist" pageId="14" pageNumber="55" pagination="170 - 172" title="An over clawed (with two enlarge chela) male crab of Ucaurvillei (Ocypodidae: Tubuca: Uca) along the coast of Karachi, Pakistan." volume="9" year="2015">Odhano et al. 2015</bibRefCitation>
: 170-171, figs 1-2 [Pakistan] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Gelasimus urvillei</taxonomicName>
H. Milne Edwards, 1852).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Deltuca) urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei" subGenus="Deltuca">Uca (Deltuca) urvillei</taxonomicName>
-
<bibRefCitation author="Hogarth, PJ" journalOrPublisher="Crustaceana" pageId="13" pageNumber="54" pagination="222 - 223" title="Occurrence of Uca (Deltuca) urvillei (H. Milne Edwards, 1852) in the Saudi Red Sea (Decapoda, Ocypodidae)." url="https://doi.org/10.1163/156854086X00737" volume="51" year="1986">Hogarth 1986</bibRefCitation>
: 222-223 [Red Sea];
<bibRefCitation author="Price, ARG" journalOrPublisher="Journal of Natural History" pageId="14" pageNumber="55" pagination="449 - 464" title="Aspects of mangal ecology along the Red Sea coast of Saudi Arabia." url="https://doi.org/10.1080/00222938700771121" volume="21" year="1987">Price et al. 1987</bibRefCitation>
: 456, 464 [Red Sea];
<bibRefCitation author="Krishnan, S" journalOrPublisher="Records of the Zoological Survey of India" pageId="13" pageNumber="54" pagination="471 - 474" title="Distribution of fiddlers in India." volume="91" year="1992">Krishnan 1992</bibRefCitation>
: 471-472 [Bombay, India] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Gelasimus urvillei</taxonomicName>
H. Milne Edwards, 1852).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Deltuca) dussumieri" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="dussumieri" subGenus="Deltuca">Uca (Deltuca) dussumieri</taxonomicName>
-
<bibRefCitation author="Krishnan, S" journalOrPublisher="Records of the Zoological Survey of India" pageId="13" pageNumber="54" pagination="471 - 474" title="Distribution of fiddlers in India." volume="91" year="1992">Krishnan 1992</bibRefCitation>
: 471-472 [Bombay, India] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus dussumieri" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="dussumieri">Gelasimus dussumieri</taxonomicName>
H. Milne Edwards, 1852)
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Tubuca) urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei" subGenus="Tubuca">Uca (Tubuca) urvillei</taxonomicName>
-
<bibRefCitation author="Beinlich, B" journalOrPublisher="Zoologische Mededelingen" pageId="11" pageNumber="52" pagination="9 - 32" title="Materials for a more stable subdivision of the genus Uca Leach." volume="80" year="2006">Beinlich and von Hagen 2006</bibRefCitation>
: 10, 14, 25, fig. 7f, k [Thailand; India] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Gelasimus urvillei</taxonomicName>
H. Milne Edwards, 1852).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Tubuca) acuta" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="acuta" subGenus="Tubuca">Uca (Tubuca) acuta</taxonomicName>
-
<bibRefCitation author="Trivedi, DJ" journalOrPublisher="Electronic Journal of Environmental Sciences" pageId="15" pageNumber="56" pagination="23 - 31" title="Crustacean fauna of Gujarat state of India: a review." volume="8" year="2015">Trivedi et al. 2015</bibRefCitation>
: 27 [Gujarat, India] (not
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus acutus" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="acutus">Gelasimus acutus</taxonomicName>
Stimpson, 1858).
</paragraph>
<paragraph pageId="6" pageNumber="47">
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca urvillei" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="urvillei">Tubuca urvillei</taxonomicName>
-
<bibRefCitation author="Shih, HT" journalOrPublisher="Raffles Bulletin of Zoology" pageId="15" pageNumber="56" pagination="139 - 175" title="Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera." volume="64" year="2016">Shih et al. 2016</bibRefCitation>
: 159, 174 [part], fig. 12A.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="47" type="materials_examined">
<paragraph pageId="6" pageNumber="47">Material examined.</paragraph>
<paragraph pageId="6" pageNumber="47">Holotype: ♂ (CW 30.1 mm, CL 17.9 mm; PL 58.2 mm) (ZRC 2017.1278), Ranong mangroves, Thailand, coll. H.-T. Shih et al., 27 May 2012. Paratypes: 2 ♂♂ (CW 22.4-29.9 mm), 1 ♀ (CW 25.1 mm) (NCHUZOOL 13661), 1 ♂ (CW 29.5 mm) (NCHUZOOL 14896), 13 ♂♂ (CW 14.7-31.2 mm), 4 ♀♀ (CW 19.9-24.1 mm), 1 ovig. ♀ (CW 25.7 mm) (NCHUZOOL 14905), same data as holotype; 1 ♂ (CW 24.6 mm), 1 ovig. ♀ (CW 14.8 mm) (ZRC 2017.1279), Kamphuan mangroves, Ranong, Thailand, 9 Sep. 2000; 1 ♂ (CW 24.0 mm) (ZRC 2001.2347), Ranong, Thailand, coll. P. Clark, 7 Nov. 2001.</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47" type="materials_examined">
<paragraph pageId="6" pageNumber="47">Other material.</paragraph>
<paragraph lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47">
Thailand: 2 ♂♂ (CW 17.8-26.1 mm) (ZRC 1988.616-617), Phuket, coll. D. H. Murphy, 12 Nov. 1987; 2 ♂♂ (CW 20.3-22.4 mm) (ZRC 1999.1131), mangrove area south east of Phuket Town ca. 8 km, W. B. Jeffries and H. K. Voris, 14 June 1990; 4 ♂♂ (CW 13.0-16.9 mm), 1 ♀ (CW 18.8 mm), 1 ovig. ♀ (CW 19.8 mm) (NCHUZOOL 14897), 6 ♂♂ (CW 9.0-12.4 mm), 3 ♀♀ (CW 11.4-14.0 mm), 2 ovig. ♀♀ (CW 13.9-14.8 mm) (NCHUZOOL 14906), Chalong Bay, Phuket, coll. H.-T. Shih et al., 28 May 2012; 4 ♂♂ (CW 4.4-16.9 mm), 1 ♀ (CW 12.8 mm), 1 ovig. ♀ (CW 14.7 mm) (NCHUZOOL 14898), Laem Tukkae, Phuket, coll. H.-T. Shih et al., 29 May 2012; 1 ♂ (CW 17.2 mm), 2 ♀♀ (CW 14.0-15.6 mm) (NCHUZOOL 14907), Tha Thiap Ruea Bang Rong, Phuket, coll. H.-T. Shih et al., 30 May 2012. India: 1 ♂ (CW 17.7 mm) (NCHUZOOL 14925), 1 ♂ (CW 19.0 mm) (NCHUZOOL 14899), 1 ♂ (CW 12.6 mm) (NCHUZOOL
<pageBreakToken pageId="7" pageNumber="48" start="start">14901</pageBreakToken>
), 1 ♀ (CW 17.5 mm) (NCHUZOOL 14902), 13 ♂♂ (CW 9.9-18.2 mm), 3 ♀♀ (CW 11.4-17.9 mm), 1 ovig. ♀ (CW 19.9 mm) (NCHUZOOL 14903), Mumbai, coll. H.-N. Chen et al., 17 Mar. 2010; 1 ♀ (CW 22.6 mm) (NCHUZOOL 14900), Diu mangroves, coll. K. Wong, 20 Mar. 2010.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="diagnosis">
<paragraph pageId="8" pageNumber="49">
<pageBreakToken pageId="8" pageNumber="49" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="49">
Male. Carapace (Figs 3A, 4A, 6A, C, G, 7A, C, E) trapezoidal, smooth; front narrow, with distinct, narrow median groove; anterolateral angle acutely triangular, directed obliquely anteriorly; anterolateral margin short to moderately long; dorsolateral margin long, definite, strongly converging; one posterolateral stria. Floor of orbit with row of 5-11 tubercles, sometimes with blunt ridge. Major cheliped (Figs 4C, 6B, D) with dactylus usually longer than palm, outer surface of dactylus and pollex each with 1 long groove proximally extending beyond midlength. Fingers of minor cheliped (Figs 3B, D, 6B, D) without conspicuous tooth on either finger. G1 (Fig. 5
<normalizedToken originalValue="A–D">A-D</normalizedToken>
) with distal tube slender, slightly curved to almost straight, distal and proximal parts subequal in width; thumb of moderate length, extending beyond base of distal tube.
</paragraph>
<caption pageId="8" pageNumber="49">
<paragraph pageId="8" pageNumber="49">
Figure 3.
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n.
<normalizedToken originalValue="A–D">A-D</normalizedToken>
holotype (CW 30.1 mm, ZRC 2017.1278) E, F ovigerous female (CW 19.8 mm, NCHUZOOL 14897; Thailand). A dorsal view B frontal view C pleon and telson D, E left minor cheliped F frontal view, with living colouration. Scale bars: 5.0 mm.
</paragraph>
</caption>
<caption pageId="8" pageNumber="49">
<paragraph pageId="8" pageNumber="49">
Figure 4.
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n. A, C holotype (CW 30.1 mm, PL 58.2 mm, ZRC 2017.1278; Thailand);
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
(H. Milne Edwards, 1852) B male (CW 29.7 mm, ZRC 1999.1107; Mayotte). A, B dorsal view C major cheliped.
</paragraph>
</caption>
<caption pageId="8" pageNumber="49">
<paragraph pageId="8" pageNumber="49">
Figure 5. Right G1.
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n.
<normalizedToken originalValue="A–D">A-D</normalizedToken>
holotype (CW 30.1 mm, ZRC 2017.1278; Thailand)
<normalizedToken originalValue="E–H">E-H</normalizedToken>
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
(H. Milne Edwards, 1852), male (CW 29.7 mm, ZRC 1999.1107; Mayotte). A, E mesial view B, F mesial view of distal part C, G lateral view D, H lateral view of distal part. Scale bars: A, C, E, G 5.0 mm B, D, F, H 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="49">Female. Anterolateral angle more broadly triangular; anterolateral margin moderately long, joining dorsolateral margin as convex structure (Fig. 7G). Floor of orbit with row of 17-19 tubercles (Figs 3F, 6F). Fingers of cheliped (Fig. 3E, F) each with conspicuous tooth on occlusal margin.</paragraph>
<caption pageId="8" pageNumber="49">
<paragraph pageId="8" pageNumber="49">
Figure 6.
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n.
<normalizedToken originalValue="A–G">A-G</normalizedToken>
variation of the live colouration. A, B holotype (CW 30.1 mm, ZRC 2017.1278; Thailand) C adult male (not collected; Phuket, Thailand) D young male (CW 13.0 mm, NCHUZOOL 14897; Thailand) E ovigerous female (CW 19.8 mm, NCHUZOOL 14897, Thailand) F, G females in the field (not captured; Phuket, Thailand) H habitat in Ranong, Thailand.
</paragraph>
</caption>
<caption pageId="8" pageNumber="49">
<paragraph pageId="8" pageNumber="49">
Figure 7. Carapace morphology. A, C, E, G
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n., B, D, F, H
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
(H. Milne Edwards, 1852). A holotype male (CW 30.1 mm, ZRC 2017.1278; Thailand) B male (CW 34.9 mm, NCHUZOOL 14895; Kenya) C male (CW 29.5 mm, NCHUZOOL 14896; Ranong, Thailand) D male (CW 29.7 mm, ZRC 1999.1107; Mayotte) E male (CW 27.8 mm, NCHUZOOL 14905; Thailand) F male (CW 27.9 mm, NCHUZOOL 14895; Kenya) G female (CW 22.6 mm, NCHUZOOL 14900; India) H female (CW 22.9 mm, SMF 19985; Kenya).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="colouration in life">
<paragraph pageId="8" pageNumber="49">Colouration in life.</paragraph>
<paragraph pageId="8" pageNumber="49">
Adults with carapace and legs brown or dark brown, posterior part gray, especially in females (Fig. 6A, C, E). Some females with anterolateral angles orange (Fig. 6E, F) or with dark blotches on blue carapace (Fig. 6G). Major cheliped with fingers white; lower palm deep yellow in large individuals, orange in young individuals; upper palm brown (Fig. 6
<normalizedToken originalValue="B–D">B-D</normalizedToken>
). Females sometimes with minor chelipeds orange, sometimes with tint of blue (Figs 3F, 6F, G).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="ecological notes">
<paragraph pageId="8" pageNumber="49">Ecological notes.</paragraph>
<paragraph pageId="8" pageNumber="49">
In western Thailand, this species inhabits muddy banks of mangroves (Fig. 6H) and is sympatric with several species of fiddler crabs, including
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Austruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austruca annulipes" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="annulipes">Austruca annulipes</taxonomicName>
(H. Milne Edwards, 1837),
<taxonomicName lsidName="A. bengali" pageId="8" pageNumber="49" rank="species" species="bengali">A. bengali</taxonomicName>
,
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca forcipata" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="forcipata">Tubuca forcipata</taxonomicName>
(Adams &amp; White, 1849) and
<taxonomicName lsidName="T. paradussumieri" pageId="8" pageNumber="49" rank="species" species="paradussumieri">T. paradussumieri</taxonomicName>
(cf.
<bibRefCitation author="Frith, DW" journalOrPublisher="Crustaceana" pageId="12" pageNumber="53" pagination="100 - 102" title="Range extensions of fiddler crabs (Decapoda, Brachyura, Ocypodidae) in the north-western Malay Peninsula area." url="https://doi.org/10.1163/156854077X00944" volume="32" year="1977 a">Frith and Frith 1977a</bibRefCitation>
,
<bibRefCitation author="Frith, DW" journalOrPublisher="Phuket Marine Biological Center Research Bulletin" pageId="12" pageNumber="53" pagination="1 - 13" title="Notes on the ecology of fiddler crab populations (Ocypodidae: genus Uca) on Phuket, Surin Nua and Yao Yai Islands, western peninsular Thailand." volume="25" year="1978">1978</bibRefCitation>
; this study). In Pakistan, this species is sympatric with
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Austruca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austruca iranica" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="iranica">Austruca iranica</taxonomicName>
(cf.
<bibRefCitation author="Saher, NU" journalOrPublisher="Crustaceana" pageId="14" pageNumber="55" pagination="666 - 673" title="On a new record of the genus Uca (Decapoda, Brachyura, Ocypodidae) found along the coast of Pakistan." url="https://doi.org/10.1163/15685403-00003317" volume="87" year="2014">Saher et al. 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="etymology">
<paragraph pageId="8" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="49">
This species is named after Alfred William Alcock, who first recorded this species from India and Pakistan as &quot;
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca urvillei" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="urvillei">Uca urvillei</taxonomicName>
&quot; (cf.
<bibRefCitation author="Alcock, A" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="11" pageNumber="52" pagination="279 - 456" title="Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea." volume="69" year="1900">Alcock 1900</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="distribution">
<paragraph pageId="8" pageNumber="49">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="49">Western Thailand, India, Pakistan, and the Red Sea (see Remarks).</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49" type="remarks">
<paragraph pageId="8" pageNumber="49">Remarks.</paragraph>
<paragraph lastPageId="9" lastPageNumber="50" pageId="8" pageNumber="49">
Although the number of tubercles on the floor of orbit and thumb morphology of G1 are sometimes useful for distinguishing species of fiddler crabs, they are too variable in
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca alcocki" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alcocki">Tubuca alcocki</taxonomicName>
sp. n. and
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
(
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
: 58-59; this study) to be used. The two species are similar, but can be morphologically distinguished by the characters of the anterolateral angle of the carapace and G1. The anterolateral angle in male
<taxonomicName lsidName="T. alcocki" pageId="8" pageNumber="49" rank="species" species="alcocki">T. alcocki</taxonomicName>
is acutely triangular and directed obliquely anteriorly (Fig. 7A, C, E) (vs. relatively broadly triangular in shape and directed more laterally in position in
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
; Fig. 7B, D, F). In female
<taxonomicName lsidName="T. alcocki" pageId="8" pageNumber="49" rank="species" species="alcocki">T. alcocki</taxonomicName>
, the anterolateral angle is broadly triangular in shape and the anterolateral margin is relatively longer and curves gently to join the dorsolateral margin (Fig. 7G) (vs. anterolateral angle acutely triangular in shape with the anterolateral margin short and merging with the dorsolateral margin in an almost straight line in
<taxonomicName lsidName="T. urvillei" pageId="8" pageNumber="49" rank="species" species="urvillei">T. urvillei</taxonomicName>
; Fig. 7H). The G1 structure is also different. The distal tube of the G1 of
<taxonomicName lsidName="T. alcocki" pageId="8" pageNumber="49" rank="species" species="alcocki">T. alcocki</taxonomicName>
is proportionately more slender, being slightly curved to almost straight, with the widths of the distal and proximal
<pageBreakToken pageId="9" pageNumber="50" start="start">parts</pageBreakToken>
subequal (Fig. 5
<normalizedToken originalValue="A–D">A-D</normalizedToken>
) (vs. distal tube relatively stouter, more distinctly curved and gently tapering towards the tip, with the distal part distinctly narrower than the proximal part in
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
; Fig. 5
<normalizedToken originalValue="E–H">E-H</normalizedToken>
).
</paragraph>
<paragraph pageId="9" pageNumber="50">
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane (1975)</bibRefCitation>
figured specimens of what she referred to as
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
from southeastern Africa and western India, and they agree with the characters of
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
and
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
, respectively. The anterolateral angles of the male lectotype of
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
(Fig. 1A;
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
: pl. 9E) and the male specimen from Tanzania (
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
: fig. 7A) are both broadly triangular. In addition, the G1 distal tubes of the lectotype of
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
as well as those from Somalia and Madagascar figured by
<bibRefCitation author="Vannini, M" journalOrPublisher=")" pageId="15" pageNumber="56" pagination="199 - 226" title="Researches on the coast of Somalia. The shore and dune of Sar Uanle. 31. Ocypodidae and Gecarcinidae (DecapodaBrachyura). Monitore Zoologico Italiano (N. S." volume="14" year="1981">Vannini and Valmori (1981</bibRefCitation>
: fig. 6F) and
<bibRefCitation author="Crosnier, A" journalOrPublisher="Faune de Madagascar" pageId="12" pageNumber="53" pagination="1 - 143" title="Crustaces Decapodes Grapsidae et Ocypodidae." volume="18" year="1965">Crosnier (1965</bibRefCitation>
: figs 195-196) are all relatively stout, curved and tapering towards the tip. As such the material from Tanzania, Somalia and Madagascar should all be referred to
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
sensu stricto.
</paragraph>
<paragraph pageId="9" pageNumber="50">
The specimen from Malabar, western India, and one of the paralectotypes of
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Gelasimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gelasimus dussumieri" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="dussumieri">Gelasimus dussumieri</taxonomicName>
(see discussion earlier), have the G1 distal tube relatively more slender, almost straight, with the distal and proximal parts subequal in width (
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
: fig. 9E) and are thus is clearly referable to
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
. The G1 structures of specimens from Pakistan (
<bibRefCitation author="Saher, NU" journalOrPublisher="PhD Thesis, University of Karachi, Karachi" pageId="14" pageNumber="55" title="Population dynamics and biology of fiddler crabs in the mangrove areas of Karachi coast. Karachi, Pakistan, University of Karachi." year="2008">Saher 2008</bibRefCitation>
: fig. 2.2;
<bibRefCitation author="Tirmizi, NM" journalOrPublisher="Publication of the Centre of Excellence in Marine Biology, University of Karachi, Karachi, Pakistan" pageId="15" pageNumber="56" title="Marine fauna of Pakistan: 5, Crustacea: Brachyura, Brachyrhyncha, Part 1. (Xanthidae, Goneplacidae, Pinnotheridae, Ocypodidae, Grapsidae)." year="1996">Tirmizi and Ghani 1996</bibRefCitation>
: fig. 39) also match that of
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
. Interestingly,
<bibRefCitation author="Hogarth, PJ" journalOrPublisher="Crustaceana" pageId="13" pageNumber="54" pagination="222 - 223" title="Occurrence of Uca (Deltuca) urvillei (H. Milne Edwards, 1852) in the Saudi Red Sea (Decapoda, Ocypodidae)." url="https://doi.org/10.1163/156854086X00737" volume="51" year="1986">Hogarth (1986)</bibRefCitation>
reported &quot;
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Uca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uca (Deltuca) urvillei" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="urvillei" subGenus="Deltuca">Uca (Deltuca) urvillei</taxonomicName>
&quot; from the Red Sea, which was a new record of this species for this region, but without any figure or description. The first author has examined specimens from the Red Sea and they are clearly
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
as well (H-T Shih and BA Kumar, in preparation). The distribution of
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
thus stretches from the northern part of the Indian Ocean (Red Sea) to the Arabian Sea and Andaman Sea.
</paragraph>
<paragraph pageId="9" pageNumber="50">
There are also colour differences between
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
and
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
. While the colouration of females, young males, and juveniles are variable in
<taxonomicName class="Malacostraca" family="Ocypodidae" genus="Tubuca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubuca" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="genus">Tubuca</taxonomicName>
species, the colouration of the adult male carapace is generally more useful (
<bibRefCitation author="Crane, J" journalOrPublisher="Princeton University Press, Princeton, New Jersey" pageId="12" pageNumber="53" title="Fiddler crabs of the world (Ocypodidae: genus Uca)." year="1975">Crane 1975</bibRefCitation>
; von
<bibRefCitation author="Hagen, HO von" journalOrPublisher="Beagle, Records of the Northern Territory Museum of Arts and Sciences" pageId="13" pageNumber="54" pagination="55 - 68" title="The fiddler crabs (Ocypodidae: Uca) of Darwin, Northern Territory, Australia." volume="6" year="1989">Hagen and Jones 1989</bibRefCitation>
;
<bibRefCitation author="Beinlich, B" journalOrPublisher="Zoologische Mededelingen" pageId="11" pageNumber="52" pagination="9 - 32" title="Materials for a more stable subdivision of the genus Uca Leach." volume="80" year="2006">Beinlich and von Hagen 2006</bibRefCitation>
). Adult male
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
sensu stricto have various degrees of blue on the carapace and ambulatory legs (Fig. 2D, E), with the palm of the major cheliped ochraceous to apricot brown (Fig. 2D); while young and females sometimes have pale and dark blotches on a blue background (Fig. 2F). In adult male
<taxonomicName lsidName="T. alcocki" pageId="9" pageNumber="50" rank="species" species="alcocki">T. alcocki</taxonomicName>
, the dorsal surface of the carapace is always dark brown (Fig. 6A, C) whereas in
<taxonomicName lsidName="T. urvillei" pageId="9" pageNumber="50" rank="species" species="urvillei">T. urvillei</taxonomicName>
, it is always blue (Fig. 2D, E).
</paragraph>
</subSubSection>
</treatment>
</document>
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