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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.181.69740" ID-GBIF-Dataset="dca39c99-b65a-42c7-accd-4f7d8e8b421a" ID-PMC="PMC8421323" ID-Pensoft-Pub="1314-2003-181-29" ID-Pensoft-UUID="25B16C1AE4375702BC69E64E28291B0A" ID-PubMed="34557054" ModsDocID="1314-2003-181-29" checkinTime="1630349217782" checkinUser="pensoft" docAuthor="Moroni, Pablo, Martinez, Agustina &amp; Simpson, Michael G." docDate="2021" docId="98C2FE90A1AE51679332AA2856FDBE05" docLanguage="en" docName="PhytoKeys 181: 29-47" docOrigin="PhytoKeys 181" docPubDate="2021-08-30" docSource="http://dx.doi.org/10.3897/phytokeys.181.69740" docTitle="Cryptantha diffusa (Phil.) I. M. Johnst., Contr. Gray Herb. 78: 52. 1927." docType="treatment" docVersion="4" id="25B16C1AE4375702BC69E64E28291B0A" lastPageNumber="29" masterDocId="25B16C1AE4375702BC69E64E28291B0A" masterDocTitle="Nomenclatural revision of Cryptantha (Boraginaceae s. str.) names linked to South American taxa" masterLastPageNumber="47" masterPageNumber="29" pageNumber="29" updateTime="1668140413250" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Nomenclatural revision of Cryptantha (Boraginaceae s. str.) names linked to South American taxa</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moroni, Pablo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5306-476X</mods:nameIdentifier>
<mods:affiliation>Instituto de Botanica Darwinion (ANCEFN-CONICET), Labarden 200, CC 22, B 1642 HYD, San Isidro, Buenos Aires, Argentina</mods:affiliation>
<mods:nameIdentifier type="email">pmoroni@darwin.edu.ar</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinez, Agustina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4768-664X</mods:nameIdentifier>
<mods:affiliation>Instituto de Botanica Darwinion (ANCEFN-CONICET), Labarden 200, CC 22, B 1642 HYD, San Isidro, Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Simpson, Michael G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6197-2132</mods:nameIdentifier>
<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>181</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>29</mods:start>
<mods:end>47</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.181.69740</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.181.69740</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-181-29</mods:identifier>
<mods:identifier type="Pensoft-UUID">25B16C1AE4375702BC69E64E28291B0A</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="185371628" LSID="urn:lsid:plazi:treatment:98C2FE90A1AE51679332AA2856FDBE05" httpUri="http://treatment.plazi.org/id/98C2FE90A1AE51679332AA2856FDBE05" lastPageNumber="29" pageId="0" pageNumber="29">
<subSubSection pageId="0" pageNumber="29" type="nomenclature">
<paragraph pageId="0" pageNumber="29">
9.
<taxonomicName LSID="98C2FE90-A1AE-5167-9332-AA2856FDBE05" authority="(Phil.) I. M. Johnst., Contr. Gray Herb. 78: 52. 1927." authorityName="(Phil.) I. M. Johnst., Contr. Gray Herb. 78: 52. 1927." class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha diffusa" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusa">Cryptantha diffusa (Phil.) I.M. Johnst., Contr. Gray Herb. 78: 52. 1927.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="reference_group">
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha diffusa" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusa">Cryptantha diffusa</taxonomicName>
≡
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium diffusum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusum">Eritrichium diffusum</taxonomicName>
Phil., Linnaea 33: 191. 1864. Type: Chile.
<normalizedToken originalValue="Región">Region</normalizedToken>
de Coquimbo: &quot;Huanta,
<normalizedToken originalValue="Baños">Banos</normalizedToken>
del Toro&quot;, 1860-1861,
<emphasis italics="true" pageId="0" pageNumber="29">H. Volckmann s.n</emphasis>
. (first-step lectotype, designated by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston 1927</bibRefCitation>
, pg. 53: SGO; second-step lectotype, designated here, SGO [bc] SGO000004064 image!; isolectotype, SGO [bc] SGO000004065 image!).
</paragraph>
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha diffusa" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusa">Cryptantha diffusa</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium micranthum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="micranthum">Eritrichium micranthum</taxonomicName>
Phil., Fl. Atac. 38. 1860. Type: Chile. [
<normalizedToken originalValue="Región">Region</normalizedToken>
de Antofagasta:]
<normalizedToken originalValue="Sandón">Sandon</normalizedToken>
, Feb. 1854,
<emphasis italics="true" pageId="0" pageNumber="29">R.A. Philippi s.n</emphasis>
. (first-step lectotype, designated by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston 1927</bibRefCitation>
, pg. 53: SGO; second-step lectotype, designated here: SGO [SGO000004107 digital image!]; isolectotype: SGO [SGO000004106 digital image!]).
</paragraph>
<paragraph pageId="0" pageNumber="29">
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha diffusa" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusa">Cryptantha diffusa</taxonomicName>
=
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha famatinae" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="famatinae">Cryptantha famatinae</taxonomicName>
Brand, Repert. Spec. Nov. Regni Veg. 20: 317. 1924. Type: Argentina. La Rioja: La Incrucijada [sic], Sierra
<taxonomicName authorityName="Ravenna" authorityYear="1972" class="Liliopsida" family="Boraginaceae" genus="Famatina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Famatina" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="genus">Famatina</taxonomicName>
, 1879,
<emphasis italics="true" pageId="0" pageNumber="29">G.H.E.W. Hieronymus &amp; G. Niederlein 466</emphasis>
(lectotype, designated here: CORD [CORD00003765 digital image!]; isolectotype: GH [GH00096383 digital image!]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="29" type="notes">
<paragraph pageId="0" pageNumber="29">Notes.</paragraph>
<paragraph pageId="0" pageNumber="29">
The protologue of
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium diffusum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="diffusum">
<emphasis italics="true" pageId="0" pageNumber="29">Eritrichium diffusum</emphasis>
</taxonomicName>
(
<bibRefCitation author="Philippi, RA" journalOrPublisher="Linnaea" pageId="0" pageNumber="29" pagination="1 - 308" refId="B36" refString="Philippi, RA, 1864. Plantarum novarum Chilensium. Linnaea 33: 1 - 308" title="Plantarum novarum Chilensium." volume="33" year="1864">Philippi 1864</bibRefCitation>
) includes a direct reference to a collection made by Herman Volckmann in Coquimbo, Chile.
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
referred to a specimen lodged at SGO as the
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
of the name, while
<bibRefCitation author="Perez-Moreau, RL" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="29" refId="B30" refString="Perez-Moreau, RL, 1976. Revision del genero Cryptantha en la Argentina (Boraginaceae). Darwiniana 20(1/2): 155-188." title="Revision del genero Cryptantha en la Argentina (Boraginaceae). Darwiniana 20 (1 / 2): 155 - 188." year="1976">
<normalizedToken originalValue="Pérez-Moreau">Perez-Moreau</normalizedToken>
(1976)
</bibRefCitation>
later stated &quot;Chile. Coquimbo: Huanta,
<normalizedToken originalValue="Baños">Banos</normalizedToken>
del Toro, 1860-61, Volckmann (tipo de
<taxonomicName lsidName="E. diffusum" pageId="0" pageNumber="29" rank="species" species="diffusum">
<emphasis italics="true" pageId="0" pageNumber="29">E. diffusum</emphasis>
</taxonomicName>
, SGO)&quot;. Despite the statements of these authors, in the general collection at SGO two sheets were found in agreement with the protologue. The specimens match the diagnosis as coined by Philippi. In this framework, the earliest statement made by
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
is here interpreted as a first-step lectotypification. In order to narrow this broad designation, the sheet SGO000004064 is here selected as a second-step lectotype.
</paragraph>
<paragraph pageId="0" pageNumber="29">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.162865" author="Philippi, RA" journalOrPublisher="INTA, Buenos Aires" pageId="0" pageNumber="29" refId="B35" refString="Philippi, RA, 1860. Florula Atacamensis seu enumeratio plantarum, quas in itinere per desertum Atacamense. Sumptibus E. Anton. https://doi.org/10.5962/bhl.title.162865" title="Florula Atacamensis seu enumeratio plantarum, quas in itinere per desertum Atacamense. Sumptibus E. Anton." url="https://doi.org/10.5962/bhl.title.162865" year="1860">Rudolph A. Philippi (1860)</bibRefCitation>
described
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium micranthum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="micranthum">
<emphasis italics="true" pageId="0" pageNumber="29">Eritrichium micranthum</emphasis>
</taxonomicName>
based on a collection he made in
<normalizedToken originalValue="Sandón">Sandon</normalizedToken>
, Chile, with no further reference. According to
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
, the type element of this species is housed at SGO.
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">
<normalizedToken originalValue="Johnston’s">Johnston's</normalizedToken>
(1927)
</bibRefCitation>
statement is here interpreted as a first-step lectotype designation since two sheets in agreement with the protologue and annotated by Philippi as &quot;
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Eritrichium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eritrichium micranthum" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="micranthum">Eritrichium micranthum</taxonomicName>
&quot; were found at SGO. The specimens involved match the protologue and, thus, the designation of
<bibRefCitation author="Johnston, IM" journalOrPublisher="Contributions from the Gray Herbarium of Harvard University" pageId="0" pageNumber="29" pagination="1 - 118" refId="B16" refString="Johnston, IM, 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contributions from the Gray Herbarium of Harvard University 78: 1 - 118" title="Studies in the Boraginaceae VI. A revision of the South American Boraginoideae." volume="78" year="1927">Johnston (1927)</bibRefCitation>
is here narrowed by selecting the sheet SGO000004107 as a second step-lectotype.
</paragraph>
<paragraph pageId="0" pageNumber="29">
In the protologue of
<taxonomicName class="Magnoliopsida" family="Boraginaceae" genus="Cryptantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cryptantha famatinae" order="Boraginales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="famatinae">
<emphasis italics="true" pageId="0" pageNumber="29">Cryptantha famatinae</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.1002/fedr.19240200617" author="Brand, A" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="0" pageNumber="29" pagination="317 - 320" refId="B3" refString="Brand, A, 1924. Decas specierum novarum quinta. Repertorium Specierum Novarum Regni Vegetabilis 20 (6-21): 317 - 320, DOI: https://doi.org/10.1002/fedr.19240200617" title="Decas specierum novarum quinta." url="https://doi.org/10.1002/fedr.19240200617" volume="20" year="1924">Brand (1924)</bibRefCitation>
cited a collection made by Georg H. E. W. Hieronymus and Gustav Niederlein in the province of La Rioja, Argentina. No original collections could be traced at B, where Brand worked (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.48631" author="Stafleu, FA" journalOrPublisher="Anales del Museo Buenos Aires" pageId="0" pageNumber="29" refId="B46" refString="Stafleu, FA, Cowan, RS, 1976. Taxonomic Literature Vol. 1, a Selective Guide to Botanical Publications and Collections with Dates, Commentaries and Types, ed. 2. Bohn, Scheltema &amp; Holkema, Utrecht. https://doi.org/10.5962/bhl.title.48631" title="Taxonomic Literature Vol. 1, a Selective Guide to Botanical Publications and Collections with Dates, Commentaries and Types, ed. 2. Bohn, Scheltema &amp; Holkema, Utrecht." url="https://doi.org/10.5962/bhl.title.48631" year="1976">Stafleu and Cowan 1976</bibRefCitation>
), but duplicates of Hieronymus and
<normalizedToken originalValue="Niederlein’s">Niederlein's</normalizedToken>
collection were located at CORD and GH. These sheets agree with the diagnosis and cited locality found in the protologue. Thus, the sheet CORD00003765 is here selected as lectotype of the name.
</paragraph>
</subSubSection>
</treatment>
</document>
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