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<mods:title id="000209E9639C4349198C2EF45FBA6ED4">Revision of the Drosophila bromeliae Species Group (Diptera: Drosophilidae): Central American, Caribbean, and Andean Species</mods:title>
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<paragraph id="EC16BC33E924FF8EFE6DFF67FCD1FF59" blockId="22.[473,871,225,251]" box="[473,871,225,251]" pageId="22" pageNumber="23">
<heading id="B75E0B5FE924FF8EFE6DFF67FCD1FF59" box="[473,871,225,251]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="5">
<taxonomicName id="2BA9C7B0E924FF8EFE6DFF67FCD1FF59" authority="Sturtevant, 1916" authorityName="Sturtevant" authorityYear="1916" box="[473,871,225,251]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="florae">
<emphasis id="DEDD6021E924FF8EFE6DFF67FD16FF59" box="[473,672,225,251]" italics="true" pageId="22" pageNumber="23">Drosophila florae</emphasis>
<bibRefCitation id="8838C1C2E924FF8EFD13FF67FCD1FF59" author="Sturtevant, A. H." box="[679,871,225,251]" pageId="22" pageNumber="23" pagination="323 - 343" refId="ref25743" refString="Sturtevant, A. H. 1916. Notes on North American Drosophilidae, with descriptions of 23 new species. Annals of the Entomological Society of America 9: 323 - 343." type="journal article" year="1916">Sturtevant, 1916</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="A4B3EFB8E924FF8EFDE5FE9FFD59FE91" box="[593,751,281,307]" pageId="22" pageNumber="23" type="description">
<paragraph id="EC16BC33E924FF8EFDE5FE9FFD59FE91" blockId="22.[593,751,281,307]" box="[593,751,281,307]" pageId="22" pageNumber="23">
<heading id="B75E0B5FE924FF8EFDE5FE9FFD59FE91" box="[593,751,281,307]" centered="true" fontSize="36" level="3" pageId="22" pageNumber="23" reason="7">
<figureCitation id="7492A0B6E924FF8EFDE5FE9FFD59FE91" box="[593,751,281,307]" captionStart="FIG" captionStartId="22.[120,162,1625,1646]" captionText="FIG. 11. Female terminalia (oviscapts and spermathecal capsules) of two species in the bromeliae group. AH. Drosophila bromeliae Sturtevant. F. Holotype specimen of D. bromeliae. I, J. Drosophila florae Sturtevant, holotype specimen. AE. Oviscapts in ventral view. FI. Same, lateral view of apex. Numbers (G, I) refer to homologous oviscapt pegs in these and subsequent diagrams. All structures to the same scale; see text for specimen numbers." pageId="22" pageNumber="23">Figures 11IJ</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4B3EFB8E924FF8EFFCCFEC6FDABFEFB" box="[120,541,320,345]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="EC16BC33E924FF8EFFCCFEC6FDABFEFB" blockId="22.[120,541,320,345]" box="[120,541,320,345]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="CCB99B1DE924FF8EFFCCFEC6FDABFEFB" box="[120,541,320,345]" pageId="22" pageNumber="23">
<treatmentCitation id="6D089A22E924FF8EFFCCFEC6FDAFFEFB" author="Sturtevant, A. H." box="[120,537,320,345]" page="339" pageId="22" pageNumber="23" year="1916">
<taxonomicName id="2BA9C7B0E924FF8EFFCCFEC6FDAFFEFB" authority="Sturtevant, 1916: 339" authorityName="Sturtevant" authorityPageNumber="339" authorityYear="1916" box="[120,537,320,345]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="florae">
<emphasis id="DEDD6021E924FF8EFFCCFEC6FE9BFEFA" box="[120,301,320,344]" italics="true" pageId="22" pageNumber="23">Drosophila florae</emphasis>
<bibRefCitation id="8838C1C2E924FF8EFE80FEC7FDAFFEFB" author="Sturtevant, A. H." box="[308,537,320,345]" pageId="22" pageNumber="23" pagination="323 - 343" refId="ref25743" refString="Sturtevant, A. H. 1916. Notes on North American Drosophilidae, with descriptions of 23 new species. Annals of the Entomological Society of America 9: 323 - 343." type="journal article" year="1916">Sturtevant, 1916: 339</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A4B3EFB8E924FF8EFF1CFEF1FE8DFDA3" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="EC16BC33E924FF8EFF1CFEF1FE8DFDA3" blockId="22.[120,1225,375,1559]" pageId="22" pageNumber="23">
DIAGNOSIS: Body yellowish, large, thorax length
<quantity id="2B5111D6E924FF8EFD55FEF1FCFDFE33" box="[737,843,375,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="22" pageNumber="23" unit="mm" value="1.11">1.11 mm</quantity>
; prescutellars large, with smaller pair laterally, prescutellar index 0.56; oviscapt long, length 5.0× the depth (in lateral view); spermathecal capsule unique in group: spherical, with short introvert and sclerotized funnel within introvert.
</paragraph>
</subSubSection>
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<paragraph id="EC16BC33E924FF8EFF1CFD8AFE39FC45" blockId="22.[120,1225,375,1559]" pageId="22" pageNumber="23">
DESCRIPTION: (based on
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,
<specimenCount id="FAAF77BAE924FF8EFDE3FD8DFDD9FD84" box="[599,623,523,550]" pageId="22" pageNumber="23" type="female"></specimenCount>
): Head: Frons dark yellow-ochraceous, pollinose on fronto-orbital plates and ocellar triangle; frontal vittae relatively broad, golden, shiny, finely striate. Frontal length
<quantity id="2B5111D6E924FF8EFE6DFDD1FDFFFDD3" box="[473,585,599,625]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="22" pageNumber="23" unit="mm" value="0.32">0.32 mm</quantity>
; frontal index 1.03, top to bottom width ratio 1.22. Ocellar triangle only slightly raised above rest of front, dark brown between ocelli; anterior apex of triangle ca. 0.7× length of front. Orbital setae dark brown-bronze in color, or1:or3 ratio 1.06, or2:or1 ratio 0.50, lengths of postocellar setae 0.47× and ocellar setae 0.65× that of frontal length; bases of ocellar setae outside of tangent between median and lateral ocelli; postocellar setae slightly convergent; vt index 1.10. Vibrissal index 0.53. Face slightly darker than front. Carina well developed, frontal edge somewhat flattened, with faint sulcus; edge width ca. 0.7× width of pedicel. Cheek index 0.11. Eye index 1.57. Occiput (not observable). Basal antennomeres slightly darker than face and front; arista with 4 dorsal and 2 ventral branches, plus small terminal fork. Proboscis very light brown; palpus lighter, with one apical and one subapical seta.
</paragraph>
<paragraph id="EC16BC33E924FF8EFF1CFC77FE2CFAB3" blockId="22.[120,1225,375,1559]" pageId="22" pageNumber="23">
Thorax length
<quantity id="2B5111D6E924FF8EFEE3FC74FE75FBAE" box="[343,451,1010,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="22" pageNumber="23" unit="mm" value="1.11">1.11 mm</quantity>
. Scutum deep ochraceous yellow, pollinose; 8 rows of acrostichal setulae. One large pair of prescutellar setae, lateral pair and pair just anterior to prescutellars barely differentiated from acrostichals; lengths of large prescutellars 0.56× that of posterior dorsocentral setae; h index 0.86. Transverse distance of dorsocentral setae 4.0× the longitudinal distance. Scutellum same color as scutum, pollinose; basal scutellars convergent, apical scutellars cruciate for ca. 0.5× their lengths; scutellar index 0.83. Pleura same color as scutum, sterno index 0.71. Halter stem same color as scutellum; bulb slightly infuscate brown. Legs lighter than scutum and pleura.
</paragraph>
<paragraph id="EC16BC33E924FF8EFF1CFA9AFC2AFA23" blockId="22.[120,1225,375,1559]" pageId="22" pageNumber="23">
Wing hyaline, veins light brown, length
<quantity id="2B5111D6E924FF8EFD3FFA9AFD41FA95" box="[651,759,1308,1335]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="mm" value="2.2">2.20 mm</quantity>
, length to width ratio 2.36. Indices: C, 2.38; ac, 2.47; hb, 0.54, 4-C, 1.00; 4V, 1.75; 5x, 1.58; M, 0.57. Wing tip not pointed; vein R
<subScript id="702DBE76E924FF8EFB1EFAC9FB7EFAC2" attach="left" box="[1194,1224,1359,1376]" fontSize="7" pageId="22" pageNumber="23">2+3</subScript>
virtually straight; section of C near Sc break with 2 thicker spinules.
</paragraph>
<paragraph id="EC16BC33E924FF8EFF1CFA0AFE47FA69" blockId="22.[120,1225,375,1559]" pageId="22" pageNumber="23">Abdomen overall color similar to that of scutum, posterior margins of tergites with faint, diffuse, light brown infuscation.</paragraph>
<paragraph id="EC16BC33E924FF8FFF1CFA51FEF1FE83" blockId="22.[120,1225,375,1559]" lastBlockId="23.[120,1225,225,663]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">Male terminalia: Unknown. Female terminalia: Valves of oviscapt acute but not pointed at tip in lateral view, ca. 400 µm long, with 20 ovisensilla pegs (4 discal and 16 marginal). Spermatheca spherical, height approximately equal to width; basal introvert reaching about 0.60× length of capsule.</paragraph>
</subSubSection>
<caption id="B8D6ECBBE924FF8EFFCCF9DFFEF6F946" pageId="22" pageNumber="23" startId="22.[120,162,1625,1646]">
<paragraph id="EC16BC33E924FF8EFFCCF9DFFEF6F946" blockId="22.[120,1224,1623,1764]" pageId="22" pageNumber="23">
FIG. 11. Female terminalia (oviscapts and spermathecal capsules) of two species in the
<taxonomicName id="2BA9C7B0E924FF8EFC56F9D1FBF5F9CD" box="[994,1091,1623,1647]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="bromeliae">
<emphasis id="DEDD6021E924FF8EFC56F9D1FBF5F9CD" box="[994,1091,1623,1647]" italics="true" pageId="22" pageNumber="23">bromeliae</emphasis>
</taxonomicName>
group.
<emphasis id="DEDD6021E924FF8EFB24F9D1FB7EF9CD" bold="true" box="[1168,1224,1623,1647]" pageId="22" pageNumber="23">AH.</emphasis>
<taxonomicName id="2BA9C7B0E924FF8EFFCCF9F2FE53F92F" authority="Sturtevant. F." authorityName="Sturtevant. F." box="[120,485,1652,1677]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="bromeliae">
<emphasis id="DEDD6021E924FF8EFFCCF9F2FEE7F92E" box="[120,337,1652,1676]" italics="true" pageId="22" pageNumber="23">Drosophila bromeliae</emphasis>
Sturtevant.
<emphasis id="DEDD6021E924FF8EFE67F9F3FE53F92F" bold="true" box="[467,485,1653,1677]" pageId="22" pageNumber="23">F.</emphasis>
</taxonomicName>
Holotype specimen of
<taxonomicName id="2BA9C7B0E924FF8EFD6AF9F3FCD6F92E" box="[734,864,1652,1676]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="bromeliae">
<emphasis id="DEDD6021E924FF8EFD6AF9F3FCD6F92E" box="[734,864,1652,1676]" italics="true" pageId="22" pageNumber="23">D. bromeliae</emphasis>
</taxonomicName>
.
<emphasis id="DEDD6021E924FF8EFCDBF9F3FC21F92F" bold="true" box="[879,919,1653,1677]" pageId="22" pageNumber="23">I, J.</emphasis>
<taxonomicName id="2BA9C7B0E924FF8EFC14F9F2FB72F92F" authority="Sturtevant" authorityName="Sturtevant" authorityYear="1916" box="[928,1220,1652,1677]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="22" pageNumber="23" phylum="Basidiomycota" rank="species" species="florae">
<emphasis id="DEDD6021E924FF8EFC14F9F2FBE6F92E" box="[928,1104,1652,1676]" italics="true" pageId="22" pageNumber="23">Drosophila florae</emphasis>
Sturtevant
</taxonomicName>
, holotype specimen.
<emphasis id="DEDD6021E924FF8EFEF2F917FECEF90B" bold="true" box="[326,376,1681,1705]" pageId="22" pageNumber="23">AE.</emphasis>
Oviscapts in ventral view.
<emphasis id="DEDD6021E924FF8EFD3AF914FD0FF908" bold="true" box="[654,697,1682,1706]" pageId="22" pageNumber="23">FI.</emphasis>
Same, lateral view of apex. Numbers (G, I) refer to homologous oviscapt pegs in these and subsequent diagrams. All structures to the same scale; see text for specimen numbers.
</paragraph>
</caption>
<subSubSection id="A4B3EFB8E925FF8FFF1CFEABFC4AFD34" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="EC16BC33E925FF8FFF1CFEABFD96FDD3" blockId="23.[120,1225,225,663]" pageId="23" pageNumber="24">
<materialsCitation id="5CC1B66EE925FF8FFF1CFEABFD96FDD3" collectingDate="1915-02" collectionCode="AMNH" collectorName="Am. Mus. Nat. Hist. Dept. Invert. &amp; D. A. G. The" country="Cuba" location="Havana" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" stateProvince="La Habana" typeStatus="holotype">
TYPE:
<typeStatus id="33120291E925FF8FFF45FEAAFED6FEE4" box="[241,352,300,326]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
,
<specimenCount id="FAAF77BAE925FF8FFED8FEADFE32FEE4" box="[364,388,299,326]" pageId="23" pageNumber="24" type="female"></specimenCount>
: “
<collectingRegion id="2E6D72D1E925FF8FFE28FEAAFE4FFEE4" box="[412,505,300,326]" country="Cuba" name="La Habana" pageId="23" pageNumber="24">Havana</collectingRegion>
,
<collectingCountry id="94BEFCA3E925FF8FFDB0FEAAFDF3FEE4" box="[516,581,300,326]" name="Cuba" pageId="23" pageNumber="24">Cuba</collectingCountry>
, Jan.
<date id="98179AF3E925FF8FFD30FEAAFD43FEE5" box="[644,757,300,327]" pageId="23" pageNumber="24" value="1915-02">
<collectingDate id="8853631BE925FF8FFD30FEAAFD43FEE5" box="[644,757,300,327]" pageId="23" pageNumber="24" value="1915-02">Feb. 1915</collectingDate>
</date>
” [printed]/ “Melon flow.[er]” [written]/ “TYPE” [printed]/ “
<collectorName id="415CD9E5E925FF8FFEDDFED4FD48FECE" box="[361,766,337,364]" pageId="23" pageNumber="24">Am. Mus. Nat. Hist. Dept. Invert.</collectorName>
Zool. No. [printed] 24141 [written]”/ “
<taxonomicName id="2BA9C7B0E925FF8FFF36FEF1FE78FE33" authority="Sturtevant" authorityName="Sturtevant" authorityYear="1916" box="[130,462,375,401]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="23" pageNumber="24" phylum="Basidiomycota" rank="species" species="florae">Drosophila florae Sturtevant</taxonomicName>
” [written in Sturtevants hand]. The specimen is double mounted, impaled with a minuten through the pleura; head was dislodged by a prior investigator, but is complete and was stored in a corked, glass microvial pinned to the specimen; it was carefully glued back on to the thorax by
<collectorName id="415CD9E5E925FF8FFE57FE61FDDEFDA3" box="[483,616,487,513]" pageId="23" pageNumber="24">D.A.G. The</collectorName>
thorax, wings, and legs are intact; the abdomen was removed and cleared (dissector unknown) and stored in a corked, glass microvial on same pin as thorax. Oviscapt valves had been separated in the original dissection, making a fully intact ventral view impossible. In
<collectionCode id="8AB824F6E925FF8FFE0AFDD1FDAAFDD3" box="[446,540,599,625]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="23" pageNumber="24">AMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="EC16BC33E925FF8FFF1CFDFAFC4AFD34" blockId="23.[120,1225,225,663]" box="[168,1020,636,663]" pageId="23" pageNumber="24">
OTHER SPECIMENS: Known with certainty only from the
<typeStatus id="33120291E925FF8FFCFFFDFBFCC8FD35" box="[843,894,637,663]" pageId="23" pageNumber="24">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
</treatment>
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