treatments-xml/data/47/52/87/475287B4FFDB2B2EFF2EFE9CFC5FF183.xml
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<document id="0215DE57473D7EF969562DBA4F3CD255" ID-CLB-Dataset="157780" ID-DOI="10.11646/zootaxa.5194.1.1" ID-GBIF-Dataset="9f0439bc-75c3-4b5f-a725-2008ecd536f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7141932" ID-ZooBank="E66D4948-BF8A-414A-9AB5-389AEF9D951B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664862474879" checkinUser="plazi" docAuthor="Haddad, Charles R. &amp; Booysen, Ruan" docDate="2022" docId="475287B4FFDB2B2EFF2EFE9CFC5FF183" docLanguage="en" docName="zootaxa.5194.1.1.pdf" docOrigin="Zootaxa 5194 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptopilos digitus Haddad &amp; Booysen 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="BB6BFFCCFFC22B34FFB9FFA1FFABF65F" masterDocTitle="The ground spider genera Leptodrassex Murphy, 2007 and Leptopilos Levy, 2009 (Araneae: Gnaphosidae) in southern Africa, including the description of a new genus and seven new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="26" updateTime="1699387069486" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="24A19EAA0C928E20596340B3AA56FB88">The ground spider genera Leptodrassex Murphy, 2007 and Leptopilos Levy, 2009 (Araneae: Gnaphosidae) in southern Africa, including the description of a new genus and seven new species</mods:title>
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<mods:namePart id="08225E7E96B302CB711FDAB8339E5AD9">Haddad, Charles R.</mods:namePart>
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<taxonomicName id="08FB4D21FFDB2B2DFF2EFE9CFEC5F707" authority="Haddad &amp; Booysen, 2022" authorityName="Haddad &amp; Booysen" authorityYear="2022" box="[151,366,317,344]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="digitus" status="sp. nov.">
<emphasis id="FD8FEAB0FFDB2B2DFF2EFE9CFEC5F707" bold="true" box="[151,366,317,344]" italics="true" pageId="25" pageNumber="26">Leptopilos digitus</emphasis>
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<figureCitation id="57C02A27FFDB2B2DFF2EFEC0FE90F724" box="[151,315,353,379]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1407,1431]" captionTargetBox="[151,1436,181,1383]" captionTargetId="figure-17@19.[151,1436,181,1383]" captionTargetPageId="19" captionText="FIGURES 8792. Dorsal habitus of Leptopilos species: 87, 88. L. butleri sp. nov. female (87) and male (88); 89, 90. L. digitus sp. nov. female (89) and male (90); 91, 92. L. vasivulva sp. nov. female (91) and male (92). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7141974" httpUri="https://zenodo.org/record/7141974/files/figure.png" pageId="25" pageNumber="26">Figures 89, 90</figureCitation>
,
<figureCitation id="57C02A27FFDB2B2DFEFFFEC0FE36F724" box="[326,413,353,379]" captionStart-0="FIGURES 93101" captionStart-1="FIGURES 102113" captionStart-2="FIGURES 114120" captionStartId-0="20.[151,264,1509,1533]" captionStartId-1="21.[151,264,1585,1609]" captionStartId-2="22.[151,264,1837,1861]" captionTargetBox-0="[151,1436,453,1486]" captionTargetBox-1="[152,1436,182,1561]" captionTargetBox-2="[177,1411,489,1814]" captionTargetId-0="figure-116@20.[151,1436,453,1486]" captionTargetId-1="figure-17@21.[151,1436,181,1561]" captionTargetId-2="figure-91@22.[177,1411,489,1814]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURES 93101. Scanning electron micrographs of Leptopilos digitus sp. nov. male (9396, 98101) and female (97): 93. Carapace, dorsal view; 94. Eye region, anterior view; 95. Mouthparts, ventral view; 96. Distal ends of chelicerae, ventral view; 97, 98. Distal ends of chelicerae, posterior views, arrows indicating cheliceral teeth; 99. Serrula teeth; 100. Sternum, ventral view; 101. Dorsal abdominal setae.Abbreviations: PES—promarginal escort setae; RES—retromarginal escort setae; RS—rake setae." captionText-1="FIGURES 102113. Scanning electron micrographs of Leptopilos digitus sp. nov. female (102104) and male (105113): 102, 105. Anterior lateral spinnerets; 103, 106. Posterior median spinnerets; 104, 107. Posterior lateral spinnerets; 108. Patella IV, retrolateral view, arrow indicating lyriform organ at proximal end of patellar indentation; 109. Same, detail of LO; 110. Tarsus I, indicating chemosensory setae (CS), feathery setae (FS), tarsal organ (TO) and trichobothria (TR); 111. Same, detail of trichobothrium base; 112. Same, tarsal organ; 113. Same, tarsal claws, retrolateral-dorsal view, arrowheads indicating claw teeth. Abbreviations: Ac—aciniform gland spigot(s); CS—chemosensory setae; Cy—cylindrical gland spigot(s); FS—feathery setae; MAmp—major ampullate gland spigot(s); mAmp—minor ampullate gland spigot(s); mPi—modified piriform gland spigot(s); n—nubbin; Pi—piriform gland spigot(s); Ta—tartipore; TO—tarsal organ; TR—trichobothria." captionText-2="FIGURES 114120. Scanning electron micrographs of Leptopilos digitus sp. nov. female (114) and male (115121): 114. Epigyne, ventral view, indicating plug (P); 115. Right palp, prolateral view; 116. Left palp, ventro-retrolateral view; 117. Right palpal patella, retrolateral view, arrowhead indicating lyriform organ; 118. Left palpal tibia, ventral view; 119. Distal end of palpal tegulum, ventro-retrolateral view; 120. Same, enlargement of anterior tegular process and embolus.Abbreviations:ATP— anterior tegular process; E—embolus; MA—median apophysis; P—plug; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; RTP—retrolateral tegular process." figureDoi-0="http://doi.org/10.5281/zenodo.7141980" figureDoi-1="http://doi.org/10.5281/zenodo.7141982" figureDoi-2="http://doi.org/10.5281/zenodo.7141984" httpUri-0="https://zenodo.org/record/7141980/files/figure.png" httpUri-1="https://zenodo.org/record/7141982/files/figure.png" httpUri-2="https://zenodo.org/record/7141984/files/figure.png" pageId="25" pageNumber="26">93120</figureCitation>
,
<figureCitation id="57C02A27FFDB2B2DFE11FEC0FDA0F724" box="[424,523,353,379]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">126130</figureCitation>
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<emphasis id="FD8FEAB0FFDB2B2DFF2EFE02FEE9F7E3" bold="true" box="[151,322,418,445]" pageId="25" pageNumber="26">Type material.</emphasis>
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a.s.l.
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&amp;
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(leaf litter, dry river bed) (NMBA 18534).
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:
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:
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(
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, burnt site 2),
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(NMBA 13573)
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.
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,
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a.s.l.
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,
<date id="BB451062FFDB2B2DFCBBFD91FC29F415" box="[770,898,560,586]" pageId="25" pageNumber="26" value="2022-03-27">
<collectingDate id="AB01E98AFFDB2B2DFCBBFD91FC29F415" box="[770,898,560,586]" pageId="25" pageNumber="26" value="2022-03-27">27.III.2022</collectingDate>
</date>
, leg.
<collectorName id="620E5374FFDB2B2DFC7AFD91FBEAF416" box="[963,1089,559,586]" pageId="25" pageNumber="26">C. Haddad</collectorName>
, R.
<collectorName id="620E5374FFDB2B2DFBCBFD91FB52F415" box="[1138,1273,560,586]" pageId="25" pageNumber="26">Booysen, L</collectorName>
. Malope &amp;
<collectorName id="620E5374FFDB2B2DFA3FFD91FF7CF432" pageId="25" pageNumber="26">S. Sibisi</collectorName>
(leaf litter, dry river bed),
<specimenCount id="D9FDFD2BFFDB2B2DFE47FDF2FD8BF432" box="[510,544,595,621]" pageId="25" pageNumber="26" type="male">2♂</specimenCount>
<specimenCount id="D9FDFD2BFFDB2B2DFD9EFDF2FDE1F432" box="[551,586,595,621]" pageId="25" pageNumber="26" type="female">3♀</specimenCount>
(NMBA 18533);
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<materialsCitation id="7F933CFFFFDB2B2DFCA9FDF2FB3EF4CF" ID-GBIF-Occurrence="3920719323" collectingDate="2001-01-15" collectingDateMax="2022-03-27" collectingDateMin="2001-01-15" collectorName="C. Haddad" country="South Africa" county="Namaqua National Park" elevation="240" latitude="-29.566668" location="Prieska district" longLatPrecision="1226" longitude="22.916666" municipality="Koeroebees" pageId="25" pageNumber="26" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Northern Cape" typeStatus="paratype">
<location id="CA246079FFDB2B2DFCA9FDF2FC11F432" LSID="urn:lsid:plazi:treatment:475287B4FFDB2B2EFF2EFE9CFC5FF183:CA246079FFDB2B2DFCA9FDF2FC11F432" box="[784,954,595,621]" country="South Africa" county="Namaqua National Park" latitude="-29.566668" longLatPrecision="1226" longitude="22.916666" municipality="Koeroebees" name="Prieska district" pageId="25" pageNumber="26" stateProvince="Northern Cape">Prieska district</location>
,
<location id="CA246079FFDB2B2DFC7DFDF2FBFDF432" LSID="urn:lsid:plazi:treatment:475287B4FFDB2B2EFF2EFE9CFC5FF183:CA246079FFDB2B2DFC7DFDF2FBFDF432" box="[964,1110,595,621]" country="South Africa" county="Namaqua National Park" latitude="-29.566668" longLatPrecision="1226" longitude="22.916666" municipality="Koeroebees" name="Green Valley" pageId="25" pageNumber="26" stateProvince="Northern Cape">
Green
<location id="CA246079FFDB2B2DFBB7FDF2FBFDF432" LSID="urn:lsid:plazi:treatment:475287B4FFDB2B2EFF2EFE9CFC5FF183:CA246079FFDB2B2DFBB7FDF2FBFDF432" box="[1038,1110,595,621]" country="South Africa" county="Namaqua National Park" latitude="-29.566668" longLatPrecision="1226" longitude="22.916666" municipality="Koeroebees" name="Valley Nuts" pageId="25" pageNumber="26" stateProvince="Northern Cape">Valley</location>
</location>
Nuts,
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,
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,
<elevation id="44D6D191FFDB2B2DFACBFDF3FF43F4CF" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="25" pageNumber="26" unit="m" value="950.0">
<quantity id="08039B47FFDB2B2DFACBFDF3FF06F4CF" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="25" pageNumber="26" unit="m" value="950.0">950 m</quantity>
a.s.l.
</elevation>
,
<date id="BB451062FFDB2B2DFF4AFDD7FECAF4CE" box="[243,353,630,657]" pageId="25" pageNumber="26" value="2001-01-15">
<collectingDate id="AB01E98AFFDB2B2DFF4AFDD7FECAF4CE" box="[243,353,630,657]" pageId="25" pageNumber="26" value="2001-01-15">15.I.2001</collectingDate>
</date>
, leg.
<collectorName id="620E5374FFDB2B2DFE20FDD7FDA4F4CF" box="[409,527,630,656]" pageId="25" pageNumber="26">C. Haddad</collectorName>
(canopy fogging, pistachio orchard),
<specimenCount id="D9FDFD2BFFDB2B2DFC14FDD7FC7BF4CF" box="[941,976,630,656]" pageId="25" pageNumber="26" type="male">1♂</specimenCount>
(NCA 2004/196)
</materialsCitation>
.
<materialsCitation id="7F933CFFFFDB2B2DFB27FDD6FC7BF489" ID-GBIF-Occurrence="3920719351" collectingDate="2021-01-19" collectorName="C. Haddad &amp; R. Booysen" country="South Africa" elevation="355" latitude="-32.398518" location="Tanqua Guesthouse" longLatPrecision="1" longitude="19.845217" municipality="Tankwa Karoo National Park" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="FD8FEAB0FFDB2B2DFB27FDD6FA9CF4CF" box="[1182,1335,630,656]" italics="true" pageId="25" pageNumber="26">
<collectingRegion id="0D3FF840FFDB2B2DFB27FDD6FA9CF4CF" box="[1182,1335,630,656]" country="South Africa" name="Western Cape" pageId="25" pageNumber="26">Western Cape</collectingRegion>
</emphasis>
:
<collectingMunicipality id="2F20ACD8FFDB2B2DFAFDFDD7FE29F4EC" pageId="25" pageNumber="26">Tankwa Karoo National Park</collectingMunicipality>
,
<location id="CA246079FFDB2B2DFE34FD3BFDC7F4EC" LSID="urn:lsid:plazi:treatment:475287B4FFDB2B2EFF2EFE9CFC5FF183:CA246079FFDB2B2DFE34FD3BFDC7F4EC" box="[397,620,665,692]" country="South Africa" latitude="-32.398518" longLatPrecision="1" longitude="19.845217" municipality="Tankwa Karoo National Park" name="Tanqua Guesthouse" pageId="25" pageNumber="26" stateProvince="Western Cape">Tanqua Guesthouse</location>
,
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,
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,
<elevation id="44D6D191FFDB2B2DFC14FD38FB9AF4EC" box="[941,1073,665,692]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="25" pageNumber="26" unit="m" value="355.0">
<quantity id="08039B47FFDB2B2DFC14FD38FC5FF4EB" box="[941,1012,665,692]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="25" pageNumber="26" unit="m" value="355.0">355 m</quantity>
a.s.l.
</elevation>
,
<date id="BB451062FFDB2B2DFB84FD38FB00F4EC" box="[1085,1195,665,691]" pageId="25" pageNumber="26" value="2021-01-19">
<collectingDate id="AB01E98AFFDB2B2DFB84FD38FB00F4EC" box="[1085,1195,665,691]" pageId="25" pageNumber="26" value="2021-01-19">19.I.2021</collectingDate>
</date>
, leg.
<collectorName id="620E5374FFDB2B2DFB5EFD38FAF4F4EC" box="[1255,1375,665,691]" pageId="25" pageNumber="26">C. Haddad</collectorName>
&amp;
<collectorName id="620E5374FFDB2B2DFA3AFD3BFF52F488" pageId="25" pageNumber="26">R. Booysen</collectorName>
(hand collecting, at night around houses),
<specimenCount id="D9FDFD2BFFDB2B2DFD6EFD1DFD51F489" box="[727,762,700,726]" pageId="25" pageNumber="26" type="female">1♀</specimenCount>
(NCA 2021/1016)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87E16529FFDB2B2DFF7EFD41FF5BF542" pageId="25" pageNumber="26" type="etymology">
<paragraph id="CF4436A2FFDB2B2DFF7EFD41FF5BF542" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
<emphasis id="FD8FEAB0FFDB2B2DFF7EFD41FEE5F4A5" bold="true" box="[199,334,736,762]" pageId="25" pageNumber="26">Etymology.</emphasis>
The species name is Latin for finger or toe (digit), referring to the shape of the apical tegular sclerite.
</paragraph>
</subSubSection>
<subSubSection id="87E16529FFDB2B2DFF7EFC87FB08F5F6" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="CF4436A2FFDB2B2DFF7EFC87FB08F5F6" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
<emphasis id="FD8FEAB0FFDB2B2DFF7EFC87FEE9F51F" bold="true" box="[199,322,806,832]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
This species is very distinct in the genus by the short finger-like apical tegular process (
<figureCitation id="57C02A27FFDB2B2DFA8AFC87FA3FF51F" box="[1331,1428,806,832]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 129</figureCitation>
) and the short, slender dorsal tibial apophysis (
<figureCitation id="57C02A27FFDB2B2DFD27FCE8FCAFF53C" box="[670,772,841,867]" captionStart="FIGURES 114120" captionStartId="22.[151,264,1837,1861]" captionTargetBox="[177,1411,489,1814]" captionTargetId="figure-91@22.[177,1411,489,1814]" captionTargetPageId="22" captionText="FIGURES 114120. Scanning electron micrographs of Leptopilos digitus sp. nov. female (114) and male (115121): 114. Epigyne, ventral view, indicating plug (P); 115. Right palp, prolateral view; 116. Left palp, ventro-retrolateral view; 117. Right palpal patella, retrolateral view, arrowhead indicating lyriform organ; 118. Left palpal tibia, ventral view; 119. Distal end of palpal tegulum, ventro-retrolateral view; 120. Same, enlargement of anterior tegular process and embolus.Abbreviations:ATP— anterior tegular process; E—embolus; MA—median apophysis; P—plug; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; RTP—retrolateral tegular process." figureDoi="http://doi.org/10.5281/zenodo.7141984" httpUri="https://zenodo.org/record/7141984/files/figure.png" pageId="25" pageNumber="26">Figs 120</figureCitation>
,
<figureCitation id="57C02A27FFDB2B2DFCA8FCE8FC97F53C" box="[785,828,841,867]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">130</figureCitation>
) of the male palp. Females can be distinguished from congeners by the paired, weakly sclerotized recurved ridges anteriorly in the epigyne, as opposed to a single median hood, and the paired longitudinal ridges forming the lateral margins of the atrium (
<figureCitation id="57C02A27FFDB2B2DFB8CFC2EFB3CF5F6" box="[1077,1175,911,937]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 126</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="87E16529FFDB2B2EFF7EFC13FEE6F1E7" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="CF4436A2FFDB2B2DFF7EFC13FEAAF269" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
<emphasis id="FD8FEAB0FFDB2B2DFF7EFC13FEF2F593" bold="true" box="[199,345,946,972]" pageId="25" pageNumber="26">Description.</emphasis>
Female (
<typeStatus id="10408800FFDB2B2DFE73FC12FD9BF592" box="[458,560,947,973]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
, Namaqua, NMBA 18534). Colouration (
<figureCitation id="57C02A27FFDB2B2DFBA1FC12FBDAF592" box="[1048,1137,947,973]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1407,1431]" captionTargetBox="[151,1436,181,1383]" captionTargetId="figure-17@19.[151,1436,181,1383]" captionTargetPageId="19" captionText="FIGURES 8792. Dorsal habitus of Leptopilos species: 87, 88. L. butleri sp. nov. female (87) and male (88); 89, 90. L. digitus sp. nov. female (89) and male (90); 91, 92. L. vasivulva sp. nov. female (91) and male (92). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7141974" httpUri="https://zenodo.org/record/7141974/files/figure.png" pageId="25" pageNumber="26">Fig. 89</figureCitation>
): carapace and chelicerae creamy-white; endites and labium pale yellow-brown; sternum creamy-white, margins brown at coxae; femora creamy-white, remaining segments creamy-yellow, metatarsi and tarsi slightly darker; abdomen white dorsally and ventrally.
</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EFBE1FCE0F29B" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">Measurements: CL 1.06, CW 0.87, AL 1.71, AW 1.25, TL 2.80. Eye diameters and interdistances: AME 0.10, ALE 0.07, PME 0.08, PLE 0.07, AMEAME 0.05, AMEALE 0.01, PMEPME 0.06, PMEPLE 0.04, ALEPLE 0.01. Leg measurements: I 0.86, 0.40, 0.67, 0.57, 0.44 = 2.94; II 0.81, 0.37, 0.62, 0.56, 0.43 = 2.79; III 0.67, 0.35, 0.49, 0.48, 0.30 = 2.29; IV 1.05, 0.40, 0.84, 0.97, 0.30 = 3.56.</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EFB6CFBB9F372" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">Leg spination: femora: I do 1, II do 1, III do 1, IV do 2 rl 1; patellae spineless; tibiae: I plv 1, II spineless, III pl 2 rl 2 plv 1, IV pl 2 rl 2 plv 1 vt 2; metatarsi: I plv 1 rlv 1, II plv 1 rlv 1, III pl 2 rl 1 plv 1 rlv 1 vt 3, IV pl 3 rl 2 plv 2 vt 3; palp: femur do 1, patella pl 1 do 1, tibia plv 1, tarsus pl 1 rl 1 plv 3 rlv 2.</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EFA97FDB5F382" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
Epigyne with paired, recurved, weakly sclerotized ridges anteriorly, longitudinal ridges laterally forming margins of atrium, with further funnel-shaped ridges medially (
<figureCitation id="57C02A27FFDB2B2DFCB8FAF8FCCFF32C" box="[769,868,1369,1395]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 126</figureCitation>
); copulatory openings small, beneath anterolateral corners of funnel-shaped ridges; copulatory ducts short, looping laterally, then anteriorly and posteriorly, entering teardrop-shaped spermathecae along their anterior margin (
<figureCitation id="57C02A27FFDB2B2DFCFCFA3EFC00F3E5" box="[837,939,1439,1466]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 127</figureCitation>
); fertilization ducts on posterior margin of spermathecae, directed posteriorly.
</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EFA47FB8EF018" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
Male (
<typeStatus id="10408800FFDB2B2DFEA9FA47FEDEF05F" box="[272,373,1510,1536]" pageId="25" pageNumber="26" type="paratype">paratype</typeStatus>
, Prieska, NCA 2004/196). Colouration (
<figureCitation id="57C02A27FFDB2B2DFC83FA47FC3BF05F" box="[826,912,1510,1536]" captionStart="FIGURES 8792" captionStartId="19.[151,264,1407,1431]" captionTargetBox="[151,1436,181,1383]" captionTargetId="figure-17@19.[151,1436,181,1383]" captionTargetPageId="19" captionText="FIGURES 8792. Dorsal habitus of Leptopilos species: 87, 88. L. butleri sp. nov. female (87) and male (88); 89, 90. L. digitus sp. nov. female (89) and male (90); 91, 92. L. vasivulva sp. nov. female (91) and male (92). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7141974" httpUri="https://zenodo.org/record/7141974/files/figure.png" pageId="25" pageNumber="26">Fig. 90</figureCitation>
): carapace and chelicerae light yellow; endites and labium creamy-yellow; sternum creamy-yellow, margins brown at coxae; femora to tibiae creamy-yellow, metatarsi and tarsi slightly darker; abdomen creamy-white dorsally and ventrally.
</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EF9F1FCE0F08B" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">Measurements: CL 1.05, CW 0.79, AL 1.81, AW 1.02, TL 2.98. Eye diameters and interdistances: AME 0.11, ALE 0.08, PME 0.09, PLE 0.08, AMEAME 0.06, AMEALE 0.01, PMEPME 0.05, PMEPLE 0.04, ALEPLE 0.01. Leg measurements: I 0.87, 0.38, 0.67, 0.57, 0.51 = 3.00; II 0.83, 0.37, 0.62, 0.56, 0.48 = 2.86; III 0.65, 0.32, 0.48, 0.46, 0.33 = 2.24; IV 1.06, 0.41, 0.83, 0.95, 0.33 = 3.58.</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EF97CFD6DF162" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">Leg spination: femora: I do 1, II do 1, III do 1 rl 1, IV pl 1 do 2 rl 1; tibiae: I plv 1 rlv 1, II spineless, III pl 2 rl 1 plv 1, IV pl 2 rl 2 plv 1 vt 2; metatarsi: I rlv 1, II rlv 1, III pl 2 rl 1 plv 1 vt 3, IV pl 2 rl 2 plv 1 vt 3; palp: femur do 1, patella and tibia spineless, tarsus plv 1 rlv 1.</paragraph>
<paragraph id="CF4436A2FFDB2B2DFF7EF8E7FC4AF1B1" blockId="25.[151,1443,418,2030]" pageId="25" pageNumber="26">
Palpal tibia very slightly longer than broad, with small spike-like dorso-retrolateral apophysis (
<figureCitation id="57C02A27FFDB2B2DFB43F8E7FA33F13F" box="[1274,1432,1862,1889]" captionStart="FIGURES 114120" captionStartId="22.[151,264,1837,1861]" captionTargetBox="[177,1411,489,1814]" captionTargetId="figure-91@22.[177,1411,489,1814]" captionTargetPageId="22" captionText="FIGURES 114120. Scanning electron micrographs of Leptopilos digitus sp. nov. female (114) and male (115121): 114. Epigyne, ventral view, indicating plug (P); 115. Right palp, prolateral view; 116. Left palp, ventro-retrolateral view; 117. Right palpal patella, retrolateral view, arrowhead indicating lyriform organ; 118. Left palpal tibia, ventral view; 119. Distal end of palpal tegulum, ventro-retrolateral view; 120. Same, enlargement of anterior tegular process and embolus.Abbreviations:ATP— anterior tegular process; E—embolus; MA—median apophysis; P—plug; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; RTP—retrolateral tegular process." figureDoi="http://doi.org/10.5281/zenodo.7141984" httpUri="https://zenodo.org/record/7141984/files/figure.png" pageId="25" pageNumber="26">Figs 115, 116</figureCitation>
,
<figureCitation id="57C02A27FFDB2B2DFF2EF8C8FF6EF1DC" box="[151,197,1897,1923]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">130</figureCitation>
); tegulum oval, with narrow finger-like prolateral apical tegular process, lobate mesal retrolateral tegular process with additional smaller subtriangular lobe, and slender strongly curved median apophysis (
<figureCitation id="57C02A27FFDB2B2DFB37F82DFB5AF1F9" box="[1166,1265,1932,1958]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 129</figureCitation>
); embolus very slender, originating proximally and entering S-shaped groove in prolateral subtegulum, continuing along prolateral margin distally (
<figureCitation id="57C02A27FFDB2B2DFEF6F872FE1BF1B1" box="[335,432,2003,2030]" captionStart="FIGURES 126130" captionStartId="26.[151,264,1484,1508]" captionTargetBox="[185,1411,195,1453]" captionTargetId="figure-18@26.[164,1423,181,1460]" captionTargetPageId="26" captionText="FIGURES 126130. Leptopilos digitus sp. nov., female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="25" pageNumber="26">Fig. 128</figureCitation>
), before entering groove in apical tegular process.
</paragraph>
<caption id="9B84662AFFD82B2EFF2EFA6DFB7FF073" ID-DOI="http://doi.org/10.5281/zenodo.7141988" ID-Zenodo-Dep="7141988" httpUri="https://zenodo.org/record/7141988/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1484,1508]" targetBox="[185,1411,195,1453]" targetPageId="26">
<paragraph id="CF4436A2FFD82B2EFF2EFA6DFB7FF073" blockId="26.[151,1437,1484,1580]" pageId="26" pageNumber="27">
<emphasis id="FD8FEAB0FFD82B2EFF2EFA6DFEDBF3BB" bold="true" box="[151,368,1484,1508]" pageId="26" pageNumber="27">FIGURES 126130.</emphasis>
<taxonomicName id="08FB4D21FFD82B2EFEC1FA6DFD9AF3BB" authority="Haddad &amp; Booysen, 2022" authorityName="Haddad &amp; Booysen" authorityYear="2022" box="[376,561,1484,1508]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="digitus" status="sp. nov.">
<emphasis id="FD8FEAB0FFD82B2EFEC1FA6DFD9AF3BB" box="[376,561,1484,1508]" italics="true" pageId="26" pageNumber="27">Leptopilos digitus</emphasis>
</taxonomicName>
<emphasis id="FD8FEAB0FFD82B2EFD83FA6DFD26F3BB" bold="true" box="[570,653,1484,1508]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="E6BC57CBFFD82B2EFD83FA6DFD26F3BB" box="[570,653,1484,1508]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female epigyne in ventral (126) and dorsal (127) views, and male palp in prolateral (128), ventral (129) and retrolateral (130) views. Abbreviations: ATP—anterior tegular process; E—embolus; MA— median apophysis; RTP—retrolateral tegular process; sRTP—secondary lobe of RTP. Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph id="CF4436A2FFD82B2EFF7EF9DFFB3CF15B" blockId="26.[151,1437,1662,2013]" pageId="26" pageNumber="27">
<emphasis id="FD8FEAB0FFD82B2EFF7EF9DFFD91F0C7" bold="true" box="[199,570,1662,1688]" pageId="26" pageNumber="27">Additional material examined.</emphasis>
<materialsCitation id="7F933CFFFFD82B2EFDFDF9DFFC1AF0BF" ID-GBIF-Occurrence="3920719313" collectingDate="2021-01-14" collectingDateMax="2022-03-27" collectingDateMin="2021-01-14" collectorName="C. Haddad &amp; R. Booysen &amp; R. Christiaan &amp; A. Stander" country="South Africa" elevation="240" latitude="-30.144716" location="Koeroebees" longLatPrecision="1" longitude="17.70295" municipality="Namaqua National Park" pageId="26" pageNumber="27" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Northern Cape">
<emphasis id="FD8FEAB0FFD82B2EFDFDF9DFFC8EF0C7" bold="true" box="[580,805,1662,1688]" pageId="26" pageNumber="27">
<collectingCountry id="B7EC7632FFD82B2EFDFDF9DFFC8BF0C7" box="[580,800,1662,1688]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<emphasis id="FD8FEAB0FFD82B2EFC96F9DFFC70F0C6" box="[815,987,1662,1689]" italics="true" pageId="26" pageNumber="27">
<collectingRegion id="0D3FF840FFD82B2EFC96F9DFFC70F0C6" box="[815,987,1662,1689]" country="South Africa" name="Northern Cape" pageId="26" pageNumber="27">Northern Cape</collectingRegion>
</emphasis>
:
<collectingMunicipality id="2F20ACD8FFD82B2EFC54F9DEFAA8F0C7" box="[1005,1283,1662,1689]" pageId="26" pageNumber="27">Namaqua National Park</collectingMunicipality>
,
<location id="CA246079FFD82B2EFAA8F9DFFA33F0C7" LSID="urn:lsid:plazi:treatment:475287B4FFDB2B2EFF2EFE9CFC5FF183:CA246079FFD82B2EFAA8F9DFFA33F0C7" box="[1297,1432,1662,1688]" country="South Africa" latitude="-30.144716" longLatPrecision="1" longitude="17.70295" municipality="Namaqua National Park" name="Koeroebees" pageId="26" pageNumber="27" stateProvince="Northern Cape">Koeroebees</location>
,
<geoCoordinate id="AACF5065FFD82B2EFF2EF903FE8DF0E3" box="[151,294,1698,1724]" degrees="30" direction="south" minutes="08.683" orientation="latitude" pageId="26" pageNumber="27" precision="1" value="-30.144716">30°08.683S</geoCoordinate>
,
<geoCoordinate id="AACF5065FFD82B2EFE8DF902FE6EF0E2" box="[308,453,1699,1725]" degrees="17" direction="east" minutes="42.177" orientation="longitude" pageId="26" pageNumber="27" precision="1" value="17.70295">17°42.177E</geoCoordinate>
,
<elevation id="44D6D191FFD82B2EFE6BF902FDF3F0E3" box="[466,600,1698,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="26" pageNumber="27" unit="m" value="240.0">
<quantity id="08039B47FFD82B2EFE6BF902FDB1F0E2" box="[466,538,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="26" pageNumber="27" unit="m" value="240.0">240 m</quantity>
a.s.l.
</elevation>
,
<date id="BB451062FFD82B2EFDDFF902FD78F0E2" box="[614,723,1699,1725]" pageId="26" pageNumber="27" value="2021-01-14">
<collectingDate id="AB01E98AFFD82B2EFDDFF902FD78F0E2" box="[614,723,1699,1725]" pageId="26" pageNumber="27" value="2021-01-14">14.I.2021</collectingDate>
</date>
, leg.
<collectorName id="620E5374FFD82B2EFCABF902FC25F0E3" box="[786,910,1698,1725]" pageId="26" pageNumber="27">C. Haddad</collectorName>
, R.
<collectorName id="620E5374FFD82B2EFC05F902FBEFF0E2" box="[956,1092,1699,1725]" pageId="26" pageNumber="27">Booysen, R</collectorName>
. Christiaan &amp;
<collectorName id="620E5374FFD82B2EFB51F902FACAF0E3" box="[1256,1377,1698,1725]" pageId="26" pageNumber="27">A. Stander</collectorName>
(leaf litter, dry river bed),
<specimenCount id="D9FDFD2BFFD82B2EFE3DF966FE0CF0BE" box="[388,423,1735,1761]" pageId="26" pageNumber="27" type="female">1♀</specimenCount>
(NCA 2021/726; epigyne lost in preparation);
</materialsCitation>
<materialsCitation id="7F933CFFFFD82B2EFC01F966FB38F15B" ID-GBIF-Occurrence="3920719345" collectingDate="2021-01-14" collectingDateMax="2022-03-27" collectingDateMin="2021-01-14" collectorName="R. Booysen &amp; L. Malope &amp; S. Sibisi" country="South Africa" elevation="240" latitude="-30.144716" location="Koeroebees" longLatPrecision="1" longitude="17.70295" municipality="Namaqua National Park" pageId="26" pageNumber="27" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Northern Cape">
<collectorName id="620E5374FFD82B2EFC01F966FC5DF0BE" box="[952,1014,1735,1761]" pageId="26" pageNumber="27">Same</collectorName>
locality,
<date id="BB451062FFD82B2EFBE6F966FB4BF0BE" box="[1119,1248,1735,1761]" pageId="26" pageNumber="27" value="2022-03-27">
<collectingDate id="AB01E98AFFD82B2EFBE6F966FB4BF0BE" box="[1119,1248,1735,1761]" pageId="26" pageNumber="27" value="2022-03-27">27.III.2022</collectingDate>
</date>
, leg.
<collectorName id="620E5374FFD82B2EFAA4F966FA33F0BF" box="[1309,1432,1734,1761]" pageId="26" pageNumber="27">C. Haddad</collectorName>
, R.
<collectorName id="620E5374FFD82B2EFF01F94AFE97F15A" box="[184,316,1771,1797]" pageId="26" pageNumber="27">Booysen, L</collectorName>
. Malope &amp;
<collectorName id="620E5374FFD82B2EFE79F94AFDB5F15B" box="[448,542,1770,1797]" pageId="26" pageNumber="27">S. Sibisi</collectorName>
(leaf litter, dry river bed),
<specimenCount id="D9FDFD2BFFD82B2EFCF0F94AFCC0F15A" box="[841,875,1771,1797]" pageId="26" pageNumber="27" type="male">2♂</specimenCount>
<specimenCount id="D9FDFD2BFFD82B2EFCCAF94AFC3EF15A" box="[883,917,1771,1797]" pageId="26" pageNumber="27" type="female">2♀</specimenCount>
(S.E.M. preparations)
</materialsCitation>
.
</paragraph>
<paragraph id="CF4436A2FFD82B2EFF7EF8AFFAFCF176" blockId="26.[151,1437,1662,2013]" box="[199,1367,1806,1833]" pageId="26" pageNumber="27">
<emphasis id="FD8FEAB0FFD82B2EFF7EF8AFFE94F177" bold="true" box="[199,319,1806,1832]" pageId="26" pageNumber="27">Variation.</emphasis>
Total length: females 2.783.32 (average 2.98, n = 7); males 2.052.98 (average 2.36, n = 4).
</paragraph>
<paragraph id="CF4436A2FFD82B2EFF7EF893FEE6F1E7" blockId="26.[151,1437,1662,2013]" pageId="26" pageNumber="27">
<emphasis id="FD8FEAB0FFD82B2EFF7EF893FE69F113" bold="true" box="[199,450,1842,1869]" pageId="26" pageNumber="27">Habitat and biology.</emphasis>
All of the females collected had the epigyne plugged (
<figureCitation id="57C02A27FFD82B2EFBF1F893FB06F112" box="[1096,1197,1842,1869]" captionStart="FIGURES 114120" captionStartId="22.[151,264,1837,1861]" captionTargetBox="[177,1411,489,1814]" captionTargetId="figure-91@22.[177,1411,489,1814]" captionTargetPageId="22" captionText="FIGURES 114120. Scanning electron micrographs of Leptopilos digitus sp. nov. female (114) and male (115121): 114. Epigyne, ventral view, indicating plug (P); 115. Right palp, prolateral view; 116. Left palp, ventro-retrolateral view; 117. Right palpal patella, retrolateral view, arrowhead indicating lyriform organ; 118. Left palpal tibia, ventral view; 119. Distal end of palpal tegulum, ventro-retrolateral view; 120. Same, enlargement of anterior tegular process and embolus.Abbreviations:ATP— anterior tegular process; E—embolus; MA—median apophysis; P—plug; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; RTP—retrolateral tegular process." figureDoi="http://doi.org/10.5281/zenodo.7141984" httpUri="https://zenodo.org/record/7141984/files/figure.png" pageId="26" pageNumber="27">Fig. 114</figureCitation>
). Recorded from the Succulent Karoo, Nama Karoo and Grassland biomes. At the
<typeStatus id="10408800FFD82B2EFCFBF8F6FCD9F12E" box="[834,882,1879,1905]" pageId="26" pageNumber="27">type</typeStatus>
locality, it was a common species collected in the litter of short shrubs in a dry river bed, with adults far more common in autumn (March) than midsummer (January) or winter (July).
</paragraph>
</subSubSection>
<subSubSection id="87E16529FFD82B2EFF7EF863FC5FF183" box="[199,1012,1986,2013]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="CF4436A2FFD82B2EFF7EF863FC5FF183" blockId="26.[151,1437,1662,2013]" box="[199,1012,1986,2013]" pageId="26" pageNumber="27">
<emphasis id="FD8FEAB0FFD82B2EFF7EF863FECAF183" bold="true" box="[199,353,1986,2012]" pageId="26" pageNumber="27">Distribution.</emphasis>
Widespread in the western half of
<collectingCountry id="B7EC7632FFD82B2EFD5EF863FCDDF183" box="[743,886,1986,2012]" name="South Africa" pageId="26" pageNumber="27">South Africa</collectingCountry>
(
<figureCitation id="57C02A27FFD82B2EFC3CF863FC4CF183" box="[901,999,1986,2012]" captionStart="FIGURE 136" captionStartId="28.[151,250,1184,1208]" captionTargetBox="[177,1411,181,1160]" captionTargetId="figure-18@28.[177,1411,181,1160]" captionTargetPageId="28" captionText="FIGURE 136. Distribution of Leptopilos butleri sp. nov. (circles), L. digitus sp. nov. (triangles) and L. vasivulva sp. nov. (stars) in southern Africa." figureDoi="http://doi.org/10.5281/zenodo.7141997" httpUri="https://zenodo.org/record/7141997/files/figure.png" pageId="26" pageNumber="27">Fig. 136</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>