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<document id="5CE08E9F48D89558337585F4008F7F86" ID-DOI="10.11646/phytotaxa.490.1.10" ID-ISSN="1179-3163" ID-Zenodo-Dep="5755516" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630385928328" checkinUser="marcus" docAuthor="Tan, Yun-Hong, Min, Dao-Zhang, Ding, Hong-Bo, Yang, Bin, Maw, Mya Bhone &amp; Li, Bo" docDate="2021" docId="21298842FFA8346B14CDD9F23176F7EB" docLanguage="en" docName="phytotaxa.490.1.10.pdf" docOrigin="Phytotaxa 490 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.490.1.10" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Premna caridantha Y. H. Tan &amp; Bo Li 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="110" masterDocId="DD10F03AFFA934681445DB463230FFC8" masterDocTitle="Premna caridantha (Lamiaceae: Permnoideae), a distinct new species from Kachin State, northern Myanmar" masterLastPageNumber="113" masterPageNumber="107" pageNumber="108" updateTime="1699255458468" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3D9CBDA68A0CB069ED265394AB97A84C">Premna caridantha (Lamiaceae: Permnoideae), a distinct new species from Kachin State, northern Myanmar</mods:title>
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<mods:namePart id="547EF8991E7968B7801E57E8BF0FB140">Tan, Yun-Hong</mods:namePart>
<mods:nameIdentifier id="2EEAE5C4257263DFEFE8E6EA7F7B92CA" type="ORCID">0000-0001-6238-2743</mods:nameIdentifier>
<mods:affiliation id="A822B1762EE767A1D16BD8BD86C486B6">Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences &amp; Centre for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, Yunnan, China. &amp; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, P. R. China. &amp; tyh @ xtbg. org. cn; https: // orcid. org / 0000 - 0001 - 6238 - 2743</mods:affiliation>
<mods:nameIdentifier id="90F03A552ED7758DD04396C10698517D" type="email">tyh@xtbg.org.cn</mods:nameIdentifier>
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<mods:namePart id="8F7CFAF749D5C1D8C82C6CFFBDC25CAF">Min, Dao-Zhang</mods:namePart>
<mods:nameIdentifier id="DDE8EC86D9FC3242436EA376C9DACE7E" type="ORCID">0000-0002-8779-5592</mods:nameIdentifier>
<mods:affiliation id="A602F550DFEBA7AA1D274E863CA24F2A">Research Center of Ecological Sciences, College of Agronomy, Jiangxi Agricultural University, Nanchang 330045, Jiangxi, China. &amp; mindaozhang @ 163. com; https: // orcid. org / 0000 - 0002 - 8779 - 5592</mods:affiliation>
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<mods:roleTerm id="0ED6DBB7F1B9CB0A1EB4D1B3C9472261">Author</mods:roleTerm>
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<mods:namePart id="16447382CDDAE260ADBF0BBAB1C7CB59">Ding, Hong-Bo</mods:namePart>
<mods:affiliation id="B0BA15722441F983A187BF173F0BAFE2">Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences &amp; Centre for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, Yunnan, China. &amp; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, P. R. China.</mods:affiliation>
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<mods:namePart id="E60457DBDA043823E56272961F7A512D">Yang, Bin</mods:namePart>
<mods:affiliation id="F1803F13DA59F371EF3B604D02840C44">Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences &amp; Centre for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, Yunnan, China. &amp; Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, Menglun, Mengla, Yunnan 666303, P. R. China.</mods:affiliation>
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<mods:namePart id="850F1F4869BF5F0CCAD560C861E73AFD">Maw, Mya Bhone</mods:namePart>
<mods:affiliation id="55100C0D7EF177EBA46AD17253804785">Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences &amp; Centre for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, Yunnan, China. &amp; University of Chinese Academy of Sciences, Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:namePart id="492E83EE23B04D5EDE73F1BEF6D5EBE9">Li, Bo</mods:namePart>
<mods:nameIdentifier id="5105E702B83A6F82DB37DD62CB71788E" type="ORCID">0000-0003-1628-8128</mods:nameIdentifier>
<mods:affiliation id="3467E3198C132276B2F4D8BA069B27CE">Research Center of Ecological Sciences, College of Agronomy, Jiangxi Agricultural University, Nanchang 330045, Jiangxi, China. &amp; hanbolijx @ 163. com; https: // orcid. org / 0000 - 0003 - 1628 - 8128</mods:affiliation>
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<mods:title id="BFF4884E15C256A05CD455272D56D64F">Phytotaxa</mods:title>
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<mods:date id="FA8E2AAD2F7AF2FE7768C117BB3C7FC3">2021</mods:date>
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<mods:number id="D6F423524B8E53233882FE7AE7928BF1">2021-03-10</mods:number>
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<mods:detail id="81555EF7D28BA6BA74C63319630EACDC" type="volume">
<mods:number id="103D56146AC1FC924FA15039413F039F">490</mods:number>
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<mods:url id="AAD51D1E2BD5CF1A9A09333F6D81B156">http://dx.doi.org/10.11646/phytotaxa.490.1.10</mods:url>
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<mods:classification id="18AA43C29F2DFA40ADB540CE9156FF61">journal article</mods:classification>
<mods:identifier id="A5239E45127C7BE0ACB1C15591EFA6FB" type="DOI">10.11646/phytotaxa.490.1.10</mods:identifier>
<mods:identifier id="3DE390AF5D2A30BFBB587167B648C91F" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="949F05D0F85233CD52E6EE065CDBFD46" type="Zenodo-Dep">5755516</mods:identifier>
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<treatment id="21298842FFA8346B14CDD9F23176F7EB" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB" httpUri="http://treatment.plazi.org/id/21298842FFA8346B14CDD9F23176F7EB" lastPageId="3" lastPageNumber="110" pageId="1" pageNumber="108">
<subSubSection id="E19A6ADFFFA8346914CDD9F2313EFD06" box="[136,782,692,719]" pageId="1" pageNumber="108" type="nomenclature">
<paragraph id="A93F3954FFA8346914CDD9F2313EFD06" blockId="1.[136,782,692,719]" box="[136,782,692,719]" pageId="1" pageNumber="108">
<heading id="F2778E38FFA8346914CDD9F2313EFD06" box="[136,782,692,719]" fontSize="11" level="2" pageId="1" pageNumber="108" reason="3">
<taxonomicName id="6E8042D7FFA8346914CDD9F23003FD06" ID-CoL="9V6HR" authority="Y.H.Tan &amp; Bo Li" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[136,563,692,719]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="108" phylum="Tracheophyta" rank="species" species="caridantha" status="sp. nov.">
<emphasis id="9BF4E546FFA8346914CDD9F23359FD07" bold="true" box="[136,361,692,719]" italics="true" pageId="1" pageNumber="108">Premna caridantha</emphasis>
Y.H.Tan &amp; Bo Li
</taxonomicName>
,
<emphasis id="9BF4E546FFA83469167AD9F230A5FD06" bold="true" box="[575,661,692,718]" italics="true" pageId="1" pageNumber="108">
<taxonomicNameLabel id="80C7583DFFA83469167AD9F230A5FD06" box="[575,661,692,718]" pageId="1" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<figureCitation id="31BB25D1FFA8346916E6D9F2313EFD06" box="[675,782,692,718]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[136,229,1915,1937]" captionStartId-1="3.[136,229,1448,1470]" captionTargetBox-0="[151,1398,265,1835]" captionTargetBox-1="[151,1436,190,1423]" captionTargetId-0="figure-24@2.[151,1436,265,1835]" captionTargetId-1="figure-18@3.[151,1436,190,1423]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Premna caridantha Y.H.Tan &amp; Bo Li, sp. nov. A. Flowering branch; B. Spike-like thyrse; C. Flower, lateral view; D. Flower, anterior view; E. Calyx." captionText-1="FIGURE 2. Premna caridantha Y.H.Tan &amp; Bo Li, sp. nov. A. Habit and habitat; B. Flowering branch; C. Adaxial surface of leaf blade; D. Abaxial surface of leaf blade; E. Flower, lateral view; F. Flower, anterior view (the arrow indicates the bright yellowish spot)." figureDoi-0="http://doi.org/10.5281/zenodo.5755518" figureDoi-1="http://doi.org/10.5281/zenodo.5755520" httpUri-0="https://zenodo.org/record/5755518/files/figure.png" httpUri-1="https://zenodo.org/record/5755520/files/figure.png" pageId="1" pageNumber="108">Figs. 1−2</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E19A6ADFFFA8346914CDD9BF379CFCFB" pageId="1" pageNumber="108" type="materials_examined">
<paragraph id="A93F3954FFA8346914CDD9BF379CFCFB" blockId="1.[136,1452,760,819]" pageId="1" pageNumber="108">
<materialsCitation id="19E83309FFA8346914CDD9BF33C6FCFB" collectingDate="2018-06-10" collectionCode="Y, H" country="Myanmar" county="Type" elevation="1400" latitude="27.687252" location="Nawngmun Township" longLatPrecision="1" longitude="97.88694" municipality="Putao District" pageId="1" pageNumber="108" specimenCount="1" stateProvince="Kachin">
<collectingCounty id="405E41D8FFA8346914CDD9BF3289FCC6" box="[136,185,761,782]" pageId="1" pageNumber="108">Type</collectingCounty>
:—
<collectingCountry id="D19779C4FFA834691490D9BF3367FCC6" box="[213,343,761,782]" name="Myanmar" pageId="1" pageNumber="108">MYANMAR</collectingCountry>
.
<collectingRegion id="6B44F7B6FFA834691527D9BF33D2FCC6" box="[354,482,761,783]" country="Myanmar" name="Kachin" pageId="1" pageNumber="108">Kachin State</collectingRegion>
,
<collectingMunicipality id="495BA32EFFA8346915ABD9BF3048FCC7" box="[494,632,761,783]" pageId="1" pageNumber="108">Putao District</collectingMunicipality>
,
<location id="AC5F6F8FFFA8346916C6D9BF316CFCC7" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB:AC5F6F8FFFA8346916C6D9BF316CFCC7" box="[643,860,761,783]" country="Myanmar" county="Type" latitude="27.687252" longLatPrecision="1" longitude="97.88694" municipality="Putao District" name="Nawngmun Township" pageId="1" pageNumber="108" stateProvince="Kachin">Nawngmun Township</location>
, from
<location id="AC5F6F8FFFA8346917DAD9BF31D4FCC7" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB:AC5F6F8FFFA8346917DAD9BF31D4FCC7" box="[927,996,761,783]" country="Myanmar" county="Type" latitude="27.687252" longLatPrecision="1" longitude="97.88694" municipality="Putao District" name="Kawlei" pageId="1" pageNumber="108" stateProvince="Kachin">Kawlei</location>
to
<location id="AC5F6F8FFFA83469104DD9BF365EFCC7" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB:AC5F6F8FFFA83469104DD9BF365EFCC7" box="[1032,1134,761,783]" country="Myanmar" county="Type" latitude="27.687252" longLatPrecision="1" longitude="97.88694" municipality="Putao District" name="Shinsanku" pageId="1" pageNumber="108" stateProvince="Kachin">Shinsanku</location>
,
<geoCoordinate id="CCB45F93FFA83469103FD9BE373CFCC7" box="[1146,1292,760,783]" degrees="27" direction="north" minutes="41" orientation="latitude" pageId="1" pageNumber="108" precision="1" seconds="14.11" value="27.687252">27°41′14.11″N</geoCoordinate>
,
<geoCoordinate id="CCB45F93FFA83469115DD9BE3798FCC7" box="[1304,1448,760,783]" degrees="97" direction="east" minutes="53" orientation="longitude" pageId="1" pageNumber="108" precision="1" seconds="12.99" value="97.88694">97°53′12.99″E</geoCoordinate>
,
<quantity id="6E7894B1FFA8346914F8D85B3336FCFA" box="[189,262,797,818]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="108" unit="m" value="1400.0">
<elevation id="22ADDE67FFA8346914F8D85B3336FCFA" box="[189,262,797,818]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="108" unit="m" value="1400.0">1400 m</elevation>
</quantity>
elev.,
<date id="DD3E1F94FFA834691501D85B33F6FCFA" box="[324,454,797,818]" pageId="1" pageNumber="108" value="2018-06-10">
<collectingDate id="CD7AE67CFFA834691501D85B33F6FCFA" box="[324,454,797,818]" pageId="1" pageNumber="108" value="2018-06-10">10 June 2018</collectingDate>
</date>
,
<emphasis id="9BF4E546FFA834691595D85833C6FCFB" box="[464,502,798,819]" italics="true" pageId="1" pageNumber="108">
<collectionCode id="CF91A191FFA834691595D85833EFFCFB" box="[464,479,798,819]" country="USA" lsid="urn:lsid:biocol.org:col:14792" name="Yale University" pageId="1" pageNumber="108" type="Herbarium">Y</collectionCode>
.
<collectionCode id="CF91A191FFA8346915A7D85833C6FCFB" box="[482,502,798,819]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="108" type="Herbarium">H</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="9BF4E546FFA8346915B3D85833CAFCFB" box="[502,506,798,819]" italics="true" pageId="1" pageNumber="108">.</emphasis>
<materialsCitation id="19E83309FFA8346915BAD8583798FCFB" box="[511,1448,797,819]" collectionCode="RAF" country="Myanmar" location="Tanet" pageId="1" pageNumber="108" specimenCode="M4286" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="9BF4E546FFA8346915BAD85830ACFCFA" box="[511,668,797,819]" italics="true" pageId="1" pageNumber="108">
<location id="AC5F6F8FFFA8346915BAD8583003FCFB" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB:AC5F6F8FFFA8346915BAD8583003FCFB" box="[511,563,798,819]" country="Myanmar" name="Tanet" pageId="1" pageNumber="108" stateProvince="Kachin">Tanet</location>
al.
<specimenCode id="F926912FFFA83469161DD85B30ACFCFA" box="[600,668,797,818]" collectionCode="M" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="108" type="Herbarium">M4286</specimenCode>
</emphasis>
(
<typeStatus id="763B87F6FFA8346916EDD85B30CEFCFB" box="[680,766,797,819]" pageId="1" pageNumber="108" type="holotype">holotype</typeStatus>
HITBC-0035004!, isotypes HITBC-0035005!, CSH-0181311!,
<collectionCode id="CF91A191FFA834691123D85B37ADFCFB" box="[1382,1437,797,819]" country="Myanmar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13034" lsid="urn:lsid:biocol.org:col:13034" name="Forest Research Institute" pageId="1" pageNumber="108" type="Herbarium">RAF</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E19A6ADFFFA8346914CDD81F31FFFBEA" pageId="1" pageNumber="108" type="diagnosis">
<paragraph id="A93F3954FFA8346914CDD81F31FFFBEA" blockId="1.[136,1452,857,2063]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914CDD81F3335FCBB" bold="true" box="[136,261,857,883]" pageId="1" pageNumber="108">Diagnosis:</emphasis>
—
<taxonomicName id="6E8042D7FFA834691564D81C33CDFCBB" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[289,509,857,883]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="108" phylum="Tracheophyta" rank="species" species="caridantha">
<emphasis id="9BF4E546FFA834691564D81C33CDFCBB" box="[289,509,857,883]" italics="true" pageId="1" pageNumber="108">Premna caridantha</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="6E8042D7FFA83469169BD81C318FFCBB" box="[734,959,857,883]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="108" phylum="Tracheophyta" rank="species" species="grandipaniculata">
<emphasis id="9BF4E546FFA83469169BD81C318FFCBB" box="[734,959,857,883]" italics="true" pageId="1" pageNumber="108">P. grandipaniculata</emphasis>
</taxonomicName>
in having large and intricate paniculiform inflorescences which branch three times with the terminal branch formed by a spike-like thyrse, but clearly differs from the latter in leaf shape (oblong to ovate with acuminate to acute apex
<emphasis id="9BF4E546FFA8346917F9D8E631EBFC71" box="[956,987,928,953]" italics="true" pageId="1" pageNumber="108">vs.</emphasis>
broadly ovate to subrounded with sharply acute apex), corolla shape (distinctly 2-lipped with a short incurved tube and a bright yellowish spot on the middle lobe of the lower lip
<emphasis id="9BF4E546FFA834691578D8A3336BFC36" box="[317,347,997,1022]" italics="true" pageId="1" pageNumber="108">vs.</emphasis>
slightly 2-lipped to nearly actinomorphic, straight corolla tube and without a yellowish spot), and stamen orientation (exerted under the upper lip
<emphasis id="9BF4E546FFA8346916D8DF4E308BFBE9" box="[669,699,1032,1057]" italics="true" pageId="1" pageNumber="108">vs.</emphasis>
nearly radially exerted).
</paragraph>
</subSubSection>
<subSubSection id="E19A6ADFFFA8346914F8DF6F3698F984" pageId="1" pageNumber="108" type="description">
<paragraph id="A93F3954FFA8346914F8DF6F36C8F9E1" blockId="1.[136,1452,857,2063]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914F8DF6F3361FB8B" bold="true" box="[189,337,1065,1091]" pageId="1" pageNumber="108">Description:</emphasis>
—Woody climber. Branches brown, terete, spreading, covered by dense white elliptic lenticels, tomentose, with an interpetiolar ridge between petioles; branchlets dark green, glabrous, base surrounded by small bracts. Leaves simple, opposite-decussate, usually unequal in size, glabrous, oblong to ovate, leathery, 9.3–18.8 ×
<quantity id="6E7894B1FFA8346914CDDFD73328FB64" box="[136,280,1169,1196]" metricMagnitude="-2" metricUnit="m" metricValue="9.55" metricValueMax="11.5" metricValueMin="7.6" pageId="1" pageNumber="108" unit="cm" value="9.55" valueMax="11.5" valueMin="7.6">7.6–11.5 cm</quantity>
, apex acuminate to acute, base cuneate to broadly cuneate, margin entire; veins 6–10 pairs, abaxially raised and adaxially slightly compressed, secondary veins curved and jointed near margin; petiole
<quantity id="6E7894B1FFA8346910AEDFF2375AFB07" box="[1259,1386,1204,1231]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.8" metricValueMin="0.4" pageId="1" pageNumber="108" unit="cm" value="1.1" valueMax="1.8" valueMin="0.4">0.4–1.8 cm</quantity>
long, slightly inflated. Inflorescences terminal, large and paniculiform branching three times,
<quantity id="6E7894B1FFA8346910C0DF9136C8FB39" box="[1157,1272,1239,1265]" metricMagnitude="-1" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="1" pageNumber="108" unit="cm" value="19.5" valueMax="23.0" valueMin="16.0">16–23 cm</quantity>
long,
<quantity id="6E7894B1FFA834691103DF91379BFB39" box="[1350,1451,1239,1266]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="1" pageNumber="108" unit="cm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 cm</quantity>
wide, peduncle minutely puberulous but nearly glabrous when mature, terminal branches spike-like thyrses, minutely brownish pubescent,
<quantity id="6E7894B1FFA834691530DE5A33EDFAFF" box="[373,477,1308,1335]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="1" pageNumber="108" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5–7.5 cm</quantity>
long, formed by sessile cymes laxly arranged on the axis; bracts ovate,
<quantity id="6E7894B1FFA834691144DE5A375CFAFF" box="[1281,1388,1308,1335]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="1" pageNumber="108" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 mm</quantity>
long, easily deciduous; bracteoles subulate, tiny. Calyx campanulate,
<quantity id="6E7894B1FFA83469170BDE79318FFA91" box="[846,959,1343,1370]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="108" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long, outside minutely brownish pubescent, strongly 2-lipped, both lips entire with obtuse apex. Corolla yellowish white, sub-glabrous,
<quantity id="6E7894B1FFA834691038DE273733FAB4" box="[1149,1283,1377,1404]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="1" pageNumber="108" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
long, 2-lipped, tube incurved, inside densely white villose around the throat; upper lip straight, deeply emarginate; lower lip 3-lobed, lobes subequal, obovate, strongly reflexed, apex subrounded, middle lobe with a bright yellowish spot near the throat. Stamens 4, subequal, exerted under the upper lip; anther purplish brown. Ovary oblong,
<quantity id="6E7894B1FFA8346910C0DE8F36C9FA2C" box="[1157,1273,1481,1508]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="108" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, glabrous, golden glandular at the top; style white, exerted, slightly longer than stamens, stigma equally 2-cleft. Fruits narrowly obovate, slightly drupaceous, shorter than the calyx, yellowish brown, only one seed fully developed.
</paragraph>
<paragraph id="A93F3954FFA8346914F8DD773698F984" blockId="1.[136,1452,857,2063]" box="[189,1192,1585,1612]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914F8DD773309F983" bold="true" box="[189,313,1585,1611]" pageId="1" pageNumber="108">Phenology</emphasis>
:—Flowering from middle May to late June; fruiting from late June to August.
</paragraph>
</subSubSection>
<subSubSection id="E19A6ADFFFA8346914F8DD1232D1F931" pageId="1" pageNumber="108" type="distribution">
<paragraph id="A93F3954FFA8346914F8DD1232D1F931" blockId="1.[136,1452,857,2063]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914F8DD1233D3F9A6" bold="true" box="[189,483,1620,1646]" pageId="1" pageNumber="108">Distribution and habitat</emphasis>
:—The natural distribution of the species is currently little known because it has been encountered only twice in the field, and we did not find any other specimens in the herbaria consulted. The species was discovered in tropical montane forests on the way from Kawlei to Shinsanku, Nawngmun Township, Putao District,
<collectingRegion id="6B44F7B6FFA8346914CDDDFA332BF91E" box="[136,283,1724,1750]" country="Myanmar" name="Kachin" pageId="1" pageNumber="108">Kachin State</collectingRegion>
, northern
<collectingCountry id="D19779C4FFA8346915CEDDFB33CAF91F" box="[395,506,1725,1751]" name="Myanmar" pageId="1" pageNumber="108">Myanmar</collectingCountry>
, at an elevation of
<quantity id="6E7894B1FFA834691689DDFA3159F91F" box="[716,873,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="108" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">1000–1500 m</quantity>
, and observed as an epiphyte on the tree of
<taxonomicName id="6E8042D7FFA834691114DDFA32EAF931" class="Magnoliopsida" family="Altingiaceae" genus="Altingia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="1" pageNumber="108" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="9BF4E546FFA834691114DDFA32EAF931" italics="true" pageId="1" pageNumber="108">Altingia excelsa</emphasis>
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="E19A6ADFFFA8346B14F8DC473176F7EB" lastPageId="3" lastPageNumber="110" pageId="1" pageNumber="108" type="etymology">
<paragraph id="A93F3954FFA8346914F8DC4730D0F8A9" blockId="1.[136,1452,857,2063]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914F8DC47330EF8D3" bold="true" box="[189,318,1793,1819]" pageId="1" pageNumber="108">Etymology</emphasis>
:—The specific epithet “
<taxonomicName id="6E8042D7FFA834691609DC4730F7F8D3" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[588,711,1793,1819]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="108" phylum="Tracheophyta" rank="species" species="caridantha">
<emphasis id="9BF4E546FFA834691609DC4730F7F8D3" box="[588,711,1793,1819]" italics="true" pageId="1" pageNumber="108">caridantha</emphasis>
</taxonomicName>
” is derived from “
<taxonomicName id="6E8042D7FFA8346917D5DC4731DCF8D3" authorityName="Dana" authorityYear="1852" box="[912,1004,1793,1819]" class="Malacostraca" family="Caridea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="108" phylum="Arthropoda" rank="family">
<emphasis id="9BF4E546FFA8346917D5DC4731DCF8D3" box="[912,1004,1793,1819]" italics="true" pageId="1" pageNumber="108">Caridea</emphasis>
</taxonomicName>
” (commonly known as caridean shrimp) and “
<emphasis id="9BF4E546FFA834691481DC62333FF8F6" box="[196,271,1828,1854]" italics="true" pageId="1" pageNumber="108">anthos</emphasis>
” (means flower), indicating that the flower of the species superficially looks like a shrimp, especially its incurved corolla tube resembles the back of a shrimp.
</paragraph>
<paragraph id="A93F3954FFA8346914F8DC2F317BF7C6" blockId="1.[136,1452,857,2063]" pageId="1" pageNumber="108">
<emphasis id="9BF4E546FFA8346914F8DC2F300AF84B" bold="true" box="[189,570,1897,1923]" pageId="1" pageNumber="108">Preliminary conservation status</emphasis>
:—
<materialsCitation id="19E83309FFA834691619DC2F3177F7C6" collectionCode="DD, IUCN" country="Myanmar" location="Further" municipality="Red List Categories" pageId="1" pageNumber="108" specimenCount="1" stateProvince="Kachin">
This species is only known from a single locality in northern
<collectingCountry id="D19779C4FFA83469114ADC2C374CF84C" box="[1295,1404,1898,1924]" name="Myanmar" pageId="1" pageNumber="108">Myanmar</collectingCountry>
and we have not gotten a clear picture of its natural distribution and/or population status, making no adequate information to assess its risk of extinction directly or indirectly. Hence, we temporarily list the species as Data Deficient (
<collectionCode id="CF91A191FFA83469113DDCE93793F801" box="[1400,1443,1967,1993]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="1" pageNumber="108" type="Herbarium">DD</collectionCode>
) according to the
<collectionCode id="CF91A191FFA83469150EDC9433BFF824" box="[331,399,2002,2028]" pageId="1" pageNumber="108">IUCN</collectionCode>
<collectingMunicipality id="495BA32EFFA8346915DCDC97304DF823" box="[409,637,2001,2027]" pageId="1" pageNumber="108">Red List Categories</collectingMunicipality>
(
<bibRefCitation id="CD1144A5FFA8346916CBDC94312BF824" author="IUCN" box="[654,795,2002,2028]" pageId="1" pageNumber="108" refId="ref3045" refString="IUCN (2012) IUCN Red List Categories and Criteria: Version 3. 1 (2 ndedn). Gland, Switzerland and Cambridge, 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
<location id="AC5F6F8FFFA834691775DC9731B2F823" LSID="urn:lsid:plazi:treatment:21298842FFA8346B14CDD9F23176F7EB:AC5F6F8FFFA834691775DC9731B2F823" box="[816,898,2001,2027]" country="Myanmar" municipality="Red List Categories" name="Further" pageId="1" pageNumber="108" stateProvince="Kachin">Further</location>
field surveys in
<collectingRegion id="6B44F7B6FFA83469100DDC9736EAF824" box="[1096,1242,2001,2028]" country="Myanmar" name="Kachin" pageId="1" pageNumber="108">Kachin State</collectingRegion>
of
<collectingCountry id="D19779C4FFA834691140DC943742F824" box="[1285,1394,2002,2028]" name="Myanmar" pageId="1" pageNumber="108">Myanmar</collectingCountry>
may provide more information on its abundance and/or distribution
</materialsCitation>
.
</paragraph>
<caption id="FDFF69DCFFAB346A14CDDC3D3340F87D" ID-DOI="http://doi.org/10.5281/zenodo.5755518" ID-Zenodo-Dep="5755518" httpUri="https://zenodo.org/record/5755518/files/figure.png" pageId="2" pageNumber="109" startId="2.[136,229,1915,1937]" targetBox="[151,1398,265,1835]" targetPageId="2">
<paragraph id="A93F3954FFAB346A14CDDC3D3340F87D" blockId="2.[136,1452,1915,1973]" pageId="2" pageNumber="109">
<emphasis id="9BF4E546FFAB346A14CDDC3D32CDF858" bold="true" box="[136,253,1915,1937]" pageId="2" pageNumber="109">FIGURE 1.</emphasis>
<taxonomicName id="6E8042D7FFAB346A1546DC3A3058F859" authority="Y.H.Tan &amp; Bo Li" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[259,616,1915,1937]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="109" phylum="Tracheophyta" rank="species" species="caridantha" status="sp. nov.">
<emphasis id="9BF4E546FFAB346A1546DC3A338CF858" box="[259,444,1915,1937]" italics="true" pageId="2" pageNumber="109">Premna caridantha</emphasis>
Y.H.Tan &amp; Bo Li
</taxonomicName>
,
<emphasis id="9BF4E546FFAB346A1637DC3A308BF859" box="[626,699,1916,1937]" italics="true" pageId="2" pageNumber="109">
<taxonomicNameLabel id="80C7583DFFAB346A1637DC3A308BF859" box="[626,699,1916,1937]" pageId="2" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="9BF4E546FFAB346A1685DC3D30E7F858" bold="true" box="[704,727,1915,1936]" pageId="2" pageNumber="109">A.</emphasis>
Flowering branch;
<emphasis id="9BF4E546FFAB346A17D7DC3D3199F858" bold="true" box="[914,937,1915,1936]" pageId="2" pageNumber="109">B.</emphasis>
Spike-like thyrse;
<emphasis id="9BF4E546FFAB346A1018DC3D3645F859" bold="true" box="[1117,1141,1915,1937]" pageId="2" pageNumber="109">C.</emphasis>
Flower, lateral view;
<emphasis id="9BF4E546FFAB346A1100DC3D376DF858" bold="true" box="[1349,1373,1915,1936]" pageId="2" pageNumber="109">D.</emphasis>
Flower, anterior view;
<emphasis id="9BF4E546FFAB346A1551DCD9331AF87C" bold="true" box="[276,298,1951,1972]" pageId="2" pageNumber="109">E.</emphasis>
Calyx.
</paragraph>
</caption>
<caption id="FDFF69DCFFAA346B14CDDEEE3771FA29" ID-DOI="http://doi.org/10.5281/zenodo.5755520" ID-Zenodo-Dep="5755520" httpUri="https://zenodo.org/record/5755520/files/figure.png" pageId="3" pageNumber="110" startId="3.[136,229,1448,1470]" targetBox="[151,1436,190,1423]" targetPageId="3">
<paragraph id="A93F3954FFAA346B14CDDEEE3771FA29" blockId="3.[136,1452,1448,1506]" pageId="3" pageNumber="110">
<emphasis id="9BF4E546FFAA346B14CDDEEE32CDFA75" bold="true" box="[136,253,1448,1470]" pageId="3" pageNumber="110">FIGURE 2.</emphasis>
<taxonomicName id="6E8042D7FFAA346B1547DEEF3056FA76" authority="Y.H.Tan &amp; Bo Li" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[258,614,1448,1470]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="caridantha" status="sp. nov.">
<emphasis id="9BF4E546FFAA346B1547DEEF338BFA75" box="[258,443,1448,1470]" italics="true" pageId="3" pageNumber="110">Premna caridantha</emphasis>
Y.H.Tan &amp; Bo Li
</taxonomicName>
,
<emphasis id="9BF4E546FFAA346B162ADEEF3088FA76" box="[623,696,1449,1470]" italics="true" pageId="3" pageNumber="110">
<taxonomicNameLabel id="80C7583DFFAA346B162ADEEF3088FA76" box="[623,696,1449,1470]" pageId="3" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="9BF4E546FFAA346B16F9DEEE30E4FA75" bold="true" box="[700,724,1448,1469]" pageId="3" pageNumber="110">A.</emphasis>
Habit and habitat;
<emphasis id="9BF4E546FFAA346B17CCDEEE31AFFA75" bold="true" box="[905,927,1448,1469]" pageId="3" pageNumber="110">B.</emphasis>
Flowering branch;
<emphasis id="9BF4E546FFAA346B101CDEEE3641FA76" bold="true" box="[1113,1137,1448,1470]" pageId="3" pageNumber="110">C.</emphasis>
Adaxial surface of leaf blade;
<emphasis id="9BF4E546FFAA346B11D1DEEE379CFA75" bold="true" box="[1428,1452,1448,1469]" pageId="3" pageNumber="110">D.</emphasis>
Abaxial surface of leaf blade;
<emphasis id="9BF4E546FFAA346B15EFDE8A33F0FA29" bold="true" box="[426,448,1484,1505]" pageId="3" pageNumber="110">E.</emphasis>
Flower, lateral view;
<emphasis id="9BF4E546FFAA346B16D7DE8A3094FA29" bold="true" box="[658,676,1484,1505]" pageId="3" pageNumber="110">F.</emphasis>
Flower, anterior view (the arrow indicates the bright yellowish spot).
</paragraph>
</caption>
<paragraph id="A93F3954FFAA346B14F8DD563176F7EB" blockId="3.[136,1452,1552,2083]" pageId="3" pageNumber="110">
<emphasis id="9BF4E546FFAA346B14F8DD563348F9E3" bold="true" box="[189,376,1552,1579]" pageId="3" pageNumber="110">Taxonomic note</emphasis>
:—In
<taxonomicName id="6E8042D7FFAA346B15F2DD57303FF9E2" authorityName="Linnaeus" authorityYear="1771" box="[439,527,1553,1578]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="genus">
<emphasis id="9BF4E546FFAA346B15F2DD57303FF9E2" box="[439,527,1553,1578]" italics="true" pageId="3" pageNumber="110">Premna</emphasis>
</taxonomicName>
, inflorescence structure and corolla shape have been tested to be of great significance for its taxonomy (
<bibRefCitation id="CD1144A5FFAA346B1510DD723385F987" author="Li, B." box="[341,437,1588,1615]" pageId="3" pageNumber="110" refId="ref3117" refString="Li, B. (2009) Phylogenetics of Wenchengia and Premna (Lamiaceae). South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, 93 pp." type="book" year="2009">Li 2009</bibRefCitation>
). There are four basic inflorescence
<typeStatus id="763B87F6FFAA346B1711DD7331BFF987" box="[852,911,1589,1615]" pageId="3" pageNumber="110">types</typeStatus>
observed in
<taxonomicName id="6E8042D7FFAA346B1061DD73364CF986" authorityName="Linnaeus" authorityYear="1771" box="[1060,1148,1589,1614]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="genus">
<emphasis id="9BF4E546FFAA346B1061DD73364CF986" box="[1060,1148,1589,1614]" italics="true" pageId="3" pageNumber="110">Premna</emphasis>
</taxonomicName>
, spike-like thyrses formed by sessile cymes, flat-topped corymbs formed by lax long petiolate cymes, capitula formed by dense short cymes, and conical panicles formed by lax cymes (
<bibRefCitation id="CD1144A5FFAA346B160FDD3A309BF95F" author="Li, B." box="[586,683,1660,1687]" pageId="3" pageNumber="110" refId="ref3117" refString="Li, B. (2009) Phylogenetics of Wenchengia and Premna (Lamiaceae). South China Botanical Garden, Chinese Academy of Sciences, Guangzhou, 93 pp." type="book" year="2009">Li 2009</bibRefCitation>
). Three species having spike-like thyrses were recognized in
<emphasis id="9BF4E546FFAA346B1129DD3A32DBF972" italics="true" pageId="3" pageNumber="110">
Flora of
<collectingCountry id="D19779C4FFAA346B14E2DDE632DBF972" box="[167,235,1696,1722]" name="China" pageId="3" pageNumber="110">China</collectingCountry>
</emphasis>
(
<bibRefCitation id="CD1144A5FFAA346B14B9DDE633C6F973" author="Chen, S. &amp; Gilbert, M. G." box="[252,502,1696,1723]" pageId="3" pageNumber="110" refId="ref2824" refString="Chen, S. L &amp; Gilbert, M. G. (1994) Verbenaceae. In: Wu, C. Y. &amp; Raven, P. H. (Eds.) Flora of China. Vol. 17. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 1 - 49. [http: // www. efloras. org / florataxon. aspx? flora _ id = 2 &amp; taxon _ id = 10941]" type="url" year="1994">Chen &amp; Gilbert 1994</bibRefCitation>
),
<emphasis id="9BF4E546FFAA346B164EDDE73002F972" box="[523,562,1697,1722]" italics="true" pageId="3" pageNumber="110">viz.</emphasis>
,
<taxonomicName id="6E8042D7FFAA346B1604DDE730FAF972" authorityName="Wallich ex Clarke" authorityYear="1885" box="[577,714,1696,1722]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="bracteata">
<emphasis id="9BF4E546FFAA346B1604DDE730FAF972" box="[577,714,1696,1722]" italics="true" pageId="3" pageNumber="110">P. bracteata</emphasis>
</taxonomicName>
,
<taxonomicName id="6E8042D7FFAA346B169EDDE7315AF972" authorityName="Wallich ex Schauer" authorityYear="1847" box="[731,874,1697,1722]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="interrupta">
<emphasis id="9BF4E546FFAA346B169EDDE7315AF972" box="[731,874,1697,1722]" italics="true" pageId="3" pageNumber="110">P. interrupta</emphasis>
</taxonomicName>
, and
<taxonomicName id="6E8042D7FFAA346B17E9DDE7379BF973" authority="Wallich ex Schauer (1847: 633)" authorityName="Schauer" authorityPageNumber="633" authorityYear="1847" box="[940,1451,1696,1723]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis id="9BF4E546FFAA346B17E9DDE73606F972" box="[940,1078,1697,1722]" italics="true" pageId="3" pageNumber="110">P. racemosa</emphasis>
Wallich ex
<bibRefCitation id="CD1144A5FFAA346B1087DDE6379BF973" author="Schauer, J. C." box="[1218,1451,1696,1723]" pageId="3" pageNumber="110" refId="ref3165" refString="Schauer, J. C. (1847) Verbenaceae. In: Candolle, A. D. (Ed.) Prodromus. Vol. 11. Victor Masson, Paris, pp. 522 - 700. https: // doi. org / 10.5962 / bhl. title. 286" type="book" year="1847">Schauer (1847: 633)</bibRefCitation>
</taxonomicName>
which has been reduced to a synonym of
<taxonomicName id="6E8042D7FFAA346B1622DD8330C7F916" authorityName="Wallich ex Schauer" authorityYear="1847" box="[615,759,1733,1758]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="interrupta">
<emphasis id="9BF4E546FFAA346B1622DD8330C7F916" box="[615,759,1733,1758]" italics="true" pageId="3" pageNumber="110">P. interrupta</emphasis>
</taxonomicName>
in
<bibRefCitation id="CD1144A5FFAA346B175BDD823676F917" author="Leeratiwong, C. &amp; Chantaranothai, P. &amp; Paton, A" box="[798,1094,1732,1759]" pageId="3" pageNumber="110" pagination="113 - 142" refId="ref3074" refString="Leeratiwong, C., Chantaranothai, P. &amp; Paton, A (2009) A synopsis of the genus Premna L. (Lamiaceae) in Thailand. The Natural History Journal of Chulalongkorn University 9: 113 - 142." type="journal article" year="2009">
Leeratiwong
<emphasis id="9BF4E546FFAA346B17F0DD8331DFF916" box="[949,1007,1732,1758]" italics="true" pageId="3" pageNumber="110">et al.</emphasis>
(2009)
</bibRefCitation>
. In
<taxonomicName id="6E8042D7FFAA346B1036DD833732F916" authorityName="Wallich ex Schauer" authorityYear="1847" box="[1139,1282,1733,1758]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="interrupta">
<emphasis id="9BF4E546FFAA346B1036DD833732F916" box="[1139,1282,1733,1758]" italics="true" pageId="3" pageNumber="110">P. interrupta</emphasis>
</taxonomicName>
, sessile cymes form a single spike-like thyrse without branches (
<figureCitation id="31BB25D1FFAA346B1685DDAE3116F8CB" box="[704,806,1768,1795]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[191,1395,190,1928]" captionTargetId="figure-24@4.[191,1395,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the leaf size and the structure of inflorescence among the four species having spike-like thyrses. A. Premna caridantha (HITBC-0035005);B.P.grandipaniculata (HITBC-0037465);C.P.bracteata (K-001329005); D. P.interrupta (K-000884640). With permissions from the herbarium of the Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences (HITBC) and the herbarium Royal Botanic Gardens, Kew (K)." figureDoi="http://doi.org/10.5281/zenodo.5755522" httpUri="https://zenodo.org/record/5755522/files/figure.png" pageId="3" pageNumber="110">Figs. 3D</figureCitation>
,
<figureCitation id="31BB25D1FFAA346B1776DDAF31BDF8CB" box="[819,909,1768,1795]" captionStart="FIGURE 4" captionStartId="5.[136,229,1054,1076]" captionTargetBox="[164,1423,190,1030]" captionTargetId="figure-18@5.[164,1423,190,1030]" captionTargetPageId="5" captionText="FIGURE 4. Comparison of inflorescences among Premna caridantha (A, a), P. grandipaniculata (B, b), P. bracteata (C, c) and P. interrupta (D, d)." figureDoi="http://doi.org/10.5281/zenodo.5755524" httpUri="https://zenodo.org/record/5755524/files/figure.png" pageId="3" pageNumber="110">4D &amp; d</figureCitation>
), while in
<taxonomicName id="6E8042D7FFAA346B104FDDAF36A2F8CA" authorityName="Wallich ex Clarke" authorityYear="1885" box="[1034,1170,1768,1794]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="bracteata">
<emphasis id="9BF4E546FFAA346B104FDDAF36A2F8CA" box="[1034,1170,1768,1794]" italics="true" pageId="3" pageNumber="110">P. bracteata</emphasis>
</taxonomicName>
, the inflorescences have 2–4 pairs of secondary branches which are spike-like thyrses (
<figureCitation id="31BB25D1FFAA346B170BDC4A3182F8EF" box="[846,946,1804,1831]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[191,1395,190,1928]" captionTargetId="figure-24@4.[191,1395,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the leaf size and the structure of inflorescence among the four species having spike-like thyrses. A. Premna caridantha (HITBC-0035005);B.P.grandipaniculata (HITBC-0037465);C.P.bracteata (K-001329005); D. P.interrupta (K-000884640). With permissions from the herbarium of the Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences (HITBC) and the herbarium Royal Botanic Gardens, Kew (K)." figureDoi="http://doi.org/10.5281/zenodo.5755522" httpUri="https://zenodo.org/record/5755522/files/figure.png" pageId="3" pageNumber="110">Figs. 3C</figureCitation>
,
<figureCitation id="31BB25D1FFAA346B17FADC4B3626F8EF" box="[959,1046,1805,1831]" captionStart="FIGURE 4" captionStartId="5.[136,229,1054,1076]" captionTargetBox="[164,1423,190,1030]" captionTargetId="figure-18@5.[164,1423,190,1030]" captionTargetPageId="5" captionText="FIGURE 4. Comparison of inflorescences among Premna caridantha (A, a), P. grandipaniculata (B, b), P. bracteata (C, c) and P. interrupta (D, d)." figureDoi="http://doi.org/10.5281/zenodo.5755524" httpUri="https://zenodo.org/record/5755524/files/figure.png" pageId="3" pageNumber="110">4C &amp; c</figureCitation>
). A recently described new species,
<taxonomicName id="6E8042D7FFAA346B14CDDC773005F883" authority="(Tan et al. 2018)" baseAuthorityName="Tan" baseAuthorityYear="2018" box="[136,565,1840,1867]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="grandipaniculata">
<emphasis id="9BF4E546FFAA346B14CDDC773358F882" box="[136,360,1840,1866]" italics="true" pageId="3" pageNumber="110">P. grandipaniculata</emphasis>
(
<bibRefCitation id="CD1144A5FFAA346B153CDC77301DF883" author="Tan, Y. H. &amp; Li, D. R. &amp; Zhou, S. S. &amp; Chen, Y. J. &amp; Bramley, G. L. &amp; Li, B." box="[377,557,1840,1867]" pageId="3" pageNumber="110" pagination="117 - 123" refId="ref3284" refString="Tan, Y. H., Li, D. R., Zhou, S. S., Chen, Y. J., Bramley, G. L. &amp; Li, B. (2018) Premna grandipaniculata (Lamiaceae, Premnoideae), a remarkable new species from north Myanmar. PhytoKeys 94: 117 - 123. https: // doi. org / 10.3897 / phytokeys. 94.22033" type="journal article" year="2018">
Tan
<emphasis id="9BF4E546FFAA346B15EBDC7733D8F882" box="[430,488,1840,1866]" italics="true" pageId="3" pageNumber="110">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
, has spike-like thyrses too, which form an intricate panicle with 4–6 pairs of secondary branches and 1–3 pairs of tertiary branches for each secondary branch (
<figureCitation id="31BB25D1FFAA346B1063DC12364EF8A7" box="[1062,1150,1876,1903]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[191,1395,190,1928]" captionTargetId="figure-24@4.[191,1395,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the leaf size and the structure of inflorescence among the four species having spike-like thyrses. A. Premna caridantha (HITBC-0035005);B.P.grandipaniculata (HITBC-0037465);C.P.bracteata (K-001329005); D. P.interrupta (K-000884640). With permissions from the herbarium of the Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences (HITBC) and the herbarium Royal Botanic Gardens, Kew (K)." figureDoi="http://doi.org/10.5281/zenodo.5755522" httpUri="https://zenodo.org/record/5755522/files/figure.png" pageId="3" pageNumber="110">Fig. 3B</figureCitation>
,
<figureCitation id="31BB25D1FFAA346B10CFDC1336D1F8A7" box="[1162,1249,1876,1903]" captionStart="FIGURE 4" captionStartId="5.[136,229,1054,1076]" captionTargetBox="[164,1423,190,1030]" captionTargetId="figure-18@5.[164,1423,190,1030]" captionTargetPageId="5" captionText="FIGURE 4. Comparison of inflorescences among Premna caridantha (A, a), P. grandipaniculata (B, b), P. bracteata (C, c) and P. interrupta (D, d)." figureDoi="http://doi.org/10.5281/zenodo.5755524" httpUri="https://zenodo.org/record/5755524/files/figure.png" pageId="3" pageNumber="110">4B &amp; b</figureCitation>
). Inflorescence of
<taxonomicName id="6E8042D7FFAA346B14CDDC3F3311F85A" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[136,289,1912,1938]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="caridantha">
<emphasis id="9BF4E546FFAA346B14CDDC3F3311F85A" box="[136,289,1912,1938]" italics="true" pageId="3" pageNumber="110">P. caridantha</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="6E8042D7FFAA346B15BFDC3F30E9F85A" box="[506,729,1912,1938]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="grandipaniculata">
<emphasis id="9BF4E546FFAA346B15BFDC3F30E9F85A" box="[506,729,1912,1938]" italics="true" pageId="3" pageNumber="110">P. grandipaniculata</emphasis>
</taxonomicName>
but the latter is characterized by having large broadly ovate to subrounded leaves with sharply acute apex (
<figureCitation id="31BB25D1FFAA346B1634DCDA30F8F87F" box="[625,712,1948,1975]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[191,1395,190,1928]" captionTargetId="figure-24@4.[191,1395,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of the leaf size and the structure of inflorescence among the four species having spike-like thyrses. A. Premna caridantha (HITBC-0035005);B.P.grandipaniculata (HITBC-0037465);C.P.bracteata (K-001329005); D. P.interrupta (K-000884640). With permissions from the herbarium of the Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences (HITBC) and the herbarium Royal Botanic Gardens, Kew (K)." figureDoi="http://doi.org/10.5281/zenodo.5755522" httpUri="https://zenodo.org/record/5755522/files/figure.png" pageId="3" pageNumber="110">Fig. 3B</figureCitation>
), while
<taxonomicName id="6E8042D7FFAA346B1765DCDB3185F87E" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[800,949,1948,1974]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="caridantha">
<emphasis id="9BF4E546FFAA346B1765DCDB3185F87E" box="[800,949,1948,1974]" italics="true" pageId="3" pageNumber="110">P. caridantha</emphasis>
</taxonomicName>
has oblong to ovate leaves with acuminate to acute apex. Besides, the corolla of
<taxonomicName id="6E8042D7FFAA346B164BDC8730DCF812" box="[526,748,1984,2010]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="grandipaniculata">
<emphasis id="9BF4E546FFAA346B164BDC8730DCF812" box="[526,748,1984,2010]" italics="true" pageId="3" pageNumber="110">P. grandipaniculata</emphasis>
</taxonomicName>
is nearly actinomorphic with extremely short and straight tube but that of
<taxonomicName id="6E8042D7FFAA346B1547DCA333A9F836" authorityName="Y. H. Tan &amp; Bo Li" authorityYear="2021" box="[258,409,2020,2046]" class="Magnoliopsida" family="Lamiaceae" genus="Premna" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="species" species="caridantha">
<emphasis id="9BF4E546FFAA346B1547DCA333A9F836" box="[258,409,2020,2046]" italics="true" pageId="3" pageNumber="110">P. caridantha</emphasis>
</taxonomicName>
is strongly 2-lipped with a short-incurved tube and a bright yellowish spot on the middle lob of the lower lip, which are distinct characteristics in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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