125 lines
16 KiB
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125 lines
16 KiB
XML
<document id="4620CC4D7779FF871A4659AC1D650629" ID-DOI="10.1111/zoj.12196" ID-ISSN="0024-4082" ID-Zenodo-Dep="5330952" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168717398" checkinUser="felipe" docAuthor="Lautenschlager, Stephan & Rauhut, Oliver W. M." docDate="2015" docId="03EA6076FF87440DFC409DA4A3C8FED9" docLanguage="en" docName="zoj12196.pdf" docOrigin="Zoological Journal of the Linnean Society 173 (1)" docSource="http://dx.doi.org/10.1111/zoj.12196" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Rauisuchus tiradentes von Huene 1938" docType="treatment" docVersion="5" lastPageNumber="83" masterDocId="FFD3180EFF9C4413FF9B9B73A227FF8F" masterDocTitle="Osteology of Rauisuchus tiradentes from the Late Triassic (Carnian) Santa Maria Formation of Brazil, and its implications for rauisuchid anatomy and phylogeny" masterLastPageNumber="91" masterPageNumber="55" pageNumber="82" updateTime="1699171848396" updateUser="plazi">
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<mods:title id="9D48513F161652C64FC71FB7ED3FCDB3">Osteology of Rauisuchus tiradentes from the Late Triassic (Carnian) Santa Maria Formation of Brazil, and its implications for rauisuchid anatomy and phylogeny</mods:title>
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<mods:namePart id="034EAF2D0253C429F3C5D2E1D0FA4F6E">Lautenschlager, Stephan</mods:namePart>
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<mods:namePart id="13639AE516DFA46584C8075A3C4B7F1B">Rauhut, Oliver W. M.</mods:namePart>
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<mods:typeOfResource id="7131CDF9303019C43A7B092D649E8553">text</mods:typeOfResource>
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<mods:title id="EE556B0A3678BD9BC513AED245B9EB30">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="C7BC156F617BB11F80D374C331836A83">
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<mods:date id="7C595A8014DA4E099B3D0260DF7C8CBD">2015</mods:date>
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<mods:number id="49E395AE4D58973488A3CB99184CAEB7">2014-12-01</mods:number>
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<mods:number id="5C8D4621CAE99D688F86F5078A0F8378">173</mods:number>
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<mods:url id="AA6A0DD623A858BF53B7346C93FB6036">http://dx.doi.org/10.1111/zoj.12196</mods:url>
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<mods:identifier id="56C037803C8C17F3FC1C37B0D04C74C7" type="DOI">10.1111/zoj.12196</mods:identifier>
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<treatment id="03EA6076FF87440DFC409DA4A3C8FED9" ID-DOI="http://doi.org/10.5281/zenodo.5398699" ID-Zenodo-Dep="5398699" LSID="urn:lsid:plazi:treatment:03EA6076FF87440DFC409DA4A3C8FED9" httpUri="http://treatment.plazi.org/id/03EA6076FF87440DFC409DA4A3C8FED9" lastPageId="30" lastPageNumber="83" pageId="27" pageNumber="82">
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<subSubSection id="C35982EBFF874408FC409DA4A628F89D" pageId="27" pageNumber="82" type="nomenclature">
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<paragraph id="8BFCD160FF874408FC409DA4A727F965" blockId="27.[941,1326,1719,1772]" box="[987,1280,1751,1772]" pageId="27" pageNumber="82">
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<heading id="D0B4660CFF874408FC409DA4A727F965" box="[987,1280,1751,1772]" centered="true" fontSize="9" level="2" pageId="27" pageNumber="82" reason="2">
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<taxonomicName id="4C43AAE3FF874408FC409DA4A727F965" authorityName="von Huene" authorityYear="1938" box="[987,1280,1751,1772]" class="Reptilia" family="Rauisuchidae" genus="Rauisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="27" pageNumber="88" phylum="Chordata" rank="species" species="tiradentes">
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<emphasis id="B9370D72FF874408FC409DA4A727F965" box="[987,1280,1751,1772]" italics="true" pageId="27" pageNumber="82">
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<emphasis id="B9370D72FF874408FC409DA4A649F965" box="[987,1134,1751,1772]" italics="true" pageId="27" pageNumber="82">RAUISUCHUS</emphasis>
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TIRADENTES
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BFCD160FF874408FCA19D8EA628F89D" blockId="27.[826,1441,1789,1903]" box="[826,1039,1789,1810]" pageId="27" pageNumber="82">
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<emphasis id="B9370D72FF874408FCA19D8EA628F89D" box="[826,1039,1789,1810]" italics="true" pageId="27" pageNumber="82">Rauisuchoid ilium</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35982EBFF87440DFCA19C6FA3C8FED9" lastPageId="30" lastPageNumber="85" pageId="27" pageNumber="82" type="description">
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<paragraph id="8BFCD160FF87440FFCA19C6FA2DAFB20" blockId="27.[826,1441,1789,1903]" lastBlockId="28.[144,759,1085,1903]" lastPageId="28" lastPageNumber="83" pageId="27" pageNumber="82">
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<materialsCitation id="3B2BDB3DFF87440FFCA19C6FA0FEFBDC" collectionCode="BSPG" country="American Samoa" lastPageId="28" lastPageNumber="83" location="Find" municipality="Compared" pageId="27" pageNumber="82" specimenCount="1" typeStatus="lectotype">
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The only known ilium (
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<collectionCode id="ED5249A5FF874408FBC99C6FA6BFF8BE" box="[1106,1176,1820,1841]" country="0" httpUri="http://grbio.org/cool/r9n2-w98n" name="Bayerische Staatssammlung fuer Palaeontologie und Geologie" pageId="27" pageNumber="82">BSPG</collectionCode>
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<collectingCountry id="F35491F0FF874408FB3A9C6FA6E5F8BE" box="[1185,1218,1820,1841]" name="American Samoa" pageId="27" pageNumber="82">AS</collectingCountry>
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XXV 88;
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<figureCitation id="1378CDE5FF874408FAA19C6FA16BF8DF" captionStart="Figure 17" captionStartId="29.[164,242,1768,1787]" captionTargetBox="[388,1218,198,1736]" captionTargetId="figure-0@29.[387,1219,195,1738]" captionTargetPageId="29" captionText="Figure 17. Left ilium (BSPG AS XXV 88) of Rauisuchoidea indet. (formerly assigned to Rauisuchus tiradentes) in lateral view (A) and medial view (B). Abbreviations: a.sr, articulation with sacral ribs; ac, acetabulum; app, anterior preacetabular process; d, depression; pep, pelvic process; ppp, posterior postacetabular process; sab, supra-acetabular buttress; sar, supraacetabular rim." figureDoi="http://doi.org/10.5281/zenodo.5370301" httpUri="https://zenodo.org/record/5370301/files/figure.png" pageId="27" pageNumber="82">Fig. 17A, B</figureCitation>
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) originally assigned to
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<taxonomicName id="4C43AAE3FF874408FBCE9C48A6C4F8C0" authorityName="von Huene" authorityYear="1938" box="[1109,1251,1850,1872]" class="Reptilia" family="Rauisuchidae" genus="Rauisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="27" pageNumber="82" phylum="Chordata" rank="species" species="tiradentes">
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<emphasis id="B9370D72FF874408FBCE9C48A6C4F8C0" box="[1109,1251,1850,1872]" italics="true" pageId="27" pageNumber="82">R. tiradentes</emphasis>
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</taxonomicName>
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is a left element and comes from the secondary fossil site as part of ‘
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<location id="8E9C87BBFF874408FAF69C2AA786F8E0" LSID="urn:lsid:plazi:treatment:03EA6076FF87440DFC409DA4A3C8FED9:8E9C87BBFF874408FAF69C2AA786F8E0" box="[1389,1441,1881,1903]" country="American Samoa" municipality="Compared" name="Find" pageId="27" pageNumber="82">Find</location>
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1025’.
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<collectingMunicipality id="6B984B1AFF80440FFF449F4EA371FBDC" box="[223,342,1085,1107]" pageId="28" pageNumber="83">Compared</collectingMunicipality>
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with the pubis of the
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<typeStatus id="54F86FC2FF80440FFDEA9F4EA0FEFBDC" box="[625,729,1085,1107]" pageId="28" pageNumber="83" type="lectotype">lectotype</typeStatus>
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</materialsCitation>
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it shows a notable difference in size and is from an individual that was at least 20–25% smaller than the
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<typeStatus id="54F86FC2FF80440FFF0B9FEAA2DDFB20" box="[144,250,1177,1199]" pageId="28" pageNumber="83" type="lectotype">lectotype</typeStatus>
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.
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</paragraph>
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<caption id="DF3C81E8FF80440FFF0B98DCA115FC72" ID-DOI="http://doi.org/10.5281/zenodo.5370299" ID-Zenodo-Dep="5370299" httpUri="https://zenodo.org/record/5370299/files/figure.png" pageId="28" pageNumber="83" startId="28.[144,220,943,962]" targetBox="[400,1166,197,912]" targetPageId="28">
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<paragraph id="8BFCD160FF80440FFF0B98DCA115FC72" blockId="28.[144,1421,943,1021]" pageId="28" pageNumber="83">
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<emphasis id="B9370D72FF80440FFF0B98DCA327FC4D" bold="true" box="[144,256,943,962]" pageId="28" pageNumber="83">Figure 16.</emphasis>
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Body osteoderms of
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<taxonomicName id="4C43AAE3FF80440FFE4898DCA096FC4D" authorityName="von Huene" authorityYear="1938" box="[467,689,943,962]" class="Reptilia" family="Rauisuchidae" genus="Rauisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="28" pageNumber="83" phylum="Chordata" rank="species" species="tiradentes">
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<emphasis id="B9370D72FF80440FFE4898DCA096FC4D" box="[467,689,943,962]" italics="true" pageId="28" pageNumber="83">Rauisuchus tiradentes</emphasis>
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</taxonomicName>
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. A, osteoderm from the cervical region (BSPG AS XXV 92), B, osteoderm (BSPG AS XXV 97) from the anterior dorsal region, C, osteoderm (BSPG AS XXV 94) from the posterior dorsal region, D, osteoderms (BSPG AS XXV 121c, f) from the caudal region.
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</paragraph>
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</caption>
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<paragraph id="8BFCD160FF80440FFF339FCBA732FA4D" blockId="28.[144,759,1085,1903]" lastBlockId="28.[806,1422,1085,1903]" pageId="28" pageNumber="83">
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The ilium consists of an acetabular area and the iliac blade with two processes: an anterior preacetabular process and a posterior postacetabular process. The lateral surface of the main body is nearly entirely occupied by a deep acetabulum. A prominent supraacetabular rim surrounds the dorsal margin. Anteriorly, the main body possesses a spike-like, dorsally flattened projection, which forms the pubic process. In this region the supra-acetabular rim is strongly developed. Anteroventrally, below the pubic process, the ilium bears a longitudinal groove with a depression on the ventralmost part, forming the articulation socket for the pubis. The parallel acetabular rim is not interrupted, indicating that the pubis only marginally contributed, or did not contribute at all, to the acetabulum. The posteroventral part of the ilium is partly broken. The area where the ischiadic process would be located is missing. The preacetabular process is short and projects anteriorly, but does not extend anterior to the pubic process. It has a blunt outline and is only slightly separated from the main body. The postacetabular process is considerably elongated and projects posteriorly and slightly laterally, although the latter might be a preservational artefact. Its cross section is triangular with a rounded lateral side. The medial(?) side is covered by a ventromedially excavated depression. Posteriorly it tapers to a blunt tip. Above the supra-acetabular rim, located between the preacetabular and the postacetabular process, the ilium possesses a prominent supra-acetabular ridge or buttress. It is perpendicular to the longitudinal axis of the iliac blade and contacts the dorsal margin to the supra-acetabular rim. According to
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<bibRefCitation id="EFD2AC91FF80440FFC229E23A67EFAE9" author="Colbert EH" box="[953,1113,1360,1382]" pageId="28" pageNumber="83" pagination="59 - 78" refId="ref21786" refString="Colbert EH. 1961. The Triassic reptile Poposaurus. Fieldiana Geology 14: 59 - 78." type="journal article" year="1961">Colbert (1961)</bibRefCitation>
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and
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<bibRefCitation id="EFD2AC91FF80440FFB099E23A770FAE9" author="Chatterjee S" box="[1170,1367,1360,1382]" pageId="28" pageNumber="83" pagination="395 - 460" refId="ref21719" refString="Chatterjee S. 1985. Postosuchus, a new thecodontian reptile from the Triassic of Texas and the origin of tyrannosaurs. Philosophical Transactions of the Royal Society, London 309: 395 - 460." type="journal article" year="1985">Chatterjee (1985)</bibRefCitation>
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, this ridge represents the attachment point for the iliofemoralis or the iliotibialis muscle, although this inference has been debated (
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<bibRefCitation id="EFD2AC91FF80440FFBEE9EDFA72EFA4D" author="Gower DJ" box="[1141,1289,1452,1474]" pageId="28" pageNumber="83" pagination="447 - 488" refId="ref22088" refString="Gower DJ. 2000. Rauisuchian archosaurs (Reptilia, Diapsida): an overview. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 218: 447 - 488." type="journal article" year="2000">Gower, 2000</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BFCD160FF80440DFCA49EB8A3C8FED9" blockId="28.[806,1422,1085,1903]" lastBlockId="30.[144,759,198,342]" lastPageId="30" lastPageNumber="85" pageId="28" pageNumber="83">
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On the medial side of the ilium, four distinct depressions are visible, which most probably represent the articulation sockets for the two-pronged sacral ribs of the sacral vertebrae. As no corresponding or overlapping elements are preserved in the
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<typeStatus id="54F86FC2FF80440FFB6A9D36A77DF9D4" box="[1265,1370,1605,1627]" pageId="28" pageNumber="83" type="lectotype">lectotype</typeStatus>
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, the ilium cannot be referred to
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<taxonomicName id="4C43AAE3FF80440FFBC49D17A6C9F9F6" authorityName="von Huene" authorityYear="1938" box="[1119,1262,1636,1657]" class="Reptilia" family="Rauisuchidae" genus="Rauisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="28" pageNumber="83" phylum="Chordata" rank="species" species="tiradentes">
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<emphasis id="B9370D72FF80440FFBC49D17A6C9F9F6" box="[1119,1262,1636,1657]" italics="true" pageId="28" pageNumber="83">R. tiradentes</emphasis>
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</taxonomicName>
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, although the following characteristics support its classification as a rauisuchoid ilium: (1) the ilium contributes to the major part of the acetabulum; (2) presence of a supraacetabular rim; (3) a short preacetabular and an elongated postacetabular process; (4) presence of a supraacetabular ridge connecting the supra-acetabular rim and the dorsal margin of the ilium; (5) closed acetabulum (
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<bibRefCitation id="EFD2AC91FF80440FFCFD9C2AA1DDF8E1" author="Gower DJ" box="[870,1018,1881,1902]" pageId="28" pageNumber="83" pagination="447 - 488" refId="ref22088" refString="Gower DJ. 2000. Rauisuchian archosaurs (Reptilia, Diapsida): an overview. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 218: 447 - 488." type="journal article" year="2000">Gower, 2000</bibRefCitation>
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;
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<bibRefCitation id="EFD2AC91FF80440FFB919C2BA68EF8E0" author="Nesbitt SJ" box="[1034,1193,1880,1903]" pageId="28" pageNumber="83" pagination="1 - 292" refId="ref22862" refString="Nesbitt SJ. 2011. The early evolution of archosaurs: relationships and the origin of major clades. Bulletin of the American Museum of Natural History 352: 1 - 292." type="journal article" year="2011">Nesbitt, 2011</bibRefCitation>
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). The fact that the acetabulum is almost entirely or even entirely placed on the ilium might indicate that it was correctly assigned to
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<taxonomicName id="4C43AAE3FF82440DFF619A77A3A3FE97" authorityName="von Huene" authorityYear="1938" box="[250,388,259,281]" class="Reptilia" family="Rauisuchidae" genus="Rauisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="30" pageNumber="85" phylum="Chordata" rank="species" species="tiradentes">
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<emphasis id="B9370D72FF82440DFF619A77A3A3FE97" box="[250,388,259,281]" italics="true" pageId="30" pageNumber="85">R. tiradentes</emphasis>
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</taxonomicName>
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, as the latter lacks an acetabular facet on the pubis; however, this can only be confirmed by additional material.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |