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<document ID-CLB-Dataset="207919" ID-DOI="http://dx.doi.org/10.3897/zookeys.1135.93800" ID-GBIF-Dataset="47a393b8-52c2-44f0-93f7-3ab89c0b1397" ID-PMC="PMC9836713" ID-Pensoft-Pub="1313-2970-1135-1" ID-Pensoft-UUID="D264615FC98353B58E89E7FD655988C6" ID-PubMed="36761800" ID-ZooBank="0915A6D8A8C54C7A95601D6EF9E14B0F" ModsDocID="1313-2970-1135-1" checkinTime="1670952524096" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Kluge, Nikita J. &amp; Gattolliat, Jean-Luc" docDate="2022" docId="02F6F7AF51A354BCA2AFC79154E564E4" docLanguage="en" docName="ZooKeys 1135: 1-59" docOrigin="ZooKeys 1135" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zookeys.1135.93800" docTitle="Branchiobaetis sabahensis Kaltenbach &amp; Kluge &amp; Gattolliat 2022, comb. nov." docType="treatment" docVersion="5" id="D264615FC98353B58E89E7FD655988C6" lastPageNumber="1" masterDocId="D264615FC98353B58E89E7FD655988C6" masterDocTitle="A widespread new genus of Baetidae (Baetidae, Ephemeroptera) from Southeast Asia" masterLastPageNumber="59" masterPageNumber="1" pageNumber="1" updateTime="1732750236106" updateUser="ExternalLinkService">
<mods:mods id="4A82868E172C3D0337DDA9C05A5474DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="009909FCF37AEACAED81AE075FD92AF4">
<mods:title id="414E45CDE830FABE6D10B9977592FDE8">A widespread new genus of Baetidae (Baetidae, Ephemeroptera) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="A743DC80D12C28714B62550A9DC607A6" type="personal">
<mods:role id="CEEA7F21058DFC53394619D1CD8D6F7C">
<mods:roleTerm id="F189B80DC7CA836AF02D12241FA7C1AD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="47C6DC24C85F5D27DEA2BFD51D28324A">Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier id="B8668E481C2039107272E88D4B1A68D8" type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation id="7C40301565617F69068AE9002E78E77C">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier id="CC7F3237D58F693C6AE61C1D022AFD9A" type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name id="64BB75D91AE33D4BB51FD6546CD67B35" type="personal">
<mods:role id="9E92D7F84442C0D08863905B9EB70775">
<mods:roleTerm id="30AD8EAF237D2C214BFB202761D41220">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="276E37B1BC861C94D79F49FCD139D4F3">Kluge, Nikita J.</mods:namePart>
<mods:nameIdentifier id="81B65699DE4D0E40F53186BD9E1ABF19" type="ORCID">https://orcid.org/0000-0001-9741-7790</mods:nameIdentifier>
<mods:affiliation id="F86A88B41847E169B7B597E403F4F23B">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
</mods:name>
<mods:name id="DBBFE378DDEED0C30F8F64DACF262DE3" type="personal">
<mods:role id="9D66C9D1CFAD06D13467A8852384A32C">
<mods:roleTerm id="931B03DECF06B216752C294802ECAFF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9470BE6B9E510CABF2B90CBEDD56E1F">Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier id="C1A3528E677B6E5EB52D7C505A60586A" type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation id="8605B398C70375FA7FFA09632878042B">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0BE473B8CF518EB299DDF626ABE013DA">text</mods:typeOfResource>
<mods:relatedItem id="A12AE74944142F36B6E2678E5F661408" type="host">
<mods:titleInfo id="3F4C50E63424382593FD8B4ED683E981">
<mods:title id="4DE99E3C7A4A6D54E75996520D1ED483">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="12516897784E17A9E45F13ECC74BD24E">
<mods:date id="95AA05F1BA51B25CC65D15304D285385">2022</mods:date>
<mods:detail id="8FAE86992EA947C49CC19312713ED317" type="pubDate">
<mods:number id="F02332B224CACAFF245241457A02D6AF">2022-12-12</mods:number>
</mods:detail>
<mods:detail id="F5957A1153FD4D17D0EE22FA396046E0" type="volume">
<mods:number id="5BEE5056C0E265734BA06DE3576ADA8A">1135</mods:number>
</mods:detail>
<mods:extent id="11ED9E32770986BD9122AB3A97928634" unit="page">
<mods:start id="10A8C837B21542908D1C9198AA107D4B">1</mods:start>
<mods:end id="648F4AB3403BAF7C86B3C5E7ED565893">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3396115E9E123E53F3368CB35DAE622A">
<mods:url id="908D80F18A024CC44C0078B0D374DCF4">http://dx.doi.org/10.3897/zookeys.1135.93800</mods:url>
</mods:location>
<mods:classification id="37C41FDE93C4ACA854E24E329A7A5A4F">journal article</mods:classification>
<mods:identifier id="E5776AC77ACA2892D89EA6EA5F88D65C" type="DOI">http://dx.doi.org/10.3897/zookeys.1135.93800</mods:identifier>
<mods:identifier id="D2B4A869047BC78F414E5C2E6B0C6050" type="Pensoft-Pub">1313-2970-1135-1</mods:identifier>
<mods:identifier id="09A7801A519833149ACC6194226E8985" type="ZooBank">0915A6D8A8C54C7A95601D6EF9E14B0F</mods:identifier>
<mods:identifier id="AE89AD14A03E2B917D5884B3D397B347" type="Pensoft-UUID">D264615FC98353B58E89E7FD655988C6</mods:identifier>
</mods:mods>
<treatment id="02F6F7AF51A354BCA2AFC79154E564E4" ID-GBIF-Taxon="206232494" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4" httpUri="http://treatment.plazi.org/id/02F6F7AF51A354BCA2AFC79154E564E4" lastPageNumber="1" pageId="0" pageNumber="1" scope_order="Ephemeroptera">
<subSubSection id="080222CB2977B65C8105122B9633A9E0" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="72410F27533043FC870627436377CF20" pageId="0" pageNumber="1">
2.
<taxonomicName id="6F10D3DA46F21770E97668376DE7E9B9" LSID="02F6F7AF-51A3-54BC-A2AF-C79154E564E4" authority="(Mueller-Liebenau, 1984)" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1984" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis" status="comb. nov.">
Branchiobaetis sabahensis (
<normalizedToken id="EFFD1BF93240D527077B87A3E1652090" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1984)
</taxonomicName>
<taxonomicNameLabel id="7E79635DE580BABA882667E763760ED7" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="CEA865B69C4FE36D53336EF7596D2CF1" pageId="0" pageNumber="1" type="description">
<paragraph id="532A6BB7ED2D4C7C582619EBF4B1FC7D" pageId="0" pageNumber="1">
<figureCitation id="970669EC17EFE0D2F234B5AD32A0F738" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Branchiobaetis sabahensis comb. nov., larva a head of male larva b labrum c prosternum and bases of forelegs, front view d labium." figureDoi="10.3897/zookeys.1135.93800.figure12" httpUri="https://binary.pensoft.net/fig/780342" pageId="0" pageNumber="1">Figs 12</figureCitation>
<figureCitation id="4A021CD8926156F74CC7D03EBB0D3F9E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="83B726E75A9658B45CEE9F263A74B004" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="D58F5F92B6AD97492389EDB159F8742A" pageId="0" pageNumber="1">
<taxonomicName id="DC214C5E7A16702BBED06C4C42B349A6" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" baseAuthorityYear="2022" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis" status="comb. nov.">Baetis sabahensis</taxonomicName>
:
<bibRefCitation id="CB70927EC63A2EB0CF0725AD391087BF" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="AE410D04381E3C2B5B9224B414D5A8B0" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: 89; figs 3, 9, 14, 14a.
</paragraph>
</subSubSection>
<subSubSection id="8823D3869502981D5B6C6AB80F9B6515" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="E0858D1B6F41467D1B0B25EB48295307" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="2BC65B39A4EDFD0E648E9EA4D337EFD6" pageId="0" pageNumber="1">
<materialsCitation id="93FE0AB0836E1F993194131BEF8D2C8F" collectingDate="1972-07-14" collectingDateMax="1972-07-16" collectingDateMin="1972-07-14" collectorName="G. F. Edmunds" country="Malaysia" location="Ranau" specimenCount="♂" stateProvince="Sabah">
<collectingCountry id="4594329FC64FA4FAF75A14DFD856A61F" name="Malaysia">Malaysia</collectingCountry>
•
<collectingRegion id="365B770B157B0EB8C51E68446F59523F" country="Malaysia" name="Sabah">Sabah</collectingRegion>
,
<location id="64A337BF55E521397628790A1DBFE61B" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4:64A337BF55E521397628790A1DBFE61B" country="Malaysia" name="Ranau" stateProvince="Sabah">Ranau</location>
;
<collectingDate id="872B8286317EC195D0F01E496356B156" value="1972-07-14" valueMax="1972-07-16" valueMin="1972-07-14">14.-16.VII.1972</collectingDate>
; leg.
<collectorName id="4090CBBA754621B9AF4342BCE69D9F2F">G. F. Edmunds</collectorName>
;
<specimenCount id="6A787F0E6F2C21D64364330ABE91EA7C">♂</specimenCount>
larva on slide; SPbU
</materialsCitation>
•
<materialsCitation id="CE5BEC5FE9E2E43724FB7066EC2A2490" collectingDate="1994-09-04" collectorName="S. C. Kang" country="Malaysia" location="Kundasang" specimenCount="♂" stateProvince="Sabah">
<collectingRegion id="E133246B5D936EA3602A812B3D0D13E6" country="Malaysia" name="Sabah">Sabah</collectingRegion>
,
<location id="E207032B3A521B3A1CA74FC5015C40DB" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4:E207032B3A521B3A1CA74FC5015C40DB" country="Malaysia" name="Kundasang" stateProvince="Sabah">Kundasang</location>
;
<collectingDate id="F1E94BCE87FEBB9D6B80A72E119AAC1B" value="1994-09-04">04.IX.1994</collectingDate>
; leg.
<collectorName id="0EE8BC2DA6BBAE1C269D76DB6B62B612">S. C. Kang</collectorName>
;
<specimenCount id="D0F9C115280FC920C396D4862743212E">♂</specimenCount>
larva on slide; SPbU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1BF5C9221F03A200D6F82C8146F196BA" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="D18ECE0E267D684302C5062D51C573E9" pageId="0" pageNumber="1">
<taxonomicName id="09072387566C1DF406646B49C7CAEB5B" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">B. cf. sabahensis</taxonomicName>
comb. nov. material examined.
</paragraph>
<paragraph id="19AB2CD99FE664A1AC8461058F362C1A" pageId="0" pageNumber="1">
<materialsCitation id="0497C13D3D94B7118AC7F97FBBFA1893" collectingDate="2000-06-16" collectorName="P. Derleth, J. - L. Gattolliat" country="Indonesia" county="River Seturan" latitude="2.9866667" location="Wok" longLatPrecision="21" longitude="116.553055" municipality="Seturan" specimenCount="3" specimenCount-larva="3" stateProvince="East Kalimantan">
<collectingCountry id="D9731A63738F2EE39234760BE31C26ED" name="Indonesia">Indonesia</collectingCountry>
•
<collectingRegion id="4FDA211985109FC39B902D486744AA5C" country="Indonesia" name="Kalimantan Timur">East Kalimantan</collectingRegion>
, Bas. Malinau,
<collectingCounty id="1EAEA4E7033D406594799E9EA9F5958A">River Seturan</collectingCounty>
, loc.
<collectingMunicipality id="5C42E1D119D2D370E5B0F963F9EC1A65">Seturan</collectingMunicipality>
(2000-block 44-45), trib.
<location id="3B2772796199FBD5AA74EE4D259257ED" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4:3B2772796199FBD5AA74EE4D259257ED" country="Indonesia" county="River Seturan" latitude="2.9866667" longLatPrecision="21" longitude="116.553055" municipality="Seturan" name="Wok" stateProvince="East Kalimantan">Wok</location>
(Sungai Guang);
<geoCoordinate id="F50346433A91E3139AD6932E410BE1CD" degrees="2" direction="north" minutes="59" orientation="latitude" precision="15" seconds="12" value="2.9866667">2°59'12&quot;N</geoCoordinate>
,
<geoCoordinate id="2AE86AA0540C8BB7730F09DCA3199510" degrees="116" direction="east" minutes="33" orientation="longitude" precision="15" seconds="11" value="116.553055">116°33'11&quot;E</geoCoordinate>
;
<collectingDate id="4CF0D451910765F182D62B553964CE44" value="2000-06-16">16.VI.2000</collectingDate>
; leg.
<collectorName id="6F71CF971AF14404AF1C933136042CC3">P. Derleth</collectorName>
&amp;
<collectorName id="4B862C5DFCA40354184FF359390735F1">J.-L. Gattolliat</collectorName>
;
<specimenCount id="E9F0622BFB6CAD34E75C5CDE53FDE700" type="larva">3 larvae</specimenCount>
on slides; GBIFCH00592470, GBIFCH00592471, GBIFCH00592495; larva in alcohol; GBIFCH00270724; MZL
</materialsCitation>
•
<materialsCitation id="1456215D0E788C476E948009FE33BA3C" collectingDate="2000-08-19" collectorName="P. Derleth, R. Schlaepfer" country="Indonesia" county="River Seturan" latitude="2.9894445" location="Bengahau" longLatPrecision="21" longitude="116.51278" municipality="Seturan" specimenCount="1" stateProvince="East Kalimantan">
<collectingRegion id="34EB51857EBD33A3D889410DD26FEBD5" country="Indonesia" name="Kalimantan Timur">East Kalimantan</collectingRegion>
, Bas. Malinau,
<collectingCounty id="3513DEC42A5558EA765A440846EEED5D">River Seturan</collectingCounty>
, loc.
<collectingMunicipality id="D8910B44BFD9BE2BCFB0020ADAC69A02">Seturan</collectingMunicipality>
(2001-block 57), trib.
<location id="953A63AC8AEEC5644DC981F8C5EC5E1C" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4:953A63AC8AEEC5644DC981F8C5EC5E1C" country="Indonesia" county="River Seturan" latitude="2.9894445" longLatPrecision="21" longitude="116.51278" municipality="Seturan" name="Bengahau" stateProvince="East Kalimantan">Bengahau</location>
;
<geoCoordinate id="C7461B8735CC7335F77F57A677D0124A" degrees="02" direction="north" minutes="59" orientation="latitude" precision="15" seconds="22" value="2.9894445">02°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="9D93069AABA9C9BAD4F88D0FD64E556E" degrees="116" direction="east" minutes="30" orientation="longitude" precision="15" seconds="46" value="116.51278">116°30'46&quot;E</geoCoordinate>
;
<collectingDate id="35E1A0C33F44B1FF0A775386F743F250" value="2000-08-19">19.VIII.2000</collectingDate>
; leg.
<collectorName id="15C64DF22A43A4CAF53C2ABB809FF53A">P. Derleth</collectorName>
&amp;
<collectorName id="DFFBD05A3EE4EAB88D8375EAF4E208E0">R. Schlaepfer</collectorName>
; larva on slide; GBIFCH00592494; larva in alcohol; GBIFCH00270710; MZL
</materialsCitation>
•
<materialsCitation id="C27AED406AE1F64D9D2ED38C0AC72A20" collectingDate="2001-04-11" collectorName="P. Derleth, B. Feldmeyer" country="Indonesia" county="Seturan" latitude="2.9894445" location="Benganau" longLatPrecision="21" longitude="116.51278" municipality="Seturan" specimenCount="1" stateProvince="East Kalimantan">
<collectingRegion id="DC1D5E81C2964B809D55759103DC6A06" country="Indonesia" name="Kalimantan Timur">East Kalimantan</collectingRegion>
, Bas. Malinau, riv.
<collectingCounty id="24DD37DB3762BE188811654B76CB0440">Seturan</collectingCounty>
, loc.
<collectingMunicipality id="EE7051F43BDEB1EFAE9F9D5E3F286E1D">Seturan</collectingMunicipality>
(2001-block 57), trib.
<location id="CDBB51D10E8CC5010C9DF723323A8B49" LSID="urn:lsid:plazi:treatment:02F6F7AF51A354BCA2AFC79154E564E4:CDBB51D10E8CC5010C9DF723323A8B49" country="Indonesia" county="Seturan" latitude="2.9894445" longLatPrecision="21" longitude="116.51278" municipality="Seturan" name="Benganau" stateProvince="East Kalimantan">Benganau</location>
;
<geoCoordinate id="1CA9FCAEA492E2DF0E390352A5066387" degrees="02" direction="north" minutes="59" orientation="latitude" precision="15" seconds="22" value="2.9894445">02°59'22&quot;N</geoCoordinate>
,
<geoCoordinate id="7DF31B368A3C5DD03705183DABF2F2BE" degrees="116" direction="east" minutes="30" orientation="longitude" precision="15" seconds="46" value="116.51278">116°30'46&quot;E</geoCoordinate>
;
<collectingDate id="AD74A5F3384ADF3F1C7A2910C8A96714" value="2001-04-11">11.IV.2001</collectingDate>
; leg.
<collectorName id="E61FCEA4ED0D3220AC5AF7A07AF224F9">P. Derleth</collectorName>
&amp;
<collectorName id="AE7DD9147082B7B1FB616854743D886D">B. Feldmeyer</collectorName>
; larva in alcohol; GBIFCH00270710; MZL
</materialsCitation>
.
</paragraph>
<caption id="CD91E74B9889BD5F28BE8D94886C90D1" doi="10.3897/zookeys.1135.93800.figure12" httpUri="https://binary.pensoft.net/fig/780342" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph id="B16E0D0BA30CEA92D5AA3ACE54B85E20" pageId="0" pageNumber="1">
<emphasis id="E910835D6C9C44C39FDD8E6196E986D8" bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName id="3F885850C5E902B08A0CC47C2B3B3C55" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1984" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="56DC3AA233F24352D2864B395F3D0C8D" italics="true" pageId="0" pageNumber="1">Branchiobaetis sabahensis</emphasis>
</taxonomicName>
comb. nov., larva
<emphasis id="A941A2AF9033D6BC88AB856EC54F9BE0" bold="true" pageId="0" pageNumber="1">a</emphasis>
head of male larva
<emphasis id="930EBA780DEE3DECB21F1D91249FF006" bold="true" pageId="0" pageNumber="1">b</emphasis>
labrum
<emphasis id="3A2C10CDA08031D57212D08B11C23C24" bold="true" pageId="0" pageNumber="1">c</emphasis>
prosternum and bases of forelegs, front view
<emphasis id="AF1E007CB31F6F54B6DAAE7F64CDC742" bold="true" pageId="0" pageNumber="1">d</emphasis>
labium.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="18EA275E7A55C0F085ADA2894761FD6D" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="87A3375095ACA775A3E45DB829189E34" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="A552B165E34053CC8D2FEC45BF9F5A15" pageId="0" pageNumber="1">
<emphasis id="5E3C62F8E135B77B344964D0F42A496A" bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters distinguish
<taxonomicName id="4F1E4401678D01E61D2FA24B838305E4" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="0C950E73BE36ED9472F097A67906A7A2" italics="true" pageId="0" pageNumber="1">B. sabahensis</emphasis>
</taxonomicName>
comb. nov. from other species of
<taxonomicName id="D836397A3D9185792AF404D4A2C5AF76" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="807A3D84D6DF7B491DEA9398018B61DA" italics="true" pageId="0" pageNumber="1">Branchiobaetis</emphasis>
</taxonomicName>
gen. nov.: A) labial palp segment II with short, thumb-like protuberance; segment III rather short and wide, ca. 0.5
<normalizedToken id="3A4A8E4044C1D8202838CF12BA3419C8" originalValue="×">x</normalizedToken>
length of segment II, ca. 0.8
<normalizedToken id="2A6FF4B34B38F229AD4515B3B15A63BD" originalValue="×">x</normalizedToken>
as long as width at base, ca. 0.7
<normalizedToken id="CB0CA8F8C33E0E3CF1C4DEE405127EE2" originalValue="×">x</normalizedToken>
as long as maximal width (Fig.
<figureCitation id="64FA259BEBEB2F24B425364F11914009" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Branchiobaetis sabahensis comb. nov., larva a head of male larva b labrum c prosternum and bases of forelegs, front view d labium." figureDoi="10.3897/zookeys.1135.93800.figure12" httpUri="https://binary.pensoft.net/fig/780342" pageId="0" pageNumber="1">12d</figureCitation>
;
<bibRefCitation id="26B843185E05CE9CF52F382B490E546D" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="B171CE2D6234DA3DD190FCB36A9A76CA" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: fig. 3b); B) incisor of right mandible with ventral denticle (
<bibRefCitation id="D22B9FC8A6079F3B7D8FFA8E90212CF1" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="47DCB5CFDE579BBE83F4C1C9D2A87108" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: fig. 3e; C) dorsal margin of femur with row of ca. 15 long, spine-like setae; no additional row of short setae along margin; no short, stout setae on surface of femur (Fig.
<figureCitation id="39FD465C7F60A7E7D71F4BD0BB17C89B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">13a</figureCitation>
;
<bibRefCitation id="D944BCF98D9536CB7D9CEC12E2F95AE6" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="264A972DCB2318F883E5958ED0A3C15A" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: fig. 3k); D) posterior margin of tergite I smooth, without spines; posterior margins of tergites II-III with rounded or triangular spines, posterior margins of tergites IV-X with triangular spines (Fig.
<figureCitation id="04C098CCC07C05FAD27C9C67BA9789C3" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">13c</figureCitation>
;
<bibRefCitation id="CD70D2820D44313EAA0D9CE59A95EB59" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="DF657354C32E558A8999DC82D5DBFD26" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: fig. 14); posterior margins of sternites: I-V smooth, without spines; VI smooth, without spines, or with some small, triangular spines; VII-IX with triangular or blunt spines (Fig.
<figureCitation id="D62B8D2C0EFE923CE425BF1A08CE39D8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">13d</figureCitation>
).
</paragraph>
<caption id="0DBD9E72B7B44EA191C097CE1B336064" doi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph id="292D0B14FE9DCC71C1F6A09CF0A76BE1" pageId="0" pageNumber="1">
<emphasis id="F9557E222C62D3C6D8795AFEB7311501" bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName id="6AA52AAE7E92573A3527044D76677122" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1984" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="FAF988BDC025799A58AD707BEA566E3E" italics="true" pageId="0" pageNumber="1">Branchiobaetis sabahensis</emphasis>
</taxonomicName>
comb. nov., larva
<emphasis id="B22068C460E2DFEEF6920FFEFC051A99" bold="true" pageId="0" pageNumber="1">a</emphasis>
apex of hind femur
<emphasis id="2D551267153E702C67EF845AB200D17A" bold="true" pageId="0" pageNumber="1">b</emphasis>
paraproct
<emphasis id="9B63A9BC05133786561B372D1D8E53C3" bold="true" pageId="0" pageNumber="1">c</emphasis>
abdominal tergum IX
<emphasis id="778E65D751C8C8BCCAEF9B996EA2CACC" bold="true" pageId="0" pageNumber="1">d</emphasis>
abdominal sternum VIII
<emphasis id="2FFD88602174CAB4045D51B071D82B65" bold="true" pageId="0" pageNumber="1">e</emphasis>
tergalius.
</paragraph>
</caption>
<paragraph id="30D0E098EDDDD5DC76A432D6780D9935" pageId="0" pageNumber="1">
<emphasis id="EEC1E04A5870C6D9A363EFEE416C02DC" bold="true" pageId="0" pageNumber="1">Imago.</emphasis>
Winged stages unknown. Judging from details revealed in last instar larva, turbinate eyes not narrowed; hind wing with costal projection; sterno-styligeral muscle present and thin.
</paragraph>
</subSubSection>
<subSubSection id="83CED5D8B4973384D497A47F7B2440FC" pageId="0" pageNumber="1" type="comments">
<paragraph id="937D60C06F3B9B69E1C92862680A7260" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="77B5C03BD7572129E4FAD7AF3DD887EC" pageId="0" pageNumber="1">
The original description of
<taxonomicName id="FADB105F28C868EEC2FC17F230DD3229" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" baseAuthorityYear="2022" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="2A532DE090EF16822C18FEF499CEF31D" italics="true" pageId="0" pageNumber="1">Baetis sabahensis</emphasis>
</taxonomicName>
<normalizedToken id="E3ED888AA0E72010B48C68155FD6A84F" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
, 1984 was based on larvae, and certain similarities of this species with
<taxonomicName id="BA4D5A2A3326853BBA6DD33F278610FA" class="Insecta" kingdom="Animalia" lsidName="B. javanicus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="891F7361F4315B05B7B66210A75FC9C4" italics="true" pageId="0" pageNumber="1">B. javanicus</emphasis>
</taxonomicName>
were reported. The single argument to separate
<taxonomicName id="919606288697F70C78F0CF72B806C04F" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="936387B016514A606B2C981E086FF19D" italics="true" pageId="0" pageNumber="1">B. sabahensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4AAAE1DF35DFC79AFF91B1DE4FCFEC94" class="Insecta" kingdom="Animalia" lsidName="B. javanicus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="A3FC41D7B65A9804FADCEB446306A415" italics="true" pageId="0" pageNumber="1">B. javanicus</emphasis>
</taxonomicName>
, was the statement that &quot;
<taxonomicName id="4FAC7ACE4A4FE3E0A8A4C59879377614" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" baseAuthorityYear="2022" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="6AD23C275FC351C0A9AE04356401E348" italics="true" pageId="0" pageNumber="1">Baetis sabahensis</emphasis>
</taxonomicName>
sp. nov. is the only species in the genus with coxal gills on the first pair of legs&quot; (
<bibRefCitation id="EC45D05850E99DB895124672532FE517" author="Mueller-Liebenau, I" journalOrPublisher="Spixiana" pageId="0" pageNumber="1" refId="B42" refString="Mueller-Liebenau, I, 1984b. Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85-89." title="Baetidae from Sabah (East Malaysia) (Ephemeroptera). In: Landa V, Soldan T, Tonner M (Eds) Proceedings of the Fourth International Conference on Ephemeroptera, Czechoslovak Academy of Sciences, Budejovice, 85 - 89." year="1984 b">
<normalizedToken id="D6C777DD8C7426871D9CB5092D478F74" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1984b
</bibRefCitation>
: 92). Larva of
<taxonomicName id="F44B1D3D722395CD21978097EF458E9E" class="Insecta" kingdom="Animalia" lsidName="B. javanicus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="6D93BCC93F87E6D22B17DDFF18D16C4A" italics="true" pageId="0" pageNumber="1">B. javanicus</emphasis>
</taxonomicName>
was described and figured by the same author earlier (
<bibRefCitation id="33DB6962E42EE48F0AB7A23A5FC71A32" author="Mueller-Liebenau, I" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="1" pagination="197 - 208" refId="B40" refString="Mueller-Liebenau, I, 1981. Review of the original materiel of the baetid genera Baetis and Pseudocloeon from the Sunda Islands and the Philippines described by G. Ulmer, with some general remarks (Insecta: Ephemeroptera). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 78: 197 - 208" title="Review of the original materiel of the baetid genera Baetis and Pseudocloeon from the Sunda Islands and the Philippines described by G. Ulmer, with some general remarks (Insecta: Ephemeroptera)." volume="78" year="1981">
<normalizedToken id="A4E8D97E44E2831F1BB59B957CD741BB" originalValue="Müller-Liebenau">Mueller-Liebenau</normalizedToken>
1981
</bibRefCitation>
), but the coxal and maxillary gills had not been reported (see above).
</paragraph>
<paragraph id="7B6487574B03A53FB992028737FA4A5B" pageId="0" pageNumber="1">
Larva of
<taxonomicName id="3E346216CC391937BF552371AD9CA44A" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1984" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="E9AE853E2F3937307536C241FE1C92EB" italics="true" pageId="0" pageNumber="1">Branchiobaetis sabahensis</emphasis>
</taxonomicName>
comb. nov. can be separated from
<taxonomicName id="8E6CCB5131E40A28602D67930A0BC1E3" class="Insecta" kingdom="Animalia" lsidName="B. javanicus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="865348D6346D8403652B3B5B3D191FDA" italics="true" pageId="0" pageNumber="1">B. javanicus</emphasis>
</taxonomicName>
comb. nov. by the following characters: A) dense spines on abdominal sternite VIII (Fig.
<figureCitation id="F4F9B513068B9A00AE176737EEA19C0B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">13d</figureCitation>
); B) presence of only one or two stout setae on paraproct, or complete absence (Fig.
<figureCitation id="028D2D0F4643330BB4A15F46723F4FA7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Branchiobaetis sabahensis comb. nov., larva a apex of hind femur b paraproct c abdominal tergum IX d abdominal sternum VIII e tergalius." figureDoi="10.3897/zookeys.1135.93800.figure13" httpUri="https://binary.pensoft.net/fig/780343" pageId="0" pageNumber="1">13b</figureCitation>
); C) incisor of right mandible with ventral denticle; D) labial palp segment III short and wide (Fig.
<figureCitation id="7E3E8B9059D3683F21DE93E236BDBB49" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Branchiobaetis sabahensis comb. nov., larva a head of male larva b labrum c prosternum and bases of forelegs, front view d labium." figureDoi="10.3897/zookeys.1135.93800.figure12" httpUri="https://binary.pensoft.net/fig/780342" pageId="0" pageNumber="1">12d</figureCitation>
; see above).
</paragraph>
<paragraph id="6518B5E154003CEF1861FCF02E40419E" pageId="0" pageNumber="1">
Judging by precursors of turbinate eyes developed in last instar male larva, male imago of
<taxonomicName id="53702F10B65597D31A73F6F441373F56" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="60EA9BA9B1BAC6F141D08BCDF8A5DC37" italics="true" pageId="0" pageNumber="1">B. sabahensis</emphasis>
</taxonomicName>
comb. nov. should differ from
<taxonomicName id="CBE0CE4CF1DDBD72E3307C621E7B7DDF" class="Insecta" kingdom="Animalia" lsidName="B. javanicus" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="054C22A1F249B88C8539F2087823E26B" italics="true" pageId="0" pageNumber="1">B. javanicus</emphasis>
</taxonomicName>
comb. nov. by usual (not narrowed) turbinate eyes (Fig.
<figureCitation id="9C4FD53870EE3D21480BE2C64700B7D8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Branchiobaetis sabahensis comb. nov., larva a head of male larva b labrum c prosternum and bases of forelegs, front view d labium." figureDoi="10.3897/zookeys.1135.93800.figure12" httpUri="https://binary.pensoft.net/fig/780342" pageId="0" pageNumber="1">12a</figureCitation>
).
</paragraph>
<paragraph id="B305E80FF4D80EC8BB3579F4A53C7574" pageId="0" pageNumber="1">
<taxonomicName id="491EE04BB29F45F3D74EF3AC6E96BF27" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2022" baseAuthorityName="Mueller-Liebenau" baseAuthorityYear="1984" class="Insecta" family="Baetidae" genus="Branchiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Branchiobaetis sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">Branchiobaetis cf. sabahensis</taxonomicName>
comb. nov. Specimens from Indonesia (Kalimantan) always have a series of stout setae along posterior margin of paraproct, contrary to specimens from Malaysia (Sabah). As there are no other differentiating characters to
<taxonomicName id="9F6D333A33708A3E214119BECAD316A3" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
<emphasis id="FD4050E59A16913BB9EF9FB73F47F677" italics="true" pageId="0" pageNumber="1">B. sabahensis</emphasis>
</taxonomicName>
comb. nov. from Malaysia (Sabah), we treat this population as
<taxonomicName id="6971F6E9DC3D0B8C0C67B812F2284C14" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">B. cf. sabahensis</taxonomicName>
comb. nov., waiting for genetic investigation of fresh material in the future.
</paragraph>
</subSubSection>
<subSubSection id="759570B54CA6E8480467C8D32C8BC1F8" pageId="0" pageNumber="1" type="distribution">
<paragraph id="843C0269779CEE31511A3FC02D8E88FE" pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph id="31AEF082A90E1BFA1BA88B1D2144119D" pageId="0" pageNumber="1">
<emphasis id="FD3431B25DF9BBF7645925203FFC8201" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="43BB7B388C2CFAEB7268B3ED3ADDB7D2" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution maps a Branchiobaetis gen. nov. in Southeast Asia b Known species of Branchiobaetis gen. nov." figureDoi="10.3897/zookeys.1135.93800.figure27" httpUri="https://binary.pensoft.net/fig/780357" pageId="0" pageNumber="1">27b</figureCitation>
).
</emphasis>
Malaysia (Borneo: Sabah);
<taxonomicName id="5D0CE6C8BC6606A329D89C4197FF64D5" class="Insecta" kingdom="Animalia" lsidName="B. sabahensis" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">B. cf. sabahensis</taxonomicName>
comb. nov. Indonesia (Borneo: Kalimantan).
</paragraph>
</subSubSection>
</treatment>
</document>
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