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<document id="A4FB967952F8B7A8121A87030D1E5DAF" ID-DOI="10.5281/zenodo.6119025" ID-ISSN="0035-418X" ID-Zenodo-Dep="6119025" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645037204323" checkinUser="carolina" docAuthor="Schwendinger, Peter J." docDate="2013" docId="D22287FFFFBA6D65FF58FCBC618DFCC6" docLanguage="en" docName="RevSuisseZool.120.4.585-663.pdf.imf" docOrigin="Revue suisse de Zoologie 120 (4)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Perania nasicornis Schwendinger 1994" docType="treatment" docVersion="3" lastPageNumber="603" masterDocId="2E1BFF87FFB56D77FFCFF862616EFFC3" masterDocTitle="A taxonomic revision of the spider genus Perania Thorell, 1890 (Araneae: Tetrablemmidae: Pacullinae) with the descriptions of eight new species" masterLastPageNumber="663" masterPageNumber="585" pageNumber="600" updateTime="1699297313336" updateUser="plazi">
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<mods:title id="D55EB86A1741D1569ABBC4DB5E61FE6C">A taxonomic revision of the spider genus Perania Thorell, 1890 (Araneae: Tetrablemmidae: Pacullinae) with the descriptions of eight new species</mods:title>
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<mods:namePart id="E0DEE823328647FA4B180AC07CCF78E0">Schwendinger, Peter J.</mods:namePart>
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<subSubSection id="12916562FFBA6D78FF58FCBC655EFB36" box="[151,1072,1245,1270]" pageId="15" pageNumber="600" type="nomenclature">
<paragraph id="5A3436E9FFBA6D78FF58FCBC655EFB36" blockId="15.[151,1072,1245,1270]" box="[151,1072,1245,1270]" pageId="15" pageNumber="600">
<heading id="017C8185FFBA6D78FF58FCBC655EFB36" box="[151,1072,1245,1270]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="600" reason="2">
<taxonomicName id="9D8B4D6AFFBA6D78FF58FCBC6324FB36" ID-CoL="76RLW" authority="Schwendinger, 1994" authorityName="Schwendinger" authorityYear="1994" box="[151,586,1245,1270]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFBA6D78FF58FCBC6006FB35" bold="true" box="[151,360,1246,1270]" italics="true" pageId="15" pageNumber="600">Perania nasicornis</emphasis>
<bibRefCitation id="3E1A4B18FFBA6D78FEA0FCBF6324FB36" author="SCHWENDINGER, P. J." box="[367,586,1245,1269]" pageId="15" pageNumber="600" pagination="447 - 464" refId="ref43375" refString="SCHWENDINGER, P. J. 1994. Four new Perania (Araneae: Tetrablemmidae, Pacullinae) from Thailand and Malaysia. Revue suisse de Zoologie 101 (2): 447 - 464." type="journal article" year="1994">Schwendinger, 1994</bibRefCitation>
</taxonomicName>
Fig. 2M-P
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12916562FFBA6D78FF58FD6060A9FAF3" pageId="15" pageNumber="600" type="reference_group">
<paragraph id="5A3436E9FFBA6D78FF58FD6060A9FAF3" blockId="15.[151,1073,1281,1328]" pageId="15" pageNumber="600">
<treatmentCitationGroup id="7A9B11C7FFBA6D78FF58FD6060A9FAF3" pageId="15" pageNumber="600">
<treatmentCitation id="DB2A10F8FFBA6D78FF58FD606318FAD4" author="SCHWENDINGER, P. J." box="[151,630,1281,1303]" page="452" pageId="15" pageNumber="600" year="1994">
<taxonomicName id="9D8B4D6AFFBA6D78FF58FD606318FAD4" ID-CoL="76RLW" authority="Schwendinger, 1994: 448 - 452" authorityName="Schwendinger" authorityPageNumber="448 - 452" authorityYear="1994" box="[151,630,1281,1303]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFBA6D78FF58FD606021FAD4" box="[151,335,1282,1303]" italics="true" pageId="15" pageNumber="600">Perania nasicornis</emphasis>
<bibRefCitation id="3E1A4B18FFBA6D78FE9AFD636318FAD4" author="SCHWENDINGER, P. J." box="[341,630,1281,1303]" pageId="15" pageNumber="600" pagination="447 - 464" refId="ref43375" refString="SCHWENDINGER, P. J. 1994. Four new Perania (Araneae: Tetrablemmidae, Pacullinae) from Thailand and Malaysia. Revue suisse de Zoologie 101 (2): 447 - 464." type="journal article" year="1994">Schwendinger, 1994: 448-452</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1-14 (description of male and female).
<bibRefCitation id="3E1A4B18FFBA6D78FF2CFD786000FAF3" author="PLATNICK, N. I." box="[227,366,1306,1328]" pageId="15" pageNumber="600" refId="ref43177" refString="PLATNICK, N. I. 2013. The world spider catalog, version 14. 0. American Museum of Natural History, online at http: // research. amnh. org / entomology / spiders / catalog / index. html DOI: 10. 5531 / db. iz. 0001." type="book" year="2013">Platnick, 2013</bibRefCitation>
(listing).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="12916562FFBA6D78FF2CFD5D624BFA66" pageId="15" pageNumber="600" type="materials_examined">
<paragraph id="5A3436E9FFBA6D78FF2CFD5D623EFAAE" blockId="15.[151,1074,1341,1445]" pageId="15" pageNumber="600">
<materialsCitation id="EAE33CB4FFBA6D78FF2CFD5D623EFAAE" collectingDate="1992-02-16" collectionCode="MHNG" collectorName="P. J. Schwendinger." country="Thailand" elevation="1250" latitude="18.489723" location="Doi Khuntan" longLatPrecision="21" longitude="99.29861" municipality="Mae Tha District" pageId="15" pageNumber="600" specimenCount="1" stateProvince="Lamphun" typeStatus="holotype">
<typeStatus id="8530884BFFBA6D78FF2CFD5D6027FA90" box="[227,329,1343,1364]" pageId="15" pageNumber="600" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="3C9AAE2CFFBA6D78FE99FD5C60F1FA97" box="[342,415,1342,1364]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="600" type="Museum">MHNG</collectionCode>
; 1 Ƌ;
<collectingCountry id="229C7679FFBA6D78FE10FD5C635AFA97" box="[479,564,1342,1364]" name="Thailand" pageId="15" pageNumber="600">Thailand</collectingCountry>
,
<collectingRegion id="984FF80BFFBA6D78FD8EFD5C6396FA97" box="[577,760,1342,1364]" country="Thailand" name="Lamphun" pageId="15" pageNumber="600">Lamphun Province</collectingRegion>
,
<collectingMunicipality id="BA50AC93FFBA6D78FCCBFD5D62C6FA97" box="[772,936,1342,1364]" pageId="15" pageNumber="600">Mae Tha District</collectingMunicipality>
,
<location id="5F546032FFBA6D78FC7BFD5C655CFA97" LSID="urn:lsid:plazi:treatment:D22287FFFFBA6D65FF58FCBC618DFCC6:5F546032FFBA6D78FC7BFD5C655CFA97" box="[948,1074,1342,1364]" country="Thailand" latitude="18.489723" longLatPrecision="21" longitude="99.29861" municipality="Mae Tha District" name="Doi Khuntan" pageId="15" pageNumber="600" stateProvince="Lamphun">Doi Khuntan</location>
(
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,
<geoCoordinate id="3FBF502EFFBA6D78FEEEFD3560FBFAAE" box="[289,405,1367,1389]" degrees="99" direction="east" minutes="17" orientation="longitude" pageId="15" pageNumber="600" precision="15" seconds="55" value="99.29861">99°1755”E</geoCoordinate>
),
<quantity id="9D739B0CFFBA6D78FE66FD35609CFAAE" box="[425,498,1367,1389]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="600" unit="m" value="1250.0">
<elevation id="D1A6D1DAFFBA6D78FE66FD35609CFAAE" box="[425,498,1367,1389]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="600" unit="m" value="1250.0">1250 m</elevation>
</quantity>
;
<date id="2E351029FFBA6D78FE30FD35630DFAAE" box="[511,611,1367,1389]" pageId="15" pageNumber="600" value="1992-02-16">
<collectingDate id="3E71E9C1FFBA6D78FE30FD35630DFAAE" box="[511,611,1367,1389]" pageId="15" pageNumber="600" value="1992-02-16">16.II.1992</collectingDate>
</date>
; leg
<collectorName id="F77E533FFFBA6D78FD5AFD3A623EFAAE" box="[661,848,1367,1389]" pageId="15" pageNumber="600">P. J. Schwendinger.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="5A3436E9FFBA6D78FF2CFD15624BFA66" blockId="15.[151,1074,1341,1445]" pageId="15" pageNumber="600">
<materialsCitation id="EAE33CB4FFBA6D78FF2CFD15624EFA66" collectingDate="6. II. 1992, 8. X. 1992" collectionCode="MHNG" collectorName="P. J. Schwendinger. No" elevation="1150" pageId="15" pageNumber="600" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<typeStatus id="8530884BFFBA6D78FF2CFD156020FA48" box="[227,334,1399,1420]" pageId="15" pageNumber="600" type="paratype">PARATYPES</typeStatus>
:
<collectionCode id="3C9AAE2CFFBA6D78FEADFD1460C5FA4F" box="[354,427,1398,1420]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="600" type="Museum">MHNG</collectionCode>
; 6 Ƌ,
<specimenCount id="4C8DFD60FFBA6D78FDCFFD146343FA4F" box="[512,557,1396,1420]" pageId="15" pageNumber="600" type="female">4 ♀</specimenCount>
; from the type locality,
<quantity id="9D739B0CFFBA6D78FCF7FD1462AEFA4F" box="[824,960,1398,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="15" pageNumber="600" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">
<elevation id="D1A6D1DAFFBA6D78FCF7FD1462AEFA4F" box="[824,960,1398,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="15" pageNumber="600" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000-1300 m</elevation>
</quantity>
;
<collectingDate id="3E71E9C1FFBA6D78FC1CFD14619FFA66" isEnumeration="true" pageId="15" pageNumber="600">
<date id="2E351029FFBA6D78FC1CFD146545FA4F" box="[979,1067,1398,1420]" pageId="15" pageNumber="600" value="1992-02-06">6.II.1992</date>
,
<date id="2E351029FFBA6D78FF58FDED619FFA66" box="[151,241,1423,1445]" pageId="15" pageNumber="600" value="1992-10-08">8.X.1992</date>
</collectingDate>
, leg.
<collectorName id="F77E533FFFBA6D78FEE8FDF26368FA66" box="[295,518,1423,1445]" pageId="15" pageNumber="600">P. J. Schwendinger. No</collectorName>
additional material available
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12916562FFBA6D67FF2CFDD462B4FE58" lastPageId="16" lastPageNumber="601" pageId="15" pageNumber="600" type="diagnosis">
<paragraph id="5A3436E9FFBA6D78FF2CFDD463D5FA0E" blockId="15.[227,699,1461,1485]" box="[227,699,1461,1485]" pageId="15" pageNumber="600">
DIAGNOSIS: See
<bibRefCitation id="3E1A4B18FFBA6D78FE59FDD763DAFA0E" author="SCHWENDINGER, P. J." box="[406,692,1461,1485]" pageId="15" pageNumber="600" pagination="447 - 464" refId="ref43375" refString="SCHWENDINGER, P. J. 1994. Four new Perania (Araneae: Tetrablemmidae, Pacullinae) from Thailand and Malaysia. Revue suisse de Zoologie 101 (2): 447 - 464." type="journal article" year="1994">Schwendinger (1994: 448)</bibRefCitation>
.
</paragraph>
<paragraph id="5A3436E9FFBA6D67FF2CFD8362B4FE58" blockId="15.[151,1074,1504,1656]" lastBlockId="16.[136,1058,227,411]" lastPageId="16" lastPageNumber="601" pageId="15" pageNumber="600">
NEW CHARACTERS: Anterior opisthosomal plate of females fragmented into microplates of different sizes. Postgenital plate in all males completely fused to posterior margin of pulmonary plate, with a sclerotised suture linking them; all microplates on posterior side of opisthosoma of males close to each other but not fused into horizontal strap-like plates. Anterior pair of ventrolateral plates of males largely free, only posterior tips fused with pulmonary plate. Preanal plate of females fragmented into microplates, some of the posterior ones slightly wider than the others. Apex of embolus distinctly split, with widely rounded subterminal lamella overlapping pointed embolic part with a quite large, lobate dorsal projection (Fig. 2M-N). Vulva with posterior dorsal margin of anterior collar very widely rounded (unlike in
<taxonomicName id="9D8B4D6AFFA56D67FC44F906657CFEB8" baseAuthorityName="Thorell" baseAuthorityYear="1898" box="[907,1042,355,379]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA56D67FC44F906657CFEB8" box="[907,1042,355,379]" italics="true" pageId="16" pageNumber="601">P. birmanica</emphasis>
</taxonomicName>
); spermathecae with lateral sides distinctly bulged in posterior portion (Fig. 2O).
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D67FF1BF9CC6343FB46" pageId="16" pageNumber="601" type="discussion">
<paragraph id="5A3436E9FFA56D67FF1BF9CC6343FB46" blockId="16.[136,1059,429,1157]" pageId="16" pageNumber="601">
REMARKS: The porepatches of all spermathecae are much larger than illustrated in the original decription (Fig. 2O-P cf.
<bibRefCitation id="3E1A4B18FFA56D67FDE1F9AF6266FE26" author="SCHWENDINGER, P. J." box="[558,776,461,485]" pageId="16" pageNumber="601" pagination="447 - 464" refId="ref43375" refString="SCHWENDINGER, P. J. 1994. Four new Perania (Araneae: Tetrablemmidae, Pacullinae) from Thailand and Malaysia. Revue suisse de Zoologie 101 (2): 447 - 464." type="journal article" year="1994">Schwendinger, 1994</bibRefCitation>
: figs 13-14), extending on the lateral sides down to the posterior corners of the spermathecae (but usually not visible in ventral view because hidden behind the bulged posterolateral corners). This and the following two new observations on
<taxonomicName id="9D8B4D6AFFA56D67FDBDFA4D63B9FD85" authorityName="Schwendinger" authorityYear="1989" box="[626,727,559,582]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FDBDFA4D63B9FD85" box="[626,727,559,582]" italics="true" pageId="16" pageNumber="601">P. nasuta</emphasis>
</taxonomicName>
make the distinction between females of
<taxonomicName id="9D8B4D6AFFA56D67FEC3FA2D60F2FDA6" authorityName="Schwendinger" authorityYear="1994" box="[268,412,590,614]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FEC3FA2D60F2FDA6" box="[268,412,590,614]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="9D8B4D6AFFA56D67FE11FA2D6328FDA5" authorityName="Schwendinger" authorityYear="1989" box="[478,582,591,614]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FE11FA2D6328FDA5" box="[478,582,591,614]" italics="true" pageId="16" pageNumber="601">P. nasuta</emphasis>
</taxonomicName>
more difficult than given in the original description. Not all
<taxonomicName id="9D8B4D6AFFA56D67FEADFA0D60A8FD45" authorityName="Schwendinger" authorityYear="1989" box="[354,454,623,646]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FEADFA0D60A8FD45" box="[354,454,623,646]" italics="true" pageId="16" pageNumber="601">P. nasuta</emphasis>
</taxonomicName>
females have a longer anterior collar of the vulva than females of
<taxonomicName id="9D8B4D6AFFA56D67FEC6FAED60F9FD66" authorityName="Schwendinger" authorityYear="1994" box="[265,407,654,678]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FEC6FAED60F9FD66" box="[265,407,654,678]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
(Fig. 2K cf. Fig. 2O).
<taxonomicName id="9D8B4D6AFFA56D67FD68FAEC623EFD65" authorityName="Schwendinger" authorityYear="1989" box="[679,848,654,678]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FD68FAEC623EFD65" box="[679,848,654,678]" italics="true" pageId="16" pageNumber="601">Perania nasuta</emphasis>
</taxonomicName>
females from Doi Suthep-Pui have spermathecae that are posteriorly slightly wider than anteriorly (Fig. 2K). Thus females of both species can be more reliably distinguished by the following:
<taxonomicName id="9D8B4D6AFFA56D67FECEFA8C60C6FCC5" authorityName="Schwendinger" authorityYear="1989" box="[257,424,750,774]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FECEFA8C60C6FCC5" box="[257,424,750,774]" italics="true" pageId="16" pageNumber="601">Perania nasuta</emphasis>
</taxonomicName>
with more or less distinctly elevated conical microplates behind dorsal scutum of opisthosoma (microplates not elevated in
<taxonomicName id="9D8B4D6AFFA56D67FC4CFB6D657CFCE6" authorityName="Schwendinger" authorityYear="1994" box="[899,1042,782,806]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FC4CFB6D657CFCE6" box="[899,1042,782,806]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
); spermathecae posteriorly narrower or slightly wider than anteriorly (distinctly wider in
<taxonomicName id="9D8B4D6AFFA56D67FF47FB2D6075FCA6" authorityName="Schwendinger" authorityYear="1994" box="[136,283,846,870]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FF47FB2D6075FCA6" box="[136,283,846,870]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
), their lateral margins not bulging outwards in posterior portion (distinctly bulged in
<taxonomicName id="9D8B4D6AFFA56D67FEA7FB0D609CFC46" authorityName="Schwendinger" authorityYear="1994" box="[360,498,878,902]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FEA7FB0D609CFC46" box="[360,498,878,902]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
), pores visible along most of lateral sides in ventral view (pores hidden by bulged posterolateral corner in ventral view in
<taxonomicName id="9D8B4D6AFFA56D67FC42FBED6577FC66" authorityName="Schwendinger" authorityYear="1994" box="[909,1049,910,934]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FC42FBED6577FC66" box="[909,1049,910,934]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
) (Fig. 2K cf. Fig. 2O). Differences were also found in the sizes of the fragments (microplates) of the anterior opisthosomal plates of females: in
<taxonomicName id="9D8B4D6AFFA56D67FD24FBAD621BFC26" authorityName="Schwendinger" authorityYear="1994" box="[747,885,974,998]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasicornis">
<emphasis id="68FFEAFBFFA56D67FD24FBAD621BFC26" box="[747,885,974,998]" italics="true" pageId="16" pageNumber="601">P. nasicornis</emphasis>
</taxonomicName>
two lateral, one central and all dorsal fragments (corresponding to the upper margin of the plate) are clearly larger than the remaining microplates; in
<taxonomicName id="9D8B4D6AFFA56D67FD09FC6D6240FBE5" authorityName="Schwendinger" authorityYear="1989" box="[710,814,1039,1062]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="68FFEAFBFFA56D67FD09FC6D6240FBE5" box="[710,814,1039,1062]" italics="true" pageId="16" pageNumber="601">P. nasuta</emphasis>
</taxonomicName>
only the two lateral fragments are enlarged. However, the latter differences are not pronounced and cannot be used as a reliable distinction between females of both species. They therefore have not been used in the key to the species.
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D67FF1BFCFB6067FB13" pageId="16" pageNumber="601" type="distribution">
<paragraph id="5A3436E9FFA56D67FF1BFCFB6067FB13" blockId="16.[136,1059,1176,1232]" pageId="16" pageNumber="601">
DISTRIBUTION: Known only from a mountain in northern
<collectingCountry id="229C7679FFA56D67FCAAFCFA62AAFB73" box="[869,964,1176,1200]" name="Thailand" pageId="16" pageNumber="601">Thailand</collectingCountry>
(Fig. 1, locality 11).
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D67FF47FC91654CFA9D" pageId="16" pageNumber="601" type="reference_group">
<paragraph id="5A3436E9FFA56D67FF47FC91654DFA86" blockId="16.[136,1059,1267,1400]" pageId="16" pageNumber="601">
<treatmentCitationGroup id="7A9B11C7FFA56D67FF47FC91654DFA86" pageId="16" pageNumber="601">
<taxonomicName id="9D8B4D6AFFA56D67FF47FC91636CFAC8" ID-CoL="6V3LV" authority="(Thorell, 1898)" baseAuthorityName="Thorell" baseAuthorityYear="1898" box="[136,514,1267,1291]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA56D67FF47FC916036FAC8" bold="true" box="[136,344,1267,1291]" italics="true" pageId="16" pageNumber="601">Perania birmanica</emphasis>
(
<bibRefCitation id="3E1A4B18FFA56D67FEA7FC916097FAC8" author="THORELL" box="[360,505,1267,1291]" pageId="16" pageNumber="601" pagination="271 - 378" refId="ref43628" refString="THORELL, 1898. Viaggio di Leonardo Fea in Birmania e regioni vicine. 80. Secondo saggio sui ragni birmani. 2. Retitelariae et Orbitelariae. Annali del Museo civico di Storia naturale di Genova (2) 19 [= 39]: 271 - 378." type="journal article" year="1898">Thorell, 1898</bibRefCitation>
)
</taxonomicName>
Fig. 3
<treatmentCitation id="DB2A10F8FFA56D67FF47FD74634DFAEF" author="THORELL" box="[136,547,1302,1324]" page="283" pageId="16" pageNumber="601" year="1898">
<taxonomicName id="9D8B4D6AFFA56D67FF47FD74634DFAEF" authority="Thorell, 1898: 282 - 283" authorityName="Thorell" authorityPageNumber="282 - 283" authorityYear="1898" box="[136,547,1302,1324]" class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA56D67FF47FD746052FAEF" box="[136,316,1302,1324]" italics="true" pageId="16" pageNumber="601">Paculla birmanica</emphasis>
<bibRefCitation id="3E1A4B18FFA56D67FE8DFD74634DFAEF" author="THORELL" box="[322,547,1302,1324]" pageId="16" pageNumber="601" pagination="271 - 378" refId="ref43628" refString="THORELL, 1898. Viaggio di Leonardo Fea in Birmania e regioni vicine. 80. Secondo saggio sui ragni birmani. 2. Retitelariae et Orbitelariae. Annali del Museo civico di Storia naturale di Genova (2) 19 [= 39]: 271 - 378." type="journal article" year="1898">Thorell, 1898: 282-283</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(description of
<specimenCount id="4C8DFD60FFA56D67FD72FD7663BFFAEF" box="[701,721,1300,1324]" pageId="16" pageNumber="601" type="female"></specimenCount>
).
<taxonomicName id="9D8B4D6AFFA56D67FF47FD52608FFA86" ID-CoL="6V3LV" authority="(Thorell, 1898)" baseAuthorityName="Thorell" baseAuthorityYear="1898" box="[136,481,1327,1349]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA56D67FF47FD52602CFA86" box="[136,322,1327,1349]" italics="true" pageId="16" pageNumber="601">Perania birmanica</emphasis>
(
<bibRefCitation id="3E1A4B18FFA56D67FE9CFD4D60B7FA86" author="THORELL" box="[339,473,1327,1349]" pageId="16" pageNumber="601" pagination="271 - 378" refId="ref43628" refString="THORELL, 1898. Viaggio di Leonardo Fea in Birmania e regioni vicine. 80. Secondo saggio sui ragni birmani. 2. Retitelariae et Orbitelariae. Annali del Museo civico di Storia naturale di Genova (2) 19 [= 39]: 271 - 378." type="journal article" year="1898">Thorell, 1898</bibRefCitation>
)
</taxonomicName>
.
<treatmentCitation id="DB2A10F8FFA56D67FDCAFD526385FA86" author="BOURNE, J. D." box="[517,747,1327,1349]" page="256" pageId="16" pageNumber="601" year="1980">
<bibRefCitation id="3E1A4B18FFA56D67FDCAFD526385FA86" author="BOURNE, J. D." box="[517,747,1327,1349]" pageId="16" pageNumber="601" pagination="249 - 260" refId="ref42857" refString="BOURNE, J. D. 1980. Revision of Thorell's type species of the family Pacullidae (Araneae). Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot; di Genova 83: 249 - 260." type="journal article" year="1980">Bourne, 1980: 256-259</bibRefCitation>
</treatmentCitation>
, figs 20-23 (re-description and
</treatmentCitationGroup>
</paragraph>
<paragraph id="5A3436E9FFA56D67FF1BFD2B654CFA9D" blockId="16.[136,1059,1267,1400]" box="[212,1058,1353,1375]" pageId="16" pageNumber="601">
illustration of
<typeStatus id="8530884BFFA56D67FE94FD2B60C1FA9C" box="[347,431,1353,1375]" pageId="16" pageNumber="601" type="holotype">holotype</typeStatus>
; transfer);
<bibRefCitation id="3E1A4B18FFA56D67FDD6FD2B63A0FA9C" author="LEHTINEN, P. T." box="[537,718,1353,1375]" pageId="16" pageNumber="601" pagination="1 - 151" refId="ref43059" refString="LEHTINEN, P. T. 1981. Spiders of the Oriental-Australian region. III. Tetrablemmidae, with a world revision. Acta Zoological Fennica 162: 1 - 151." type="journal article" year="1981">Lehtinen, 1981: 16</bibRefCitation>
(mention; transfer);
<bibRefCitation id="3E1A4B18FFA56D67FC5BFD2B654CFA9D" author="SCHWENDINGER, P. J." box="[916,1058,1353,1375]" pageId="16" pageNumber="601" pagination="571 - 582" refId="ref43337" refString="SCHWENDINGER, P. J. 1989. On three new armoured spiders (Araneae: Tetrablemmidae, Pacullinae) from Indonesia and Thailand. Revue suisse de Zoologie 96 (3): 571 - 582." type="journal article" year="1989">Schwendinger,</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D67FF1BFD0062BFFABB" box="[212,977,1378,1400]" pageId="16" pageNumber="601" type="description">
<paragraph id="5A3436E9FFA56D67FF1BFD0062BFFABB" blockId="16.[136,1059,1267,1400]" box="[212,977,1378,1400]" pageId="16" pageNumber="601">
1989: fig. 21 (illustration of outlines of spermathecae);
<bibRefCitation id="3E1A4B18FFA56D67FD22FD006216FABB" author="PLATNICK, N. I." box="[749,888,1378,1400]" pageId="16" pageNumber="601" refId="ref43177" refString="PLATNICK, N. I. 2013. The world spider catalog, version 14. 0. American Museum of Natural History, online at http: // research. amnh. org / entomology / spiders / catalog / index. html DOI: 10. 5531 / db. iz. 0001." type="book" year="2013">Platnick, 2013</bibRefCitation>
(listing).
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D67FF1BFDE560DCFA0D" pageId="16" pageNumber="601" type="materials_examined">
<paragraph id="5A3436E9FFA56D67FF1BFDE560DCFA0D" blockId="16.[136,1059,1412,1487]" pageId="16" pageNumber="601">
<materialsCitation id="EAE33CB4FFA56D67FF1BFDE560DCFA0D" collectionCode="MCSN" collectorName="Leonardo Fea." country="Myanmar" county="Bia-po" latitude="19.083334" location="Carin Ceba" longLatPrecision="1273" longitude="96.583336" municipality="Carin Cheba" pageId="16" pageNumber="601" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="8530884BFFA56D67FF1BFDE56054FA58" box="[212,314,1415,1436]" pageId="16" pageNumber="601" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="3C9AAE2CFFA56D67FE83FDE460FEFA5F" box="[332,400,1414,1436]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="16" pageNumber="601">MCSN</collectionCode>
;
<specimenCount id="4C8DFD60FFA56D67FE6DFDE660D8FA5F" box="[418,438,1412,1436]" pageId="16" pageNumber="601" type="female"></specimenCount>
; “
<collectingCountry id="229C7679FFA56D67FE1CFDE46342FA5F" box="[467,556,1414,1436]" name="Myanmar" pageId="16" pageNumber="601">Birmania</collectingCountry>
,
<location id="5F546032FFA56D67FDF2FDE463C1FA5F" LSID="urn:lsid:plazi:treatment:D22287FFFFBA6D65FF58FCBC618DFCC6:5F546032FFA56D67FDF2FDE463C1FA5F" box="[573,687,1414,1436]" country="Myanmar" county="Bia-po" latitude="19.083334" longLatPrecision="1273" longitude="96.583336" municipality="Carin Cheba" name="Carin Ceba" pageId="16" pageNumber="601">Carin Ceba</location>
,
<location id="5F546032FFA56D67FD0FFDE46397FA5F" LSID="urn:lsid:plazi:treatment:D22287FFFFBA6D65FF58FCBC618DFCC6:5F546032FFA56D67FD0FFDE46397FA5F" box="[704,761,1414,1436]" country="Myanmar" county="Bia-po" latitude="19.083334" longLatPrecision="1273" longitude="96.583336" municipality="Carin Cheba" name="Biapo" pageId="16" pageNumber="601">Biapó</location>
” [on original labels], “
<collectingMunicipality id="BA50AC93FFA56D67FC22FDE461ABFA76" pageId="16" pageNumber="601">Carin Chebà</collectingMunicipality>
” or “
<collectingCounty id="B3554E65FFA56D67FEC6FDFD6024FA76" box="[265,330,1439,1461]" pageId="16" pageNumber="601">Bia-pó</collectingCounty>
” mountains [in original description], approximally
<geoCoordinate id="3FBF502EFFA56D67FCAAFDFD62D6FA76" box="[869,952,1439,1461]" degrees="19" direction="north" minutes="05" orientation="latitude" pageId="16" pageNumber="601" precision="925" value="19.083334">19°05N</geoCoordinate>
,
<geoCoordinate id="3FBF502EFFA56D67FC04FDFD6572FA76" box="[971,1052,1439,1461]" degrees="96" direction="east" minutes="35" orientation="longitude" pageId="16" pageNumber="601" precision="925" value="96.583336">96°35E</geoCoordinate>
; 1885-1889; leg.
<collectorName id="F77E533FFFA56D67FEE8FDDB60DCFA0D" box="[295,434,1465,1487]" pageId="16" pageNumber="601">Leonardo Fea.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA56D66FF1BFD8063EFFA8D" lastPageId="17" lastPageNumber="602" pageId="16" pageNumber="601" type="diagnosis">
<paragraph id="5A3436E9FFA56D67FF1BFD80637DF9BA" blockId="16.[136,1059,1505,1657]" pageId="16" pageNumber="601">
DIAGNOSIS: Female
<typeStatus id="8530884BFFA56D67FE7AFD83637CFA3A" box="[437,530,1505,1529]" pageId="16" pageNumber="601" type="holotype">holotype</typeStatus>
(male unknown) distinguished from females of other congeners by a dorsal opisthosomal scutum with a quite regularly eliptical margin (Fig. 3C;
<bibRefCitation id="3E1A4B18FFA56D67FE8EFE4060BDF9FA" author="BOURNE, J. D." box="[321,467,1569,1593]" pageId="16" pageNumber="601" pagination="249 - 260" refId="ref42857" refString="BOURNE, J. D. 1980. Revision of Thorell's type species of the family Pacullidae (Araneae). Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot; di Genova 83: 249 - 260." type="journal article" year="1980">Bourne, 1980</bibRefCitation>
: fig. 21), and by largely fused spermathecae with a flat common ventral surface, a widely arched common anterior margin and distinct trenches in anterior and lateral sides.
</paragraph>
<paragraph id="5A3436E9FFA46D66FD88FD5B63EFFA8D" blockId="17.[151,1074,1336,1465]" box="[583,641,1336,1358]" pageId="17" pageNumber="602">FIG. 3</paragraph>
</subSubSection>
<subSubSection id="12916562FFA46D65FF58FD3A6366FE5B" lastPageId="18" lastPageNumber="603" pageId="17" pageNumber="602" type="reference_group">
<paragraph id="5A3436E9FFA46D66FF58FD3A60DDFA7B" blockId="17.[151,1074,1336,1465]" pageId="17" pageNumber="602">
<taxonomicName id="9D8B4D6AFFA46D66FF58FD3A608FFAAE" ID-CoL="6V3LV" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" box="[151,481,1367,1389]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="602" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA46D66FF58FD3A603FFAAE" box="[151,337,1367,1389]" italics="true" pageId="17" pageNumber="602">Perania birmanica</emphasis>
<bibRefCitation id="3E1A4B18FFA46D66FE94FD35608FFAAE" author="THORELL" box="[347,481,1367,1389]" pageId="17" pageNumber="602" pagination="271 - 378" refId="ref43628" refString="THORELL, 1898. Viaggio di Leonardo Fea in Birmania e regioni vicine. 80. Secondo saggio sui ragni birmani. 2. Retitelariae et Orbitelariae. Annali del Museo civico di Storia naturale di Genova (2) 19 [= 39]: 271 - 378." type="journal article" year="1898">Thorell, 1898</bibRefCitation>
</taxonomicName>
, female
<typeStatus id="8530884BFFA46D66FDF3FD3563FEFAAE" box="[572,656,1367,1389]" pageId="17" pageNumber="602" type="holotype">holotype</typeStatus>
. (A) Anterior portion of carapace, dorsal view. (B) Same, lateral view. (C) Dorsal scutum of opisthosoma, dorsal view. (D) Vulva, dorsal view. (E) Same, ventral view. (F) Same, anterior view. Abbreviation: t = trench. Scale lines 1.0 mm (A-B; C),
<quantity id="9D739B0CFFA46D66FEE9FDC1601EFA7B" box="[294,368,1443,1465]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="602" unit="mm" value="0.5">0.5 mm</quantity>
(D-F).
</paragraph>
<paragraph id="5A3436E9FFA46D65FF2CFDBD6366FE5B" blockId="17.[151,1074,1503,1653]" lastBlockId="18.[136,1059,227,408]" lastPageId="18" lastPageNumber="603" pageId="17" pageNumber="602">
NEW CHARACTERS: Vulva with ventral side uniformly pigmented (no unpigmented zone between spermathecae and genital atrium; Fig. 3D-E) and uniformly flat (spermathecae not bulged; Fig. 3F). Vulva similar to that of
<taxonomicName id="9D8B4D6AFFA46D66FC83FE7D62F0F9F5" ID-CoL="773M7" authorityName="Schwendinger" authorityYear="2013" box="[844,926,1566,1590]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="602" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
<emphasis id="68FFEAFBFFA46D66FC83FE7D62F0F9F5" box="[844,926,1566,1590]" italics="true" pageId="17" pageNumber="602">P. ferox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73CC5780FFA46D66FC6AFE7C62B1F9F6" box="[933,991,1566,1589]" pageId="17" pageNumber="602" rank="species">sp. n.</taxonomicNameLabel>
, distinguished by left and right spermathecae without depression or unpigmented zone between them; anterior and lateral sides of spermathecae with narrow porepatches lying in distinct trenches (Fig. 3F); anterior collar with deep median invagination in posterior margin (Fig. 3D; possibly a condition of the
<typeStatus id="8530884BFFA76D65FD1DF9606241FED9" box="[722,815,258,282]" pageId="18" pageNumber="603" type="holotype">holotype</typeStatus>
and not a character of the species). Outlines of the holotypes spermathecae shining through the intact genital plate illustrated by
<bibRefCitation id="3E1A4B18FFA76D65FE9BF923635FFE9A" author="SCHWENDINGER, P. J." box="[340,561,321,345]" pageId="18" pageNumber="603" pagination="571 - 582" refId="ref43337" refString="SCHWENDINGER, P. J. 1989. On three new armoured spiders (Araneae: Tetrablemmidae, Pacullinae) from Indonesia and Thailand. Revue suisse de Zoologie 96 (3): 571 - 582." type="journal article" year="1989">Schwendinger (1989</bibRefCitation>
: fig. 21). Preanal plate fragmented into microplates, the three posterior ones laterally connected with each other. Anterior opisthosomal plate completely fragmented.
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA76D65FF1BF9C9604BFDBC" pageId="18" pageNumber="603" type="description">
<paragraph id="5A3436E9FFA76D65FF1BF9C9604BFDBC" blockId="18.[136,1059,426,639]" pageId="18" pageNumber="603">
RELATIONSHIPS: Strong similarities in copulatory organs of females and close geographical proximity indicate that
<taxonomicName id="9D8B4D6AFFA76D65FDD4F9A963CBFE21" authorityName="Thorell" authorityYear="1898" box="[539,677,458,482]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA76D65FDD4F9A963CBFE21" box="[539,677,458,482]" italics="true" pageId="18" pageNumber="603">P. birmanica</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="9D8B4D6AFFA76D65FC43F9A962B0FE21" authorityName="Schwendinger" authorityYear="2013" box="[908,990,458,482]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="ferox" status="sp. nov.">
<emphasis id="68FFEAFBFFA76D65FC43F9A962B0FE21" box="[908,990,458,482]" italics="true" pageId="18" pageNumber="603">P. ferox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73CC5780FFA76D65FC28F9A9654DFE21" box="[999,1059,459,482]" pageId="18" pageNumber="603" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="9D8B4D6AFFA76D65FF74F9886003FDC2" authorityName="Schwendinger" authorityYear="2013" box="[187,365,489,513]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="quadrifurcata" status="sp. nov.">
<emphasis id="68FFEAFBFFA76D65FF74F9886003FDC2" box="[187,365,489,513]" italics="true" pageId="18" pageNumber="603">P. quadrifurcata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73CC5780FFA76D65FEB5F98860D6FDC2" box="[378,440,490,513]" pageId="18" pageNumber="603" rank="species">sp. n.</taxonomicNameLabel>
Therefore it is quite likely that the unknown male of
<taxonomicName id="9D8B4D6AFFA76D65FF47FA686061FDE2" authorityName="Thorell" authorityYear="1898" box="[136,271,521,545]" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA76D65FF47FA686061FDE2" box="[136,271,521,545]" italics="true" pageId="18" pageNumber="603">P. birmanica</emphasis>
</taxonomicName>
also has a long clypeal process.
<bibRefCitation id="3E1A4B18FFA76D65FDABFA6B6256FDE2" author="LEHTINEN, P. T." box="[612,824,521,545]" pageId="18" pageNumber="603" pagination="1 - 151" refId="ref43059" refString="LEHTINEN, P. T. 1981. Spiders of the Oriental-Australian region. III. Tetrablemmidae, with a world revision. Acta Zoological Fennica 162: 1 - 151." type="journal article" year="1981">Lehtinen (1981: 16)</bibRefCitation>
speculated that
<taxonomicName id="9D8B4D6AFFA76D65FC2EFA6861B9FD83" authorityName="Thorell" authorityYear="1898" class="Arachnida" family="Pacullidae" genus="Perania" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="birmanica">
<emphasis id="68FFEAFBFFA76D65FC2EFA6861B9FD83" italics="true" pageId="18" pageNumber="603">P. birmanica</emphasis>
</taxonomicName>
could be close to
<taxonomicName id="9D8B4D6AFFA76D65FE6EFA4B6364FD83" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[417,522,553,576]" class="Arachnida" family="Pacullidae" genus="Paculla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="603" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="68FFEAFBFFA76D65FE6EFA4B6364FD83" box="[417,522,553,576]" italics="true" pageId="18" pageNumber="603">P. armata</emphasis>
</taxonomicName>
from
<collectingRegion id="984FF80BFFA76D65FD81FA4A63C9FD83" box="[590,679,552,576]" country="Indonesia" name="Sumatera Utara" pageId="18" pageNumber="603">Sumatra</collectingRegion>
, because
<bibRefCitation id="3E1A4B18FFA76D65FCDCFA4A62DFFD83" author="THORELL" box="[787,945,552,576]" pageId="18" pageNumber="603" pagination="271 - 378" refId="ref43628" refString="THORELL, 1898. Viaggio di Leonardo Fea in Birmania e regioni vicine. 80. Secondo saggio sui ragni birmani. 2. Retitelariae et Orbitelariae. Annali del Museo civico di Storia naturale di Genova (2) 19 [= 39]: 271 - 378." type="journal article" year="1898">Thorell (1898)</bibRefCitation>
compared the first with the second species, but there is no evidence for a close relationship between them.
</paragraph>
</subSubSection>
<subSubSection id="12916562FFA76D65FF1BFAED618DFCC6" pageId="18" pageNumber="603" type="distribution">
<paragraph id="5A3436E9FFA76D65FF1BFAED618DFCC6" blockId="18.[136,1059,654,773]" pageId="18" pageNumber="603">
<materialsCitation id="EAE33CB4FFA76D65FF1BFAED61B2FCC6" county="Karen" location="Toungoo" municipality="State" pageId="18" pageNumber="603" specimenCount="1" stateProvince="Kayin">
DISTRIBUTION: Known only from the
<typeStatus id="8530884BFFA76D65FD4DFAED63DEFD64" box="[642,688,655,679]" pageId="18" pageNumber="603">type</typeStatus>
locality in the mountains of the
<collectingCounty id="B3554E65FFA76D65FF47FACD61A7FD05" box="[136,201,687,710]" pageId="18" pageNumber="603">Karen</collectingCounty>
(=
<collectingRegion id="984FF80BFFA76D65FF3DFACC605DFD05" box="[242,307,686,710]" country="Myanmar" name="Kayin" pageId="18" pageNumber="603">Kayin</collectingRegion>
)
<collectingMunicipality id="BA50AC93FFA76D65FE8BFACC6017FD05" box="[324,377,686,710]" pageId="18" pageNumber="603">State</collectingMunicipality>
, northeast of
<location id="5F546032FFA76D65FDDFFACD6300FD04" LSID="urn:lsid:plazi:treatment:D22287FFFFBA6D65FF58FCBC618DFCC6:5F546032FFA76D65FDDFFACD6300FD04" box="[528,622,687,711]" county="Karen" municipality="State" name="Toungoo" pageId="18" pageNumber="603" stateProvince="Kayin">Toungoo</location>
(Fig. 1, locality 12).
<location id="5F546032FFA76D65FC97FACC62D2FD05" LSID="urn:lsid:plazi:treatment:D22287FFFFBA6D65FF58FCBC618DFCC6:5F546032FFA76D65FC97FACC62D2FD05" box="[856,956,686,710]" county="Karen" municipality="State" name="This area" pageId="18" pageNumber="603" stateProvince="Kayin">This area</location>
has been off limits to foreign visitors without a special permit for the last four and a half decades
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>