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<document id="E66F29BB0BEC96DAC9B91FE2C888B5A5" ID-CLB-Dataset="4084" ID-DOI="10.11646/palaeoentomology.4.3.5" ID-GBIF-Dataset="963caafd-cffd-47ba-85d2-3bb469d24dd2" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507869" ID-ZooBank="C625EE39-050C-49CC-9182-220F212BCFB4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631645642513" checkinUser="carolina" docAuthor="NEL, ANDRÉ" docDate="2021" docId="8208B456FFC1581BFF02AA34266F809D" docLanguage="en" docName="palaeoentomology.4.3.5.pdf" docOrigin="Palaeoentomology 4 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.3.5" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Dakotaeschnidium spedeni NEL 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="210" masterDocId="7E31CC2EFFC05819FF8AA81A25668441" masterDocTitle="Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous" masterLastPageNumber="212" masterPageNumber="209" pageNumber="210" updateTime="1699448782307" updateUser="ExternalLinkService">
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<mods:title id="17EAC60E5B076541CCCD0603454A7EEE">Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous</mods:title>
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<mods:namePart id="528888D1CBBB2B5CC712C6564882DC67">NEL, ANDRÉ</mods:namePart>
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<mods:title id="BA15BE3215FA4020CE400B811079690A">Palaeoentomology</mods:title>
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<mods:date id="C04899AB92223CF29D52EF193DF62C96">2021</mods:date>
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<mods:classification id="D8E5DF6540AB9E389D13BEDD80BFD268">journal article</mods:classification>
<mods:identifier id="4267B55E7B500C4C47FD1AC2C1908310" type="CLB-Dataset">4084</mods:identifier>
<mods:identifier id="1C0120A881B05D33F72CA456C386BD57" type="DOI">10.11646/palaeoentomology.4.3.5</mods:identifier>
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<treatment id="8208B456FFC1581BFF02AA34266F809D" ID-DOI="http://doi.org/10.5281/zenodo.5507873" ID-GBIF-Taxon="188607635" ID-Zenodo-Dep="5507873" LSID="urn:lsid:plazi:treatment:8208B456FFC1581BFF02AA34266F809D" httpUri="http://treatment.plazi.org/id/8208B456FFC1581BFF02AA34266F809D" lastPageId="2" lastPageNumber="210" pageId="1" pageNumber="210">
<subSubSection id="42BB56CBFFC15818FF02AA3427478608" box="[136,545,558,585]" pageId="1" pageNumber="210" type="nomenclature">
<paragraph id="0A1E0540FFC15818FF02AA3427478608" blockId="1.[136,778,558,694]" box="[136,545,558,585]" pageId="1" pageNumber="210">
<heading id="5156B22CFFC15818FF02AA3427478608" bold="true" box="[136,545,558,585]" fontSize="10" level="2" pageId="1" pageNumber="210" reason="3">
<emphasis id="38D5D952FFC15818FF02AA3427478608" bold="true" box="[136,545,558,585]" pageId="1" pageNumber="210">
<taxonomicName id="CDA17EC3FFC15818FF02AA3424A68608" authority="NEL, 2021" authorityName="NEL" authorityYear="2021" box="[136,448,558,585]" class="Insecta" family="Aeschnidiidae" genus="Dakotaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="spedeni" status="sp. nov.">
<emphasis id="38D5D952FFC15818FF02AA3424A68608" bold="true" box="[136,448,558,585]" italics="true" pageId="1" pageNumber="210">Dakotaeschnidium spedeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="23E66429FFC15818FE4DAA3527478608" box="[455,545,559,585]" pageId="1" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FF02AA4F245F86F7" pageId="1" pageNumber="210" type="description">
<paragraph id="0A1E0540FFC15818FF02AA4F245F86F7" blockId="1.[136,778,558,694]" pageId="1" pageNumber="210">
(
<figureCitation id="929A19C5FFC15818FF1AAA4F25A8862C" box="[144,206,597,621]" captionStart="FIGURE 1" captionStartId="1.[136,235,1962,1986]" captionTargetBox="[142,772,1156,1938]" captionTargetId="figure-662@1.[142,772,1156,1938]" captionTargetPageId="1" captionText="FIGURE 1. Dakotaeschnidium spedeni gen. et sp. nov., holotype YPM IP 028154. A, Part. B, Counterpart. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5507871" httpUri="https://zenodo.org/record/5507871/files/figure.png" pageId="1" pageNumber="210">Fig. 1</figureCitation>
)
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urn:lsid:zoobank.org:pub:
<uuid id="7E073F95FFC15818FE2DAA63245F86F7" pageId="1" pageNumber="210">C625EE39-050C-49CC-9182- 220F212BCFB4</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FF02AAFD247E872C" pageId="1" pageNumber="210" type="materials_examined">
<paragraph id="0A1E0540FFC15818FF02AAFD247E872C" blockId="1.[136,779,743,1132]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FF02AAFD259486BE" bold="true" box="[136,242,743,767]" pageId="1" pageNumber="210">Material.</emphasis>
HolotypeYPM IP 028154 (collector Ian G. Speden, 1962, IP number 28154; lot count 1; part/counterpart). Invertebrate Paleontology, Yale Peabody Museum, New Haven,
<collectingCountry id="72B645D0FFC15818FF51AB4F2472872C" box="[219,276,853,877]" name="United States of America" pageId="1" pageNumber="210">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FF37AB63243687F7" pageId="1" pageNumber="210" type="etymology">
<paragraph id="0A1E0540FFC15818FF37AB63243687F7" blockId="1.[136,779,743,1132]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FF37AB63245887D0" bold="true" box="[189,318,889,913]" pageId="1" pageNumber="210">Etymology.</emphasis>
Named after Ian G. Speden who collected the
<typeStatus id="D51ABBE2FFC15818FF25AB8425BB87F7" box="[175,221,926,950]" pageId="1" pageNumber="210">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FF37ABD8259787BE" pageId="1" pageNumber="210" type="diagnosis">
<paragraph id="0A1E0540FFC15818FF37ABD8259787BE" blockId="1.[136,779,743,1132]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FF37ABD82454879B" bold="true" box="[189,306,962,986]" pageId="1" pageNumber="210">Diagnosis.</emphasis>
As for the genus. Wing membrane darkened.
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FF37AC11219E872B" pageId="1" pageNumber="210" type="description">
<paragraph id="0A1E0540FFC15818FF37AC11219E872B" blockId="1.[136,779,743,1132]" lastBlockId="1.[810,1452,159,2074]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FF37AC11242E8062" bold="true" box="[189,328,1035,1059]" pageId="1" pageNumber="210">Description.</emphasis>
A fragment of hind wing showing the discoidal triangle, hypertriangle, a fragment of the postdiscoidal area, the submedian space and basal part of anal area,
<quantity id="CD59A8A5FFC15818FCEFA88526A384F6" box="[869,965,159,183]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="210" unit="mm" value="11.7">11.7 mm</quantity>
long, 21.0 mm wide; whole membrane very dark with yellow veins; part of wing basal to hypertriangle missing; hypertriangle subdivided into
<emphasis id="38D5D952FFC15818FB43A8F3218F8541" box="[1225,1257,233,256]" italics="true" pageId="1" pageNumber="210">ca.</emphasis>
40 small irregular cells,
<quantity id="CD59A8A5FFC15818FCE2A91626DF8565" box="[872,953,268,292]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="210" unit="mm" value="1.9">1.9 mm</quantity>
wide,
<quantity id="CD59A8A5FFC15818FB8BA91621398565" box="[1025,1119,268,292]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="210" unit="mm" value="11.6">11.6 mm</quantity>
long; discoidal triangle clearly transverse, subdivided into more than 27 irregular cells, with vein MAb more than 7.0 mm long, curved, vein separating discoidal triangle from hypertriangle
<quantity id="CD59A8A5FFC15818FB30A963206D85D0" box="[1210,1291,377,401]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="210" unit="mm" value="3.1">3.1 mm</quantity>
long, and vein MP+CuA
<quantity id="CD59A8A5FFC15818FC1FA984268285F4" box="[917,996,414,438]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="1" pageNumber="210" unit="mm" value="6.6">6.6 mm</quantity>
long; postdiscoidal area very broad at base, with more than 13 rows of cells between MAa and MP; anal area very broad,
<quantity id="CD59A8A5FFC15818FC52A9FC215285BF" box="[984,1076,486,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="1" pageNumber="210" unit="mm" value="16.2">16.2 mm</quantity>
wide; only three posterior branches of AA in anal area; no infrasubdiscoidal space basal to basal subdiscoidal space; basal subdiscoidal space transverse, posteriorly opened because vein AA1a poorly defined, distal subdiscoidal space transverse, with a shape of acute triangle, very broad and long, subdivided into
<emphasis id="38D5D952FFC15818FB4EAA8721BB86F5" box="[1220,1245,669,692]" italics="true" pageId="1" pageNumber="210">ca</emphasis>
. 70 irregular cells, no angle between AAspl and AA1a veins; PsA vein clearly visible vanishing one cell basal to antero-basal angle of discoidal triangle; submedian space subdivided into more than 11 irregular cells; area between RP and MA subdivided into numerous cells disposed in three rows.
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC15818FCD5AB6D20608061" pageId="1" pageNumber="210" type="materials_examined">
<paragraph id="0A1E0540FFC15818FCD5AB6D20608061" blockId="1.[810,1452,159,2074]" pageId="1" pageNumber="210">
<materialsCitation id="BAC90F1DFFC15818FCD5AB6D20608061" ID-GBIF-Occurrence="3382555301" country="United States of America" county="Dewey County" location="Long's Ridge" municipality="Fox Hills Formation" pageId="1" pageNumber="210" specimenCount="1" stateProvince="South Dakota" typeStatus="holotype">
<emphasis id="38D5D952FFC15818FCD5AB6D21EF87CF" bold="true" box="[863,1161,886,911]" pageId="1" pageNumber="210">
<typeStatus id="D51ABBE2FFC15818FCD5AB6D26F187CE" box="[863,919,887,911]" pageId="1" pageNumber="210">Type</typeStatus>
locality and horizon.
</emphasis>
<collectingMunicipality id="EA7A9F3AFFC15818FB1AAB6D200887CF" box="[1168,1390,886,910]" pageId="1" pageNumber="210">Fox Hills Formation</collectingMunicipality>
, Trail City Member.
<collectingCountry id="72B645D0FFC15818FC47AB81216087F2" box="[973,1030,923,947]" name="United States of America" pageId="1" pageNumber="210">USA</collectingCountry>
;
<collectingRegion id="C865CBA2FFC15818FB9EAB8121C887F2" box="[1044,1198,923,947]" country="United States of America" name="South Dakota" pageId="1" pageNumber="210">South Dakota</collectingRegion>
;
<collectingCounty id="E37F7DCCFFC15818FB37AB81200387F2" box="[1213,1381,923,947]" pageId="1" pageNumber="210">Dewey County</collectingCounty>
; float, conc. 5, face on SW side of SSE-trending spur,
<quantity id="CD59A8A5FFC15818FA95ABA520EB8796" box="[1311,1421,959,983]" metricMagnitude="3" metricUnit="m" metricValue="2.01168" pageId="1" pageNumber="210" unit="mi" value="1.25">1.25 miles</quantity>
W of E end of
<location id="0F7E539BFFC15818FC27ABFE212487BD" LSID="urn:lsid:plazi:treatment:8208B456FFC1581BFF02AA34266F809D:0F7E539BFFC15818FC27ABFE212487BD" box="[941,1090,996,1020]" country="United States of America" county="Dewey County" municipality="Fox Hills Formation" name="Long's Ridge" pageId="1" pageNumber="210" stateProvince="South Dakota">Long’s Ridge</location>
,
<location id="0F7E539BFFC15818FBC4ABFE218487BD" LSID="urn:lsid:plazi:treatment:8208B456FFC1581BFF02AA34266F809D:0F7E539BFFC15818FBC4ABFE218487BD" box="[1102,1250,996,1020]" country="United States of America" county="Dewey County" municipality="Fox Hills Formation" name="Long's Ridge" pageId="1" pageNumber="210" stateProvince="South Dakota">Long’s Ridge</location>
, and
<locationDeviation id="1BAC1C62FFC15818FA97ABFE21218061" location="Whitehorse. Maastrichtian" pageId="1" pageNumber="210">
<quantity id="CD59A8A5FFC15818FA97ABFE200B87BD" box="[1309,1389,996,1020]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="1" pageNumber="210" unit="mi" value="6.0">6 miles</quantity>
W of Whitehorse. Maastrichtian
</locationDeviation>
,
<location id="0F7E539BFFC15818FBD8AC1320608061" LSID="urn:lsid:plazi:treatment:8208B456FFC1581BFF02AA34266F809D:0F7E539BFFC15818FBD8AC1320608061" box="[1106,1286,1032,1056]" country="United States of America" county="Dewey County" municipality="Fox Hills Formation" name="Late Cretaceous." pageId="1" pageNumber="210" stateProvince="South Dakota">Late Cretaceous.</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="42BB56CBFFC1581BFCD5AC37266F809D" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210" type="discussion">
<paragraph id="0A1E0540FFC15818FCD5AC37217B8227" blockId="1.[810,1452,159,2074]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FCD5AC3726A88004" bold="true" box="[863,974,1069,1093]" pageId="1" pageNumber="210">Remarks.</emphasis>
The very broad anal of this fossil is typical of that of a hind wing of an
<taxonomicName id="CDA17EC3FFC15818FBF2AC4B20758028" authorityName="Needham" authorityYear="1903" box="[1144,1299,1105,1129]" class="Insecta" family="Aeschnidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="family">Aeschnidiidae</taxonomicName>
, with a basal subdiscoidal space and a distal subdiscoidal space. The transverse discoidal triangle is also only found in this family (
<bibRefCitation id="6E3078B1FFC15818FC09ACA421068097" author="Fleck, G. &amp; Nel, A." box="[899,1120,1214,1238]" pageId="1" pageNumber="210" pagination="1 - 180" refId="ref2691" refString="Fleck, G. &amp; Nel, A. (2003) Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica, 153, 1 - 180." type="journal article" year="2003">Fleck &amp; Nel, 2003</bibRefCitation>
). The particular shape of the distal subdiscoidal space of YPM IP 028154 is also found in
<taxonomicName id="CDA17EC3FFC15818FCA0AD1D20C1815E" authority="Nel &amp; Martinez-Delclos, 1993" authorityName="Nel &amp; Martinez-Delclos" authorityYear="1993" box="[810,1447,1287,1311]" class="Insecta" family="Aeschnidiidae" genus="Gigantoaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="38D5D952FFC15818FCA0AD1D2104815E" box="[810,1122,1287,1311]" italics="true" pageId="1" pageNumber="210">Gigantoaeschnidium ibericus</emphasis>
Nel &amp; Martinez-Delclòs, 1993
</taxonomicName>
, but the basal subdiscoidal space of
<taxonomicName id="CDA17EC3FFC15818FB38AD3120F68102" authorityName="Nel &amp; Martinez-Delclos" authorityYear="1993" box="[1202,1424,1323,1347]" class="Insecta" family="Aeschnidiidae" genus="Gigantoaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="38D5D952FFC15818FB38AD3120F68102" box="[1202,1424,1323,1347]" italics="true" pageId="1" pageNumber="210">Gigantoaeschnidium</emphasis>
</taxonomicName>
is much smaller than that of YPM IP 028154, and it has an angle between AAspl and AA1a veins (Fleck &amp; Nel, 1993: fig. 63a).
<taxonomicName id="CDA17EC3FFC15818FC13AD82201C81F1" authority="Fleck &amp; Nel, 1993" authorityName="Fleck &amp; Nel" authorityYear="1993" box="[921,1402,1432,1456]" class="Insecta" family="Aeschnidiidae" genus="Angloaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="toyei">
<emphasis id="38D5D952FFC15818FC13AD8221F881F1" box="[921,1182,1432,1456]" italics="true" pageId="1" pageNumber="210">Angloaeschnidium toyei</emphasis>
Fleck &amp; Nel, 1993
</taxonomicName>
and
<taxonomicName id="CDA17EC3FFC15818FCA0ADA620538194" authority="Fleck &amp; Nel, 1993" authorityName="Fleck &amp; Nel" authorityYear="1993" box="[810,1333,1468,1493]" class="Insecta" family="Aeschnidiidae" genus="Cooperaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="durandi">
<emphasis id="38D5D952FFC15818FCA0ADA621388195" box="[810,1118,1468,1492]" italics="true" pageId="1" pageNumber="210">Cooperaeschnidium durandi</emphasis>
Fleck &amp; Nel, 1993
</taxonomicName>
have their distal subdiscoidal spaces similar to those of YPM IP 028154 but their distal subdiscoidal spaces are much smaller than that of YPM IP 028154. They also have only one row of cells in their hypertriangles.
</paragraph>
<paragraph id="0A1E0540FFC15818FCD5AE6926178393" blockId="1.[810,1452,159,2074]" pageId="1" pageNumber="210">
Even if
<taxonomicName id="CDA17EC3FFC15818FC4AAE6920CD82CB" authority="Martill &amp; Nel, 1996" authorityName="Martill &amp; Nel" authorityYear="1996" box="[960,1451,1650,1675]" class="Insecta" family="Aeschnidiidae" genus="Santanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="gabbotti">
<emphasis id="38D5D952FFC15818FC4AAE6921D082CB" box="[960,1206,1650,1674]" italics="true" pageId="1" pageNumber="210">Santanoptera gabbotti</emphasis>
<bibRefCitation id="6E3078B1FFC15818FB49AE6820CD82CB" author="Martill, D. M. &amp; Nel, A." box="[1219,1451,1650,1675]" pageId="1" pageNumber="210" pagination="279 - 292" refId="ref2963" refString="Martill, D. M. &amp; Nel, A. (1996) A new dragonfly from the Crato Formation (Lower Cretaceous, Aptian) of N. E. Brazil. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 1996, 279 - 292. https: // doi. org / 10.1127 / njgpm / 1996 / 1996 / 279" type="journal article" year="1996">Martill &amp; Nel, 1996</bibRefCitation>
</taxonomicName>
(Aptian, Crato Formation,
<collectingCountry id="72B645D0FFC15818FBCCAE8D21EF82EE" box="[1094,1161,1687,1711]" name="Brazil" pageId="1" pageNumber="210">Brazil</collectingCountry>
) is based on a forewing, its distal subdiscoidal space is similar to that of YPM IP 028154, and it has no angle between AAspl and AA1a veins (Fleck &amp; Nel, 1993: fig. 89). They also share the presence of two rows of cells in the hypertriangle, two in the submedian space, and two-three in the basal part of area between RP and MA. Nevertheless,
<taxonomicName id="CDA17EC3FFC15818FC4BAF6B213783C9" authorityName="Martill &amp; Nel" authorityYear="1996" box="[961,1105,1905,1928]" class="Insecta" family="Aeschnidiidae" genus="Santanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="38D5D952FFC15818FC4BAF6B213783C9" box="[961,1105,1905,1928]" italics="true" pageId="1" pageNumber="210">Santanoptera</emphasis>
</taxonomicName>
has a smaller basal subdiscoidal space that is posteriorly closed by a well-defined vein AA1a.
</paragraph>
<caption id="5EDE55C8FFC15818FF02AFB025B88C4B" ID-DOI="http://doi.org/10.5281/zenodo.5507871" ID-Zenodo-Dep="5507871" httpUri="https://zenodo.org/record/5507871/files/figure.png" pageId="1" pageNumber="210" startId="1.[136,235,1962,1986]" targetBox="[142,772,1156,1938]" targetPageId="1">
<paragraph id="0A1E0540FFC15818FF02AFB025B88C4B" blockId="1.[136,777,1962,2058]" pageId="1" pageNumber="210">
<emphasis id="38D5D952FFC15818FF02AFB024688383" bold="true" box="[136,270,1962,1986]" pageId="1" pageNumber="210">FIGURE 1.</emphasis>
<taxonomicName id="CDA17EC3FFC15818FE94AFB1275E8382" authority="NEL, 2021" authorityName="NEL" authorityYear="2021" box="[286,568,1963,1987]" class="Insecta" family="Aeschnidiidae" genus="Dakotaeschnidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="210" phylum="Arthropoda" rank="species" species="spedeni" status="gen. et sp. nov.">
<emphasis id="38D5D952FFC15818FE94AFB1275E8382" box="[286,568,1963,1987]" italics="true" pageId="1" pageNumber="210">Dakotaeschnidium spedeni</emphasis>
</taxonomicName>
<emphasis id="38D5D952FFC15818FDC3AFB126658382" bold="true" box="[585,771,1963,1987]" pageId="1" pageNumber="210">
<taxonomicNameLabel id="23E66429FFC15818FDC3AFB126658382" box="[585,771,1963,1987]" pageId="1" pageNumber="210" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype YPM IP 028154.
<emphasis id="38D5D952FFC15818FE2AAFD424D483A7" bold="true" box="[416,434,1998,2022]" pageId="1" pageNumber="210">A</emphasis>
, Part.
<emphasis id="38D5D952FFC15818FE73AFD5276C83A6" bold="true" box="[505,522,1999,2023]" pageId="1" pageNumber="210">B</emphasis>
, Counterpart. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="0A1E0540FFC1581BFCD5AFC4266F809D" blockId="1.[810,1452,159,2074]" lastBlockId="2.[136,778,1076,1244]" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210">
Despite the fact that the specimen YPM IP 028154 is reduced to the basal third of a hind wing, it has a series of characters allowing to consider that it corresponds to a new genus and species, possibly related to the South American genus
<taxonomicName id="CDA17EC3FFC2581BFF41AC67243D80D5" authorityName="Martill &amp; Nel" authorityYear="1996" box="[203,347,1149,1172]" class="Insecta" family="Aeschnidiidae" genus="Santanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="38D5D952FFC2581BFF41AC67243D80D5" box="[203,347,1149,1172]" italics="true" pageId="2" pageNumber="211">Santanoptera</emphasis>
</taxonomicName>
, as they share the very particular absence of an angle between AAspl and AA1a veins, and presence of several rows of cells in hypertriangle, submedian space, etc.
</paragraph>
</subSubSection>
</treatment>
</document>
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