treatments-xml/data/16/5F/87/165F8783FFEDFFEFFF2E0C7C4BA6FCDD.xml
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<document id="B6D93E5217A5FFCAF76FB1CFFD99A53F" ID-DOI="10.11646/zootaxa.3635.4.4" ID-GBIF-Dataset="5b27dcc4-7605-425c-847d-41f2a8dd86d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="219563" ID-ZooBank="2426930B-7131-49DA-B1BF-682BE3826928" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460586800093" checkinUser="plazi" docAuthor="Jia, Fenglong, Gentili, Elio &amp; Fikáček, Martin" docDate="2013" docId="165F8783FFEDFFEFFF2E0C7C4BA6FCDD" docLanguage="en" docName="zt03635p418.pdf" docOrigin="Zootaxa 3635 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laccobius (Glyptolaccobius) qinlingensis Jia, Gentili &amp; Fikáček, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="406" masterDocId="EA66FFFBFFECFFEBFFB9094B4F71FFFB" masterDocTitle="The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae)" masterLastPageNumber="418" masterPageNumber="402" pageNumber="403" updateTime="1698325284964" updateUser="plazi">
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<mods:title id="2BCA368CC20CEB53CEBF4BA4A528579D">The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae)</mods:title>
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<mods:name id="ED55A3B549F6AE22B9A9838400590899" type="personal">
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<mods:namePart id="481F04E2103CF05FBE1A4B554D9D7002">Jia, Fenglong</mods:namePart>
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<mods:namePart id="D0EB9F4F9F13CCDC79B03B58BB4C574C">Gentili, Elio</mods:namePart>
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<mods:namePart id="A9E2D3BDF2496D03DD2DFB86DDDD43F4">Fikáček, Martin</mods:namePart>
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<mods:title id="7C308AAB9675FAB01C8D0BF18A5FB634">Zootaxa</mods:title>
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<mods:part id="BB3C2A101EF4AFD9D0912B49B2082E57">
<mods:date id="88F348B24ECD0F6FB0D17B9736814847">2013</mods:date>
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<mods:identifier id="368D24C0D16A4E2850E065E372525253" type="DOI">10.11646/zootaxa.3635.4.4</mods:identifier>
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<treatment id="165F8783FFEDFFEFFF2E0C7C4BA6FCDD" ID-DOI="http://doi.org/10.5281/zenodo.6164571" ID-GBIF-Taxon="119564702" ID-Zenodo-Dep="6164571" LSID="urn:lsid:plazi:treatment:165F8783FFEDFFEFFF2E0C7C4BA6FCDD" httpUri="http://treatment.plazi.org/id/165F8783FFEDFFEFFF2E0C7C4BA6FCDD" lastPageId="4" lastPageNumber="406" pageId="1" pageNumber="403">
<subSubSection id="D6EC651EFFEDFFEAFF2E0C7C4E63FA89" pageId="1" pageNumber="403" type="nomenclature">
<paragraph id="9E493695FFEDFFEAFF2E0C7C4D9CFAAA" blockId="1.[151,749,1335,1394]" box="[151,749,1335,1361]" pageId="1" pageNumber="403">
<heading id="C50181F9FFEDFFEAFF2E0C7C4D9CFAAA" bold="true" box="[151,749,1335,1361]" fontSize="11" level="1" pageId="1" pageNumber="403" reason="1">
<emphasis id="AC82EA87FFEDFFEAFF2E0C7C4D9CFAAA" bold="true" box="[151,749,1335,1361]" pageId="1" pageNumber="403">
<taxonomicName id="59F64D16FFEDFFEAFF2E0C7C4DF8FAAA" ID-CoL="3RLL6" box="[151,649,1335,1361]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="403" phylum="Arthropoda" rank="species" species="qinlingensis" status="sp. nov." subGenus="Glyptolaccobius">
<emphasis id="AC82EA87FFEDFFEAFF2E0C7C4E62FAAA" bold="true" box="[151,275,1335,1361]" italics="true" pageId="1" pageNumber="403">Laccobius</emphasis>
(
<emphasis id="AC82EA87FFEDFFEAFE9D0C7C4E97FAAA" bold="true" box="[292,486,1335,1361]" italics="true" pageId="1" pageNumber="403">Glyptolaccobius</emphasis>
)
<emphasis id="AC82EA87FFEDFFEAFE4F0C7C4DF8FAAA" bold="true" box="[502,649,1335,1361]" italics="true" pageId="1" pageNumber="403">qinlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B157FCFFEDFFEAFD290C7C4D9CFAAA" box="[656,749,1335,1361]" pageId="1" pageNumber="403" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E493695FFEDFFEAFF2E0C114E63FA89" blockId="1.[151,749,1335,1394]" box="[151,274,1370,1394]" pageId="1" pageNumber="403">
(
<figureCitation id="06CD2A10FFEDFFEAFF260C114E78FA89" box="[159,265,1370,1394]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="1" pageNumber="403">Figs. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFEDFFEAFF2E0CE84B16F8BD" pageId="1" pageNumber="403" type="materials_examined">
<paragraph id="9E493695FFEDFFEAFF2E0CE84E10F9DD" blockId="1.[151,1437,1441,1970]" pageId="1" pageNumber="403">
<emphasis id="AC82EA87FFEDFFEAFF2E0CE84E47FA40" bold="true" box="[151,310,1442,1467]" pageId="1" pageNumber="403">
<typeStatus id="414D8837FFEDFFEAFF2E0CE84FA1FA40" box="[151,208,1443,1467]" pageId="1" pageNumber="403">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E6E17605FFEDFFEAFEFB0CE94EF9FA41" box="[322,392,1442,1466]" name="China" pageId="1" pageNumber="403">China</collectingCountry>
, Shaanxi Province, Qinlingshan Mts., Fengyukou, 34°01.4ʹN 108°49.2ʹE,
<quantity id="590E9B70FFEDFFEAFB660CE94A59FA41" box="[1247,1320,1442,1466]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="403" unit="m" value="580.0">580 m</quantity>
a.s.l. (the series including the
<typeStatus id="414D8837FFEDFFEAFEC10C8D4EACFA25" box="[376,477,1478,1502]" pageId="1" pageNumber="403" type="holotype">holotype</typeStatus>
, collected by the first author, was taken simultaneously and in the same spot that was collected by Balke &amp; Hájek, hence we provide the complete data on the
<typeStatus id="414D8837FFEDFFEAFC680CA04B70F9F8" box="[977,1025,1515,1539]" pageId="1" pageNumber="403">type</typeStatus>
locality combining the label data of both these series).
</paragraph>
<paragraph id="9E493695FFEDFFEAFF700F784B16F8BD" blockId="1.[151,1437,1441,1970]" pageId="1" pageNumber="403">
<emphasis id="AC82EA87FFEDFFEAFF700F784E8BF9B0" bold="true" box="[201,506,1586,1611]" pageId="1" pageNumber="403">
<typeStatus id="414D8837FFEDFFEAFF700F784E75F9B0" box="[201,260,1587,1611]" pageId="1" pageNumber="403">Type</typeStatus>
material.
<typeStatus id="414D8837FFEDFFEAFE3D0F784E84F9B0" box="[388,501,1587,1611]" pageId="1" pageNumber="403" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂ (SYSU):
<collectingCountry id="E6E17605FFEDFFEAFD360F794D9AF9B1" box="[655,747,1586,1610]" name="China" pageId="1" pageNumber="403">CHINA</collectingCountry>
, Shaanxi, Qinling, Qinlingliang, Fengyukou,
<date id="EA481055FFEDFFEAFB400F794A19F9B1" box="[1273,1384,1586,1610]" pageId="1" pageNumber="403" value="2011-05-12">12.v.2011</date>
, lgt. Fenglong Jia [transcribed from Chinese].
<emphasis id="AC82EA87FFEDFFEAFDCE0F1C4C40F994" bold="true" box="[631,817,1622,1647]" pageId="1" pageNumber="403">
<typeStatus id="414D8837FFEDFFEAFDCE0F1C4D9EF994" box="[631,751,1623,1647]" pageId="1" pageNumber="403" type="paratype">Paratypes</typeStatus>
(58):
</emphasis>
32 spec. (SYSU): same data as the
<typeStatus id="414D8837FFEDFFEAFB610F1D4A4FF995" box="[1240,1342,1622,1646]" pageId="1" pageNumber="403" type="holotype">holotype</typeStatus>
; 2 spec. (MSNV):
<collectingCountry id="E6E17605FFEDFFEAFEB40F314E22F969" box="[269,339,1658,1682]" name="China" pageId="1" pageNumber="403">China</collectingCountry>
, Shaanxi, Prov. / Huashan, rock of old road /
<date id="EA481055FFEDFFEAFCC00F314C96F969" box="[889,999,1658,1682]" pageId="1" pageNumber="403" value="2011-05-12">12.v.2011</date>
/ Fenglong Jia leg.; 9 spec. (NMPC, MSNV):
<collectingCountry id="E6E17605FFEDFFEAFEBB0FD54E2EF94D" box="[258,351,1694,1718]" name="China" pageId="1" pageNumber="403">CHINA</collectingCountry>
: Shaanxi prov.
<date id="EA481055FFEDFFEAFDB60FD54D0DF94D" box="[527,636,1694,1718]" pageId="1" pageNumber="403" value="2011-05-12">12.v.2011</date>
/ ca.
<quantity id="590E9B70FFEDFFEAFD070FD54C76F94D" box="[702,775,1694,1718]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="403" unit="km" value="30.0">30 km</quantity>
SSW of Xian / Qinlingshan Mts., Fengyukou / (wet cliff, waterfall), 34°01.4ʹN / 108°49.2ʹE,
<quantity id="590E9B70FFEDFFEAFD9F0F894D1CF921" box="[550,621,1730,1754]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="403" unit="m" value="580.0">580 m</quantity>
/ M. Balke &amp; J. Hájek lgt.; 12 spec. (NMPC, MSNV):
<collectingCountry id="E6E17605FFEDFFEAFB630F894A46F921" box="[1242,1335,1730,1754]" name="China" pageId="1" pageNumber="403">CHINA</collectingCountry>
: Shaanxi Prov.
<date id="EA481055FFEDFFEAFF610FAD4E34F905" box="[216,325,1766,1790]" pageId="1" pageNumber="403" value="2011-05-10">10.v.2011</date>
/
<quantity id="590E9B70FFEDFFEAFEE30FAD4EDFF905" box="[346,430,1766,1790]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="403" unit="km" value="110.0">110 km</quantity>
NNE Xian, Huayin vill. / Hua Shan Mt. (river valley), / ca.
<quantity id="590E9B70FFEDFFEAFBF60FAD4BE9F905" box="[1103,1176,1766,1790]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="403" unit="m" value="530.0">530 m</quantity>
, 34°31.2ʹN 10°04.6ʹE / M. Balke &amp; J. Hájek lgt.; 3 spec. (NMPC):
<collectingCountry id="E6E17605FFEDFFEAFD3D0E414D90F8D9" box="[644,737,1802,1826]" name="China" pageId="1" pageNumber="403">CHINA</collectingCountry>
: Shaanxi prov.,
<date id="EA481055FFEDFFEAFC2C0E414B61F8D9" box="[917,1040,1802,1826]" pageId="1" pageNumber="403" value="2011-05-08" valueMax="2011-05-09" valueMin="2011-05-08">89.v.2011</date>
/
<quantity id="590E9B70FFEDFFEAFB9E0E414B0DF8D9" box="[1063,1148,1802,1826]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="403" unit="km" value="110.0">110 km</quantity>
NEE Xian, Huyain vill. / Hua Shan Mt. (granite cliff) /
<quantity id="590E9B70FFEDFFEAFE5C0E654D4DF8BD" box="[485,572,1838,1862]" metricMagnitude="3" metricUnit="m" metricValue="1.275" pageId="1" pageNumber="403" unit="m" value="1275.0">1275 m</quantity>
, 34°29.5ʹN 110°05.1ʹE / M. Balke &amp; J. Hájek lgt.
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFEDFFEFFF7E0E194AEDFE1D" lastPageId="4" lastPageNumber="406" pageId="1" pageNumber="403" type="description">
<paragraph id="9E493695FFEDFFEAFF7E0E194EDFF849" blockId="1.[151,1437,1441,1970]" pageId="1" pageNumber="403">
<emphasis id="AC82EA87FFEDFFEAFF7E0E194E28F890" bold="true" box="[199,345,1874,1899]" pageId="1" pageNumber="403">Description.</emphasis>
Total length
<quantity id="590E9B70FFEDFFEAFE570E194D07F891" box="[494,630,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="1" pageNumber="403" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
(
<typeStatus id="414D8837FFEDFFEAFD3C0E194D9CF891" box="[645,749,1874,1898]" pageId="1" pageNumber="403" type="holotype">holotype</typeStatus>
:
<quantity id="590E9B70FFEDFFEAFD410E194C22F891" box="[760,851,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="403" unit="mm" value="2.8">2.8 mm</quantity>
), total width
<quantity id="590E9B70FFEDFFEAFC5E0E194B1EF891" box="[999,1135,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="403" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
(
<typeStatus id="414D8837FFEDFFEAFBC70E194B97F891" box="[1150,1254,1874,1898]" pageId="1" pageNumber="403" type="holotype">holotype</typeStatus>
:
<quantity id="590E9B70FFEDFFEAFB480E194A3DF891" box="[1265,1356,1874,1898]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="403" unit="mm" value="1.5">1.5 mm</quantity>
). Body oval, moderately convex, maximum width in anterior elytral third. Total length / total width ratio = 1.9. Dorsal surface finely pubescent.
</paragraph>
<caption id="CA89661DFFEEFFE9FF2E0E754DABF84B" httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="2" pageNumber="404" targetBox="[156,1435,193,1825]" targetPageId="2">
<paragraph id="9E493695FFEEFFE9FF2E0E754DABF84B" blockId="2.[151,1436,1854,1969]" pageId="2" pageNumber="404">
<emphasis id="AC82EA87FFEEFFE9FF2E0E754E4DF8AF" bold="true" box="[151,316,1854,1877]" pageId="2" pageNumber="404">FIGURES 17.</emphasis>
<taxonomicName id="59F64D16FFEEFFE9FEFD0E744D9CF8AE" box="[324,749,1855,1877]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="404" phylum="Arthropoda" rank="species" species="qinlingensis" status="sp. nov." subGenus="Glyptolaccobius">
<emphasis id="AC82EA87FFEEFFE9FEFD0E744EDDF8AE" box="[324,428,1855,1877]" italics="true" pageId="2" pageNumber="404">Laccobius</emphasis>
(
<emphasis id="AC82EA87FFEEFFE9FE050E744D13F8AE" box="[444,610,1855,1877]" italics="true" pageId="2" pageNumber="404">Glyptolaccobius</emphasis>
)
<emphasis id="AC82EA87FFEEFFE9FDCB0E744D9CF8AE" box="[626,749,1855,1877]" italics="true" pageId="2" pageNumber="404">qinlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B157FCFFEEFFE9FD4C0E744C36F8AF" box="[757,839,1855,1876]" pageId="2" pageNumber="404" rank="species">
<emphasis id="AC82EA87FFEEFFE9FD4C0E744C36F8AF" bold="true" box="[757,839,1855,1876]" pageId="2" pageNumber="404">sp. nov.</emphasis>
</taxonomicNameLabel>
and its localities. 12: general habitus (1: dorsal view; 2: lateral view). 35: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting
<taxonomicName id="59F64D16FFEEFFE9FC4F0E364BFFF869" box="[1014,1166,1916,1938]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="404" phylum="Arthropoda" rank="species" species="qinlingensis">
<emphasis id="AC82EA87FFEEFFE9FC4F0E364BFFF869" box="[1014,1166,1916,1938]" italics="true" pageId="2" pageNumber="404">L. qinlingensis</emphasis>
</taxonomicName>
and
<taxonomicName id="59F64D16FFEEFFE9FB7B0E374A1EF869" box="[1218,1391,1916,1938]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="404" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="AC82EA87FFEEFFE9FB7B0E374A1EF869" box="[1218,1391,1916,1938]" italics="true" pageId="2" pageNumber="404">Aspidytes wrasei</emphasis>
</taxonomicName>
at a granite cliff near Huayin village. Photos 67 by J. Hájek.
</paragraph>
</caption>
<paragraph id="9E493695FFEFFFE8FF7009D34D36FD37" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF7009D34E7CFF4B" box="[201,269,152,176]" italics="true" pageId="3" pageNumber="405">Head.</emphasis>
Coloration black, without pale preocular spots, surface with dense and coarse punctures, without microsculpture; systematic punctures indistinct; periocular sulci vaguely defined, reaching anterolateral branches of frontoclypeal suture. Labrum blackish, about 2.3× as wide as long, without specula in both sexes; anterior margin very feebly emarginated medially; lateral portions sharply angulated. Anterior margin of clypeus deeply excised. Eyes oblong, oblique, closest to each other posteriorly, slightly protruding laterad, separated by distance equal to 2.35× of width of one eye in dorsal view; posterior margin of eye nearly straight in dorsal view, distinctly reniform in lateral view; postocular portion of tempora short. Mentum strongly rugose, bearing few coarse punctures, not emarginate anteriorly. Submentum not microgranulate. Gula microgranulate, gular sutures widely separated, tentorial pits moderately large, very distinct. Maxillary palpi yellowish brown, dark at apex; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length, palpomere 3 dilated at apex; palpomere 4 elongate, nearly 1.4× as long as palpomere 3, asymmetrical, inner margin straight and outer margin convex, apex truncate. Antenna with eight antennomeres; scape (antennomere 1) longer than antennomeres 24 combined; pedicel (antennomere 2) cone-shaped; two intermediate antennomeres (antennomeres 34) very short; cupule (antennomere 5) asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, bearing densely arranged setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 widest, antennomere 7 shortest and narrowest.
</paragraph>
<paragraph id="9E493695FFEFFFE8FF700B934E20FB87" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF700B934E50FD0B" box="[201,289,728,752]" italics="true" pageId="3" pageNumber="405">Thorax.</emphasis>
Pronotum transverse, slightly impressed sublaterally, 0.4× as long as wide; black with yellowish lateral margins; yellow area about 0.5× as wide as posterior margin of eye along base, sometimes yellow edge vaguely defined from black disc; surface smooth as that on head, without microsculpture; punctures coarse and dense, bearing decumbent whitish setae. Prosternum black, tectiform, with longitudinal keel. Scutellar shield equilaterally triangular, black, with few punctures. Mesoventrite with longitudinal, anteriorly arrow-shaped keel. Elytra combined slightly elongate, ca. 1.2× as long as wide, blackish with yellowish lateral margins and apices; without pale yellow basal spot; 10 longitudinal rows of regularly impressed primary punctures alternating with 10 rows (interstriae) of sparser and more or less irregular secondary punctures; secondary punctures of same size as primary punctures; all punctures bearing long whitish semierect setae. Parasutural furrow reaching anterior third of elytral length, very conspicuous on posterior elytral third and elytral apex. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous with an oval median glabrous area and a longitudinal sulcus posteromedially.
</paragraph>
<paragraph id="9E493695FFEFFFE8FF700DC24E7CFA67" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF700DC24E74FB5B" box="[201,261,1161,1184]" italics="true" pageId="3" pageNumber="405">Legs.</emphasis>
Procoxae and trochanters pubescent; profemora setiferous on basal half, smooth distally, with tibial grooves; protibiae smooth, each bearing stiff setae and two apical spurs. Mesofemora smooth, with tibial grooves; ventral side of mesotibiae with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Metatrochanters smooth and shining, only very finely pubescent mesally; metafemora smooth, each with scarce punctures and tibial grooves; metatibiae curved, ventral side of each tibia with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges. Natatory setae nearly absent on dorsal surface of tarsi. Metatarsi with second tarsomere as long as third and fourth combined; fourth metatarsomere about 0.65× as long as third tarsomere.
</paragraph>
<paragraph id="9E493695FFEFFFE8FF700CE34E1AFA1F" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF700CE34E4BFA3B" box="[201,314,1448,1472]" italics="true" pageId="3" pageNumber="405">Abdomen.</emphasis>
Ventrites 14 smooth, without microsculpture, with transverse series of setae; last two ventrites densely pubescent.
</paragraph>
<paragraph id="9E493695FFEFFFE8FF700CBB4B93F9D7" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF700CBB4E47F9F3" box="[201,310,1520,1544]" italics="true" pageId="3" pageNumber="405">Aedeagus</emphasis>
(
<figureCitation id="06CD2A10FFEFFFE8FEFD0CBB4EC1F9F3" box="[324,432,1520,1544]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="3" pageNumber="405">Figs. 35</figureCitation>
). Total length
<quantity id="590E9B70FFEFFFE8FDE80CBB4DDBF9F3" box="[593,682,1520,1544]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="405" unit="mm" value="0.6">0.6 mm</quantity>
. Parameres nearly 1.65× as long as phallobase. Median lobe almost as long as parameres, with a series of stout spines laterally, its apex rounded, subdivided medially.
</paragraph>
<paragraph id="9E493695FFEFFFE8FF700F7C4FAEF843" blockId="3.[151,1437,151,1976]" pageId="3" pageNumber="405">
<emphasis id="AC82EA87FFEFFFE8FF700F7C4EA8F9AB" bold="true" box="[201,473,1591,1616]" pageId="3" pageNumber="405">Differential diagnosis.</emphasis>
<taxonomicName id="59F64D16FFEFFFE8FE500F734D85F9AB" box="[489,756,1592,1616]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="species" species="qinlingensis" status="sp. nov.">
<emphasis id="AC82EA87FFEFFFE8FE500F734D85F9AB" box="[489,756,1592,1616]" italics="true" pageId="3" pageNumber="405">Laccobius qinlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B157FCFFEFFFE8FCBD0F734C16F9AB" box="[772,871,1592,1616]" pageId="3" pageNumber="405" rank="species">
<emphasis id="AC82EA87FFEFFFE8FCBD0F734C16F9AB" bold="true" box="[772,871,1592,1616]" pageId="3" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
is easily distinguished from other species of
<taxonomicName id="59F64D16FFEFFFE8FF2E0F174E3EF98F" box="[151,335,1628,1652]" class="Insecta" family="Hydrophilidae" genus="Glyptolaccobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="AC82EA87FFEFFFE8FF2E0F174E3EF98F" box="[151,335,1628,1652]" italics="true" pageId="3" pageNumber="405">Glyptolaccobius</emphasis>
</taxonomicName>
by the combination of following characters: body large (
<quantity id="590E9B70FFEFFFE8FBAB0F174BD2F98F" box="[1042,1187,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="3" pageNumber="405" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
long); head without preocular spots; pronotum with pale lateral margins; mesoventrite with longitudinal keel, arrow-shaped anteriorly; parasutural stria of elytra distinct, reaching anterior third of elytral length; elytra with 10 rows of distinct primary punctures and somewhat irregular secondary rows of punctures that are of the same size as primary rows; lateral margins of elytra and elytral apex pale; pale spots near scutellum absent; elytral rows of punctures not sulciform; mentum rugose. Besides the external characters,
<taxonomicName id="59F64D16FFEFFFE8FD7B0E5B4CB4F8D3" box="[706,965,1808,1832]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="species" species="qinlingensis" status="sp. nov.">
<emphasis id="AC82EA87FFEFFFE8FD7B0E5B4CB4F8D3" box="[706,965,1808,1832]" italics="true" pageId="3" pageNumber="405">Laccobius qinlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B157FCFFEFFFE8FC740E5B4B59F8D3" box="[973,1064,1808,1832]" pageId="3" pageNumber="405" rank="species">
<emphasis id="AC82EA87FFEFFFE8FC740E5B4B59F8D3" bold="true" box="[973,1064,1808,1832]" pageId="3" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all
<taxonomicName id="59F64D16FFEFFFE8FB5D0E5B4AEDF8D3" box="[1252,1436,1808,1832]" class="Insecta" family="Hydrophilidae" genus="Glyptolaccobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="AC82EA87FFEFFFE8FB5D0E5B4AEDF8D3" box="[1252,1436,1808,1832]" italics="true" pageId="3" pageNumber="405">Glyptolaccobius</emphasis>
</taxonomicName>
by the median lobe bearing lateral subapical rows of stout spines (
<figureCitation id="06CD2A10FFEFFFE8FC160E7F4B68F8B7" box="[943,1049,1844,1868]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="3" pageNumber="405">Fig. 3, 5</figureCitation>
). These spines are unique for
<taxonomicName id="59F64D16FFEFFFE8FA3C0E7E4E5DF88B" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="species" species="quinlingensis">
<emphasis id="AC82EA87FFEFFFE8FA3C0E7E4E5DF88B" italics="true" pageId="3" pageNumber="405">L. quinlingensis</emphasis>
</taxonomicName>
; no similar structure is known in any other
<taxonomicName id="59F64D16FFEFFFE8FCFF0E134CC8F88B" box="[838,953,1880,1904]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="AC82EA87FFEFFFE8FCFF0E134CC8F88B" box="[838,953,1880,1904]" italics="true" pageId="3" pageNumber="405">Laccobius</emphasis>
</taxonomicName>
species except
<taxonomicName id="59F64D16FFEFFFE8FBC00E124E5DF86F" authority="Fauvel, 1883" authorityName="Fauvel" authorityYear="1883" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="405" phylum="Arthropoda" rank="species" species="elevatus" subGenus="Yateberosus">
<emphasis id="AC82EA87FFEFFFE8FBC00E124BE1F88B" box="[1145,1168,1881,1904]" italics="true" pageId="3" pageNumber="405">L.</emphasis>
(
<emphasis id="AC82EA87FFEFFFE8FB1C0E134A59F88B" box="[1189,1320,1880,1904]" italics="true" pageId="3" pageNumber="405">Yateberosus</emphasis>
)
<emphasis id="AC82EA87FFEFFFE8FAF90E134AEDF88B" box="[1344,1436,1880,1904]" italics="true" pageId="3" pageNumber="405">elevatus</emphasis>
Fauvel, 1883
</taxonomicName>
from
<collectingCountry id="E6E17605FFEFFFE8FECC0E364D56F86F" box="[373,551,1916,1940]" name="New Caledonia" pageId="3" pageNumber="405">New Caledonia</collectingCountry>
, although the spines are much thinner in that species (for details, see Gentili 1988).
</paragraph>
<paragraph id="9E493695FFE8FFEFFF7009DC4E0CFF2F" blockId="4.[151,1436,151,212]" pageId="4" pageNumber="406">
The key to the species of the subgenus
<taxonomicName id="59F64D16FFE8FFEFFD2509D34C22FF4B" box="[668,851,152,176]" class="Insecta" family="Hydrophilidae" genus="Glyptolaccobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="AC82EA87FFE8FFEFFD2509D34C22FF4B" box="[668,851,152,176]" italics="true" pageId="4" pageNumber="406">Glyptolaccobius</emphasis>
</taxonomicName>
published by Gentili &amp; Fikáček (2009) should be modified as follows:
</paragraph>
<paragraph id="9E493695FFE8FFEFFF2E08484AEDFE1D" blockId="4.[151,1437,259,486]" pageId="4" pageNumber="406">
4. Entire elytra shining black, without yellow lateral margins. Apex of median lobe widely expanded (Gentili &amp; Fikáček 2009:
<figureCitation id="06CD2A10FFE8FFEFFF58086A4E6AFECD" box="[225,283,289,310]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="4" pageNumber="406">Fig. 4</figureCitation>
). Body length
<quantity id="590E9B70FFE8FFEFFE10086A4D6AFECD" box="[425,539,289,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="4" pageNumber="406" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.32.5 mm</quantity>
........................................
<taxonomicName id="59F64D16FFE8FFEFFBBE086A4AEDFECD" authority="Kamite, Ogata &amp; Hikida, 2007" authorityName="Kamite, Ogata &amp; Hikida" authorityYear="2007" box="[1031,1436,289,310]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="406" phylum="Arthropoda" rank="species" species="moriyai">
<emphasis id="AC82EA87FFE8FFEFFBBE086A4B1FFECD" bold="true" box="[1031,1134,289,310]" italics="true" pageId="4" pageNumber="406">L. moriyai</emphasis>
Kamite, Ogata &amp; Hikida, 2007
</taxonomicName>
At least lateral margins and apex of elytra yellowish. Apex of median lobe widened or not. Body length less than
<quantity id="590E9B70FFE8FFEFFA8E08754AF0FEA8" box="[1335,1409,318,339]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="406" unit="mm" value="2.3">2.3 mm</quantity>
or more than
<quantity id="590E9B70FFE8FFEFFEFE08104EE0FE8B" box="[327,401,347,368]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="406" unit="mm" value="2.7">2.7 mm</quantity>
.................................................................................... 4a 4a. Body length
<quantity id="590E9B70FFE8FFEFFEE608324EA6FE75" box="[351,471,377,398]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="4" pageNumber="406" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
. Posterior pronotal margin yellowish laterally, median lobe somewhat widened before apex, with a series or large spine-like projections laterally (
<figureCitation id="06CD2A10FFE8FFEFFD2708DD4D8BFE50" box="[670,762,406,427]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="4" pageNumber="406">Figs. 3, 5</figureCitation>
)....................................
<emphasis id="AC82EA87FFE8FFEFFB0F08DD4AEAFE50" bold="true" box="[1206,1435,406,427]" pageId="4" pageNumber="406">
<taxonomicName id="59F64D16FFE8FFEFFB0F08DD4A38FE50" box="[1206,1353,406,427]" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="406" phylum="Arthropoda" rank="species" species="qinlingensis" status="sp. nov.">
<emphasis id="AC82EA87FFE8FFEFFB0F08DD4A38FE50" bold="true" box="[1206,1353,406,427]" italics="true" pageId="4" pageNumber="406">L. qinlingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7B157FCFFE8FFEFFAF608DD4AEAFE50" box="[1359,1435,406,427]" pageId="4" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Body length smaller than
<quantity id="590E9B70FFE8FFEFFE6E08F84D53FE33" box="[471,546,435,456]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="406" unit="mm" value="2.3">2.3 mm</quantity>
. Posterior pronotal margin yellowish or entirely black, median lobe without a series of spinelike projections laterally................................................................................ 5
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE8FFEFFF7E0B584F84FDB5" pageId="4" pageNumber="406" type="etymology">
<paragraph id="9E493695FFE8FFEFFF7E0B584F84FDB5" blockId="4.[151,1437,530,807]" pageId="4" pageNumber="406">
<emphasis id="AC82EA87FFE8FFEFFF7E0B584E3FFDD0" bold="true" box="[199,334,531,555]" pageId="4" pageNumber="406">Etymology.</emphasis>
The species name refers to the Qinlingshan Mts. in which all known localities of this species are situated.
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE8FFEFFF7E0B114B7DFCF9" pageId="4" pageNumber="406" type="biology_ecology">
<paragraph id="9E493695FFE8FFEFFF7E0B114B7DFCF9" blockId="4.[151,1437,530,807]" pageId="4" pageNumber="406">
<emphasis id="AC82EA87FFE8FFEFFF7E0B114E56FD88" bold="true" box="[199,295,602,627]" pageId="4" pageNumber="406">Biology.</emphasis>
Part of the
<typeStatus id="414D8837FFE8FFEFFE130B104EABFD88" box="[426,474,603,627]" pageId="4" pageNumber="406">type</typeStatus>
series was collected on wet granite rocks at the bottom of the waterfall in Fengyukou (
<figureCitation id="06CD2A10FFE8FFEFFF260B354F93FD6D" box="[159,226,638,662]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="4" pageNumber="406">Fig. 6</figureCitation>
) or in small hygropetric “streamlets” flowing over the granite cliffs of the Huashan Mts. near Huayin village (
<figureCitation id="06CD2A10FFE8FFEFFF260BE94F92FD41" box="[159,227,674,698]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[156,1435,193,1825]" captionTargetId="figure@2.[151,1436,193,1833]" captionTargetPageId="2" captionText="FIGURES 1 7. Laccobius (Glyptolaccobius) qinlingensis sp. nov. and its localities. 1 2: general habitus (1: dorsal view; 2: lateral view). 3 5: aedeagus (3: dorsal view; 4: lateral view; 5: ventral view). 6: waterfall at Fengyukou, type locality, site where the specimens were collected is marked by an arrow. 7: M. Balke collecting L. qinlingensis and Aspidytes wrasei at a granite cliff near Huayin village. Photos 6 7 by J. Hájek." httpUri="https://zenodo.org/record/219564/files/figure.png" pageId="4" pageNumber="406">Fig. 7</figureCitation>
) (in the latter case together with
<taxonomicName id="59F64D16FFE8FFEFFDEE0BE84B1BFD41" authority="Balke, Ribera &amp; Beutel, 2003" authorityName="Balke, Ribera &amp; Beutel" authorityYear="2003" box="[599,1130,674,699]" class="Insecta" family="Aspidytidae" genus="Aspidytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="406" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="AC82EA87FFE8FFEFFDEE0BE84C65FD40" box="[599,788,675,699]" italics="true" pageId="4" pageNumber="406">Aspidytes wrasei</emphasis>
Balke, Ribera &amp; Beutel, 2003
</taxonomicName>
(Adephaga:
<taxonomicName id="59F64D16FFE8FFEFFB420BE94AF6FD41" box="[1275,1415,674,698]" class="Insecta" family="Aspidytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="406" phylum="Arthropoda" rank="family">Aspidytidae</taxonomicName>
)). Second part of the
<typeStatus id="414D8837FFE8FFEFFED30B8C4EEBFD24" box="[362,410,711,735]" pageId="4" pageNumber="406">type</typeStatus>
series (in Fengyukou and in river valley near Huayin village) was collected on wet rocks at sides of the roads (the rocks were exposed during the construction of the road).
</paragraph>
</subSubSection>
<subSubSection id="D6EC651EFFE8FFEFFF7E0A454BA6FCDD" box="[199,1239,782,807]" pageId="4" pageNumber="406" type="distribution">
<paragraph id="9E493695FFE8FFEFFF7E0A454BA6FCDD" blockId="4.[151,1437,530,807]" box="[199,1239,782,807]" pageId="4" pageNumber="406">
<emphasis id="AC82EA87FFE8FFEFFF7E0A454E11FCDC" bold="true" box="[199,352,782,807]" pageId="4" pageNumber="406">Distribution.</emphasis>
Only known from three localities in the granite massif of the Qinlingshan Mts.
</paragraph>
</subSubSection>
</treatment>
</document>