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<document id="CE85B334266C6C152CE8B4006A3D81F4" ID-DOI="10.1080/00222933.2011.623358" ID-ISSN="1464-5262" ID-Zenodo-Dep="5199547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628903809132" checkinUser="felipe" docAuthor="Delicado, Diana, Machordom, Annie &amp; Ramos, Marian A." docDate="2011" docId="03C9996EFFCA2C22EB4389BDFDAAFA63" docLanguage="en" docName="JNatHist.46.1-2.25-89.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 46 (1 - 2)" docSource="http://dx.doi.org/10.1080/00222933.2011.623358" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Pseudamnicola (Corrosella) falkneri" docType="treatment" docVersion="3" lastPageNumber="44" masterDocId="FFF0E116FFC62C36EA0F8C1FFF9AFFD8" masterDocTitle="Underestimated diversity of hydrobiid snails. The case of Pseudamnicola (Corrosella) (Mollusca: Caenogastropoda: Hydrobiidae)" masterLastPageNumber="89" masterPageNumber="25" pageNumber="36" updateTime="1699055545266" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="66D4371AE18077B01B3DD43291EFD81F">Underestimated diversity of hydrobiid snails. The case of Pseudamnicola (Corrosella) (Mollusca: Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="81ADA776879C9980EBCD81829430215E">Delicado, Diana</mods:namePart>
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<mods:namePart id="FFDC272989187FB4E81ED51F5B1B88DA">Machordom, Annie</mods:namePart>
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<mods:namePart id="69C563C4478E1AA32B37963EB17BFAAC">Ramos, Marian A.</mods:namePart>
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<mods:title id="1734C8E7BB59593FBA818297DA6D522A">Journal of Natural History</mods:title>
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<mods:date id="27EF1F45367E6C57A251C9774F58AFAD">2011</mods:date>
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<mods:title id="D1E6B1008231BCA686F2894FAF93549D">J. Nat. Hist.</mods:title>
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<paragraph id="8BDF2878FFCA2C3AEB4389BDFC08FA63" blockId="12.[332,914,1442,1467]" box="[332,914,1442,1467]" pageId="12" pageNumber="36">
<heading id="D0979F14FFCA2C3AEB4389BDFC08FA63" box="[332,914,1442,1467]" centered="true" fontSize="36" level="2" pageId="12" pageNumber="36" reason="7">
<taxonomicName id="4C6053FBFFCA2C3AEB4389BDFC08FA63" ID-CoL="4NJVX" authority="(Boeters, 1970)" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[332,914,1442,1467]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="36" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFCA2C3AEB4389BDFD79FA63" bold="true" box="[332,739,1442,1467]" italics="true" pageId="12" pageNumber="36">Pseudamnicola (Corrosella) falkneri</emphasis>
(
<bibRefCitation id="EFF15589FFCA2C3AE8FD89BCFC11FA63" author="Boeters HD" box="[754,907,1443,1467]" pageId="12" pageNumber="36" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters, 1970</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A7BF3FFCA2C3AEA8089F7FC17F9D8" box="[143,909,1512,1536]" pageId="12" pageNumber="36" type="reference_group">
<paragraph id="8BDF2878FFCA2C3AEA8089F7FC17F9D8" blockId="12.[143,909,1512,1536]" box="[143,909,1512,1536]" pageId="12" pageNumber="36">
<taxonomicName id="4C6053FBFFCA2C3AEA8089F7FD8BF9D8" ID-CoL="YHNY" authority="(Boeters, 1970)" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[143,529,1512,1536]" class="Gastropoda" family="Hydrobiidae" genus="Corrosella" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="36" phylum="Mollusca" rank="species" species="falkneri">
<emphasis id="B914F46AFFCA2C3AEA8089F7FEC4F9D8" box="[143,350,1512,1536]" italics="true" pageId="12" pageNumber="36">Corrosella falkneri</emphasis>
(
<bibRefCitation id="EFF15589FFCA2C3AEB6289F7FD93F9D8" author="Boeters HD" box="[365,521,1512,1536]" pageId="12" pageNumber="36" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters, 1970</bibRefCitation>
)
</taxonomicName>
: p.6566, figs 1, 3, 5, 8, 10 and 11.
</paragraph>
</subSubSection>
<subSubSection id="C37A7BF3FFCA2C22EA808A24FDAAFA63" lastPageId="20" lastPageNumber="44" pageId="12" pageNumber="36" type="materials_examined">
<paragraph id="8BDF2878FFCA2C3AEA808A24FE84F98A" blockId="12.[143,286,1594,1618]" box="[143,286,1594,1618]" pageId="12" pageNumber="36">
<emphasis id="B914F46AFFCA2C3AEA808A24FE84F98A" box="[143,286,1594,1618]" italics="true" pageId="12" pageNumber="36">Type locality</emphasis>
</paragraph>
<paragraph id="8BDF2878FFCA2C38EA808A7DFF7DF972" blockId="12.[143,1103,1634,1690]" lastBlockId="14.[143,1104,1586,1706]" lastPageId="14" lastPageNumber="38" pageId="12" pageNumber="36">
<materialsCitation id="3B082225FFCA2C38EA808A7DFF79F972" collectingDate="1967-09-28" collectorName="Falkner &amp; The Cerro de la Virgen" country="Spain" county="Boeters" elevation="120" lastPageId="14" lastPageNumber="38" location="Granada" municipality="In" pageId="12" pageNumber="36" specimenCount="2">
According to its original description this species was collected by
<collectorName id="26954DAEFFCA2C3AE9C58A7DFBB8F9A2" box="[970,1058,1634,1658]" pageId="12" pageNumber="36">Falkner</collectorName>
on
<date id="FFDE0EB8FFCA2C3AEA808A9DFEF7F942" box="[143,365,1666,1690]" pageId="12" pageNumber="36" value="1967-09-28">
<collectingDate id="EF9AF750FFCA2C3AEA808A9DFEF7F942" box="[143,365,1666,1690]" pageId="12" pageNumber="36" value="1967-09-28">
28 September
<quantity id="4C98859DFFCA2C3AEB378A9DFEF7F942" box="[312,365,1666,1690]" metricMagnitude="1" metricUnit="m" metricValue="4.99618" pageId="12" pageNumber="36" unit="in" value="1967.0">1967</quantity>
</collectingDate>
</date>
in two streams, separated by
<quantity id="4C98859DFFCA2C3AE8DB8A9DFCBBF942" box="[724,801,1666,1690]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="36" unit="m" value="120.0">
<elevation id="004DCF4BFFCA2C3AE8DB8A9DFCBBF942" box="[724,801,1666,1690]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="36" unit="m" value="120.0">120 m</elevation>
</quantity>
, at “Cerro de la Virgen”,
<location id="8EBF7EA3FFC82C38EA808A2DFF6DF992" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFC82C38EA808A2DFF6DF992" box="[143,247,1586,1610]" country="Spain" county="Boeters" municipality="In" name="Granada" pageId="14" pageNumber="38">Granada</location>
,
<collectingCountry id="F37768E8FFC82C38EB0D8A2DFEDCF992" box="[258,326,1586,1610]" name="Spain" pageId="14" pageNumber="38">Spain</collectingCountry>
.
<location id="8EBF7EA3FFC82C38EB408A2DFEE1F992" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFC82C38EB408A2DFEE1F992" box="[335,379,1586,1610]" country="Spain" county="Boeters" municipality="In" name="The" pageId="14" pageNumber="38">The</location>
eastern flow, between the source and a watering place, is the
<typeStatus id="54DB96DAFFC82C38EE108A2CFBD5F993" box="[1055,1103,1587,1611]" pageId="14" pageNumber="38">type</typeStatus>
locality.
<collectorName id="26954DAEFFC82C38EAE38A4DFE73F9B2" box="[236,489,1618,1642]" pageId="14" pageNumber="38">
<location id="8EBF7EA3FFC82C38EAE38A4DFE73F9B2" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFC82C38EAE38A4DFE73F9B2" box="[236,489,1618,1642]" country="Spain" county="Boeters" municipality="In" name="The Cerro de la Virgen" pageId="14" pageNumber="38">The Cerro de la Virgen</location>
</collectorName>
is located
<locationDeviation id="9A6D315AFFC82C38E8538A4DFCEDF9B2" box="[604,887,1618,1642]" firstLocation="Galera" pageId="14" pageNumber="38" secondLocation="Orce">between Galera and Orce</locationDeviation>
above the southern edge of the
<location id="8EBF7EA3FFC82C38EB018A6DFEE4F952" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFC82C38EB018A6DFEE4F952" box="[270,382,1650,1674]" country="Spain" county="Boeters" municipality="In" name="river Orce" pageId="14" pageNumber="38">river Orce</location>
(
<bibRefCitation id="EFF15589FFC82C38EB848A6DFDBAF952" author="Boeters HD" box="[395,544,1650,1674]" pageId="14" pageNumber="38" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
).
<collectingMunicipality id="6BBBB202FFC82C38E83E8A6CFDD0F952" box="[561,586,1651,1674]" pageId="14" pageNumber="38">In</collectingMunicipality>
1988,
<collectingCounty id="62BE50F4FFC82C38E89D8A6CFD7DF952" box="[658,743,1651,1674]" pageId="14" pageNumber="38">Boeters</collectingCounty>
added UTM WG 47 to this
<typeStatus id="54DB96DAFFC82C38EE108A6CFBD5F953" box="[1055,1103,1651,1675]" pageId="14" pageNumber="38">type</typeStatus>
locality
</materialsCitation>
.
</paragraph>
<caption id="DF1F78F0FFCB2C3BEADC89FEFD82F997" ID-DOI="http://doi.org/10.5281/zenodo.5199557" ID-Zenodo-Dep="5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="13" pageNumber="37" startId="13.[211,278,1505,1527]" targetBox="[211,1170,169,1472]" targetPageId="13">
<paragraph id="8BDF2878FFCB2C3BEADC89FEFD82F997" blockId="13.[211,1172,1505,1615]" pageId="13" pageNumber="37">
Figure 5. Shells of
<taxonomicName id="4C6053FBFFCB2C3BEB9B89FEFD2AFA2F" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[404,688,1505,1527]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="37" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFCB2C3BEB9B89FEFD2AFA2F" box="[404,688,1505,1527]" italics="true" pageId="13" pageNumber="37">Pseudamnicola (C.) falkneri</emphasis>
</taxonomicName>
. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture.
</paragraph>
</caption>
<caption id="DF1F78F0FFC82C38EA808996FF53FA2F" ID-DOI="http://doi.org/10.5281/zenodo.5199559" ID-Zenodo-Dep="5199559" httpUri="https://zenodo.org/record/5199559/files/figure.png" pageId="14" pageNumber="38" startId="14.[143,210,1417,1439]" targetBox="[143,1102,168,1384]" targetPageId="14">
<paragraph id="8BDF2878FFC82C38EA808996FF53FA2F" blockId="14.[143,1104,1417,1527]" pageId="14" pageNumber="38">
Figure 6. Operculum and radula of
<taxonomicName id="4C6053FBFFC82C38EBF48996FC8DFA47" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[507,791,1417,1439]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="38" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFC82C38EBF48996FC8DFA47" box="[507,791,1417,1439]" italics="true" pageId="14" pageNumber="38">Pseudamnicola (C.) falkneri</emphasis>
</taxonomicName>
from La Armada spring, Orce, Granada (MNCN 15.05/49161). (A) Internal side of the operculum; (B): external side of the operculum; (C): radula; (D) rows of teeth of the radula; (E) central tooth; (F) outer marginal teeth.
</paragraph>
</caption>
<caption id="DF1F78F0FFC92C39EADC8A37FBA0F96B" ID-DOI="http://doi.org/10.5281/zenodo.5199561" ID-Zenodo-Dep="5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="15" pageNumber="39" startId="15.[211,278,1576,1598]" targetBox="[213,1167,164,1543]" targetPageId="15">
<paragraph id="8BDF2878FFC92C39EADC8A37FBA0F96B" blockId="15.[211,1172,1576,1715]" pageId="15" pageNumber="39">
Figure 7. Anatomy of
<taxonomicName id="4C6053FBFFC92C39EBCC8A37FD7CF9E6" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[451,742,1576,1598]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="39" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFC92C39EBCC8A37FD7CF9E6" box="[451,742,1576,1598]" italics="true" pageId="15" pageNumber="39">Pseudamnicola (C.) falkneri</emphasis>
</taxonomicName>
from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology.
</paragraph>
</caption>
<caption id="DF1F78F0FFD62C26EA80881EFC5AFBB7" ID-DOI="http://doi.org/10.5281/zenodo.5199563" ID-Zenodo-Dep="5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="16" pageNumber="40" startId="16.[143,210,1025,1047]" targetBox="[143,1102,169,992]" targetPageId="16">
<paragraph id="8BDF2878FFD62C26EA80881EFC5AFBB7" blockId="16.[143,1104,1025,1135]" pageId="16" pageNumber="40">
Figure 8. Shells of several populations of
<taxonomicName id="4C6053FBFFD62C26E845881EFCF4FBCF" baseAuthorityName="Boeters" baseAuthorityYear="1970" box="[586,878,1025,1047]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="40" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD62C26E845881EFCF4FBCF" box="[586,878,1025,1047]" italics="true" pageId="16" pageNumber="40">Pseudamnicola (C.) falkneri</emphasis>
</taxonomicName>
. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163).
</paragraph>
</caption>
<paragraph id="8BDF2878FFD62C26EA8088B0FEB2FB1F" blockId="16.[143,296,1199,1223]" box="[143,296,1199,1223]" pageId="16" pageNumber="40">
<emphasis id="B914F46AFFD62C26EA8088B0FEB2FB1F" box="[143,296,1199,1223]" italics="true" pageId="16" pageNumber="40">Type material</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD62C26EA8088C9FEE4FAF6" blockId="16.[143,1104,1238,1390]" pageId="16" pageNumber="40">
<materialsCitation id="3B082225FFD62C26EA8088C9FEA1FB36" box="[143,315,1238,1262]" collectionCode="SMF" pageId="16" pageNumber="40" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DB96DAFFD62C26EA8088C9FF60FB36" box="[143,250,1238,1262]" pageId="16" pageNumber="40" type="holotype">Holotype</typeStatus>
<collectionCode id="ED71B0BDFFD62C26EB0E88C9FEA1FB36" box="[257,315,1238,1262]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B082225FFD62C26EB4888C8FD33FB36" box="[327,681,1238,1263]" collectionCode="SMF, MP, RNHL" pageId="16" pageNumber="40" specimenCount="1" typeStatus="paratype">
<typeStatus id="54DB96DAFFD62C26EB4888C8FE2FFB37" box="[327,437,1239,1263]" pageId="16" pageNumber="40" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED71B0BDFFD62C26EBB388C9FE6CFB36" box="[444,502,1238,1262]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF</collectionCode>
,
<collectionCode id="ED71B0BDFFD62C26E80D88C8FDB6FB36" box="[514,556,1239,1262]" country="0" name="Mohonk Preserve, Inc." pageId="16" pageNumber="40">MP</collectionCode>
,
<collectionCode id="ED71B0BDFFD62C26E83888C8FD16FB36" box="[567,652,1239,1262]" pageId="16" pageNumber="40">RNHL</collectionCode>
, F
</materialsCitation>
,
<materialsCitation id="3B082225FFD62C26E8BB88C9FC81FB36" box="[692,795,1238,1262]" collectionCode="BOE" pageId="16" pageNumber="40" specimenCode="BOE 222" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBC68003FFD62C26E8BB88C9FC81FB36" box="[692,795,1238,1262]" collectionCode="BOE" pageId="16" pageNumber="40">BOE 222</specimenCode>
</materialsCitation>
et 223 (
<bibRefCitation id="EFF15589FFD62C26E97B88C9FB91FB36" author="Boeters HD" box="[884,1035,1238,1262]" pageId="16" pageNumber="40" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
). [
<collectionCode id="ED71B0BDFFD62C26EE2B88C8FBD5FB36" box="[1060,1103,1239,1262]" country="0" name="Mohonk Preserve, Inc." pageId="16" pageNumber="40">MP</collectionCode>
(=
<collectionCode id="ED71B0BDFFD62C26EAA288E9FE96FAD6" box="[173,268,1270,1294]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="40">MNHN</collectionCode>
),
<collectionCode id="ED71B0BDFFD62C26EB1088E8FEEBFAD6" box="[287,369,1271,1294]" pageId="16" pageNumber="40">RNHL</collectionCode>
(=
<collectionCode id="ED71B0BDFFD62C26EB9A88E9FE69FAD6" box="[405,499,1270,1294]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="40" type="Museum">RMNH</collectionCode>
), F (=
<collectionCode id="ED71B0BDFFD62C26E83488E9FD1CFAD6" box="[571,646,1270,1294]" pageId="16" pageNumber="40">FALK</collectionCode>
or Falkner, according to original writing in collection labels)].
</paragraph>
<paragraph id="8BDF2878FFD62C26EAB58928FD5DFAB6" blockId="16.[143,1104,1238,1390]" pageId="16" pageNumber="40">
<materialsCitation id="3B082225FFD62C26EAB58928FD59FAB6" collectionCode="MNHN, FALK, RMNH" pageId="16" pageNumber="40" specimenCode="SMF 219026, SMF 219027, BOE 222" specimenCount="2" typeStatus="holotype">
Types:
<typeStatus id="54DB96DAFFD62C26EB028929FEE2FA96" box="[269,376,1334,1358]" pageId="16" pageNumber="40" type="holotype">Holotype</typeStatus>
<specimenCode id="DBC68003FFD62C26EB8C8929FD86FA96" box="[387,540,1334,1358]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF 219026</specimenCode>
,
<typeStatus id="54DB96DAFFD62C26E8248928FD03FA97" box="[555,665,1335,1359]" pageId="16" pageNumber="40" type="paratype">Paratypes</typeStatus>
<specimenCode id="DBC68003FFD62C26E8AB8929FCA3FA96" box="[676,825,1334,1358]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF 219027</specimenCode>
/1, 219028/10,
<collectionCode id="ED71B0BDFFD62C26E9E18928FBD1FA96" box="[1006,1099,1335,1358]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="40">MNHN</collectionCode>
,
<collectionCode id="ED71B0BDFFD62C26EA808949FF47FAB6" box="[143,221,1366,1390]" pageId="16" pageNumber="40">FALK</collectionCode>
,
<collectionCode id="ED71B0BDFFD62C26EAE78948FEDFFAB6" box="[232,325,1367,1390]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="40" type="Museum">RMNH</collectionCode>
,
<specimenCode id="DBC68003FFD62C26EB5F8949FE2DFAB6" box="[336,439,1366,1390]" collectionCode="BOE" pageId="16" pageNumber="40">BOE 222</specimenCode>
and 223 (
<bibRefCitation id="EFF15589FFD62C26E82A8949FD26FAB6" author="Boeters HD" box="[549,700,1366,1390]" pageId="16" pageNumber="40" pagination="181 - 261" refId="ref22923" refString="Boeters HD. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal. Archiv Molluskenkd. 118 (4 / 6): 181 - 261." type="journal article" year="1988">Boeters 1988</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD62C26EA8089A4FEFBFA0B" blockId="16.[143,353,1467,1491]" box="[143,353,1467,1491]" pageId="16" pageNumber="40">
<emphasis id="B914F46AFFD62C26EA8089A4FEFBFA0B" box="[143,353,1467,1491]" italics="true" pageId="16" pageNumber="40">Material examined</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD62C27EA8089FCFC6AFEE7" blockId="16.[143,1104,1506,1690]" lastBlockId="17.[211,1172,167,1690]" lastPageId="17" lastPageNumber="41" pageId="16" pageNumber="40">
<emphasis id="B914F46AFFD62C26EA8089FCFEB6FA23" box="[143,300,1507,1531]" italics="true" pageId="16" pageNumber="40">Type material</emphasis>
.
<materialsCitation id="3B082225FFD62C26EB3089FCFC96F9E2" collectionCode="SMF, BOE" collectorName="Falkner" country="Grenada" location="Abfluss" municipality="Quellaustritts" pageId="16" pageNumber="40" specimenCode="SMF 219026, BOE 222, SMF 219028" specimenCount="67" typeStatus="holotype">
We examined a photograph of the
<typeStatus id="54DB96DAFFD62C26E8D589FDFCA4FA22" box="[730,830,1506,1530]" pageId="16" pageNumber="40" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBC68003FFD62C26E95F89FDFC7DFA22" box="[848,999,1506,1530]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF 219026</specimenCode>
) labelled as “
<location id="8EBF7EA3FFD62C26EAB48A1DFE89F9C2" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD62C26EAB48A1DFE89F9C2" box="[187,275,1538,1562]" country="Grenada" municipality="Quellaustritts" name="Abfluss" pageId="16" pageNumber="40">Abfluss</location>
des östlichen
<collectingMunicipality id="6BBBB202FFD62C26EBB48A1DFDCCF9C2" box="[443,598,1538,1562]" pageId="16" pageNumber="40">Quellaustritts</collectingMunicipality>
, Cerro de la Virgen,
<collectingCountry id="F37768E8FFD62C26E9408A1DFC22F9C2" box="[847,952,1538,1562]" name="Grenada" pageId="16" pageNumber="40">Granada</collectingCountry>
.
<collectorName id="26954DAEFFD62C26E9C98A1DFB84F9C2" box="[966,1054,1538,1562]" pageId="16" pageNumber="40">Falkner</collectorName>
leg. 28/9/
<specimenCount id="9D66E3F1FFD62C26EADF8A3DFE82F9E2" box="[208,280,1570,1594]" pageId="16" pageNumber="40" type="generic">67 ex.</specimenCount>
<specimenCode id="DBC68003FFD62C26EB2B8A3DFE0CF9E2" box="[292,406,1570,1594]" collectionCode="BOE" pageId="16" pageNumber="40">BOE 222</specimenCode>
”, five shells from
<specimenCode id="DBC68003FFD62C26E87A8A3DFC96F9E2" box="[629,780,1570,1594]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="40">SMF 219028</specimenCode>
</materialsCitation>
/
<materialsCitation id="3B082225FFD62C26E9178A3DFD4DF9A2" collectionCode="BOE, NHMW" country="Spain" location="Spanien" pageId="16" pageNumber="40" specimenCode="BOE 223, NHMW 107332" specimenCount="1" typeStatus="paratype">
10 labelled as “
<typeStatus id="54DB96DAFFD62C26E9DD8A3DFBD1F9E2" box="[978,1099,1570,1594]" pageId="16" pageNumber="40" type="paratype">Paratypen</typeStatus>
,
<collectingCountry id="F37768E8FFD62C26EA808A5DFF74F982" box="[143,238,1602,1626]" name="Spain" pageId="16" pageNumber="40">Spanien</collectingCountry>
:
<collectingCountry id="F37768E8FFD62C26EB0E8A5DFEF0F982" box="[257,362,1602,1626]" name="Grenada" pageId="16" pageNumber="40">Granada</collectingCountry>
: Cerro de la Virgen: W Quellenaustritt, G. Falkner 28.9.67 (
<specimenCode id="DBC68003FFD62C26EA988A7DFE93F9A2" box="[151,265,1634,1658]" collectionCode="BOE" pageId="16" pageNumber="40">BOE 223</specimenCode>
b)”, 10 shells from the
<specimenCode id="DBC68003FFD62C26E8148A7CFD4DF9A2" box="[539,727,1634,1658]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="40" type="Museum">NHMW 107332</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD62C27E8EC8A7DFB8FFF67" collectingDate="1967-09-28" collectionCode="BOE, NHMW" collectorName="W. Balsa" country="Grenada" lastPageId="17" lastPageNumber="41" location="Cerro de la Virgen" pageId="16" pageNumber="40" specimenCode="BOE 223, NHMW 107331" specimenCount="1" typeStatus="paratype">
labelled as “
<typeStatus id="54DB96DAFFD62C26E97D8A7DFC72F9A2" box="[882,1000,1634,1658]" pageId="16" pageNumber="40" type="paratype">Paratypen</typeStatus>
ex
<specimenCode id="DBC68003FFD62C26EE188A7DFF23F942" collectionCode="BOE" pageId="16" pageNumber="40">BOE 223</specimenCode>
b.
<taxonomicName id="4C6053FBFFD62C26EADB8A9DFDC3F942" authority="Quellenaustritt" authorityName="Quellenaustritt" box="[212,601,1666,1690]" class="Gastropoda" family="Hydrobiidae" genus="Corrosella" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="40" phylum="Mollusca" rank="species" species="falkneri">
<emphasis id="B914F46AFFD62C26EADB8A9DFE3FF942" box="[212,421,1666,1690]" italics="true" pageId="16" pageNumber="40">Corrosella falkneri</emphasis>
Quellenaustritt
</taxonomicName>
der
<collectorName id="26954DAEFFD62C26E8808A9CFD6DF942" box="[655,759,1666,1690]" pageId="16" pageNumber="40">W. Balsa</collectorName>
,
<location id="8EBF7EA3FFD62C26E90B8A9DFC47F942" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD62C26E90B8A9DFC47F942" box="[772,989,1666,1690]" country="Grenada" name="Cerro de la Virgen" pageId="16" pageNumber="40">Cerro de la Virgen</location>
,
<collectingCountry id="F37768E8FFD62C26E9E58A9DFBD5F942" box="[1002,1103,1666,1690]" name="Grenada" pageId="16" pageNumber="40">Granada</collectingCountry>
FALKNER leg
<date id="FFDE0EB8FFD72C27EB848CB8FD94FF67" box="[395,526,167,191]" pageId="17" pageNumber="41" value="1967-09-28">
<collectingDate id="EF9AF750FFD72C27EB848CB8FD94FF67" box="[395,526,167,191]" pageId="17" pageNumber="41" value="1967-09-28">28 IX 1967</collectingDate>
</date>
”, four shells (dry specimens)
<specimenCode id="DBC68003FFD72C27E9528CB7FB8FFF67" box="[861,1045,167,191]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="41" type="Museum">NHMW 107331</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD72C27EE128CB8FC71FEE7" collectingDate="1967-09-28" collectingDateMax="2011-03-20" collectingDateMin="1967-09-28" collectionCode="BOE, MNCN" collectorName="Falkner" country="Grenada" county="Rio Orce" location="Cerro de la Virgen" municipality="Orce" pageId="17" pageNumber="41" specimenCode="BOE 223, MNCN 15.05" specimenCount="1" typeStatus="paratype">
labelled as “
<typeStatus id="54DB96DAFFD72C27EAD28CD8FEC9FF07" box="[221,339,199,223]" pageId="17" pageNumber="41" type="paratype">Paratypen</typeStatus>
ex
<specimenCode id="DBC68003FFD72C27EB738CD8FE7FFF07" box="[380,485,199,223]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
b
<taxonomicName id="4C6053FBFFD72C27EBF48CD7FB0EFF07" authority="Westl. Quellenaustritt" authorityName="Westl. Quellenaustritt" box="[507,1172,199,224]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="41" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD72C27EBF48CD7FC0EFF38" box="[507,916,200,224]" italics="true" pageId="17" pageNumber="41">Pseudamnicola (Corrosella) falkneri</emphasis>
Westl. Quellenaustritt
</taxonomicName>
am
<location id="8EBF7EA3FFD72C27EAF18CF8FE55FF27" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD72C27EAF18CF8FE55FF27" box="[254,463,231,255]" country="Grenada" county="Rio Orce" municipality="Orce" name="Cerro de la Virgen" pageId="17" pageNumber="41">Cerro de la Virgen</location>
zw.
<location id="8EBF7EA3FFD72C27E80F8CF8FDD7FF27" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD72C27E80F8CF8FDD7FF27" box="[512,589,231,255]" country="Grenada" county="Rio Orce" municipality="Orce" name="Galera" pageId="17" pageNumber="41">Galera</location>
u.
<collectingMunicipality id="6BBBB202FFD72C27E87D8CF8FD32FF27" box="[626,680,231,255]" pageId="17" pageNumber="41">Orce</collectingMunicipality>
oberh. des Süduffers der Vega d.
<collectingCounty id="62BE50F4FFD72C27EE298CF8FB15FF27" box="[1062,1167,231,255]" pageId="17" pageNumber="41">Rio Orce</collectingCounty>
,
<collectingCountry id="F37768E8FFD72C27EADC8D18FEA2FEC7" box="[211,312,263,287]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
leg.
<collectorName id="26954DAEFFD72C27EB618D18FE5CFEC7" box="[366,454,263,287]" pageId="17" pageNumber="41">Falkner</collectorName>
<date id="FFDE0EB8FFD72C27EBC28D18FDA5FEC7" box="[461,575,263,287]" pageId="17" pageNumber="41" value="1967-09-28">
<collectingDate id="EF9AF750FFD72C27EBC28D18FDA5FEC7" box="[461,575,263,287]" pageId="17" pageNumber="41" value="1967-09-28">28.9.1967</collectingDate>
</date>
”, 15 shells (most of them juveniles) recently collected at 223b (D.M., N.M. and E.M.,
<date id="FFDE0EB8FFD72C27E8328D38FD73FEE7" box="[573,745,295,319]" pageId="17" pageNumber="41" value="2011-03-20">
<collectingDate id="EF9AF750FFD72C27E8328D38FD73FEE7" box="[573,745,295,319]" pageId="17" pageNumber="41" value="2011-03-20">20 March 2011</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD72C27E8FB8D38FC0AFEE7" box="[756,912,295,319]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN 15.05</specimenCode>
/49430)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD72C27EAF18D58FEB1FE26" blockId="17.[211,1172,167,1690]" pageId="17" pageNumber="41">
All published papers (
<bibRefCitation id="EFF15589FFD72C27EBF68D58FD0AFE87" author="Boeters HD" box="[505,656,327,351]" pageId="17" pageNumber="41" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
,
<bibRefCitation id="EFF15589FFD72C27E8928D58FD49FE87" author="Boeters HD" box="[669,723,327,351]" pageId="17" pageNumber="41" pagination="181 - 261" refId="ref22923" refString="Boeters HD. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal. Archiv Molluskenkd. 118 (4 / 6): 181 - 261." type="journal article" year="1988">1988</bibRefCitation>
) mentioned the existence of more than one sample collected at locality 223. The presence of material labelled as “
<collectionCode id="ED71B0BDFFD72C27EE018D78FBD3FEA7" box="[1038,1097,359,383]" pageId="17" pageNumber="41">BOE</collectionCode>
223b” in
<collectionCode id="ED71B0BDFFD72C27EAFC8D99FEB4FE46" box="[243,302,390,414]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="17" pageNumber="41">SMF</collectionCode>
and as “ex
<collectionCode id="ED71B0BDFFD72C27EBB28D99FE6FFE46" box="[445,501,390,414]" pageId="17" pageNumber="41">BOE</collectionCode>
223b” in
<collectionCode id="ED71B0BDFFD72C27E8638D98FD56FE46" box="[620,716,391,414]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="41" type="Museum">NHMW</collectionCode>
alerted us to the existence of a sample 223a. Boeters (personal communication) kindly clarified that the description of
<taxonomicName id="4C6053FBFFD72C27EE4B8DB8FEB2FE07" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="41" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD72C27EE4B8DB8FEB2FE07" italics="true" pageId="17" pageNumber="41">P. (C.) falkneri</emphasis>
</taxonomicName>
was based on specimens of three samples received from Gerhard Falkner as follows:
</paragraph>
<paragraph id="8BDF2878FFD72C27EAF18E18FBBBFD46" blockId="17.[211,1172,167,1690]" pageId="17" pageNumber="41">
-
<materialsCitation id="3B082225FFD72C27EB1B8E19FD1BFD86" collectionCode="BOE" pageId="17" pageNumber="41" specimenCode="BOE 222" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBC68003FFD72C27EB1B8E19FE1BFDC6" box="[276,385,518,542]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 222</specimenCode>
is the type locality (“écoulement oriental”, according to Falkner “Abfluss des östlichen Quellaustritts” [flow from the eastern spring emergence]). Type material from this sample:
<typeStatus id="54DB96DAFFD72C27E8158E59FD1BFD86" box="[538,641,582,606]" pageId="17" pageNumber="41" type="holotype">holotype</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B082225FFD72C27E89D8E59FC35FD87" box="[658,943,582,607]" collectionCode="SMF" pageId="17" pageNumber="41" specimenCode="SMF 219026" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBC68003FFD72C27E89D8E59FCB7FD86" box="[658,813,582,606]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="17" pageNumber="41">SMF 219026</specimenCode>
,
<typeStatus id="54DB96DAFFD72C27E9318E58FC35FD87" box="[830,943,583,607]" pageId="17" pageNumber="41" type="paratype">paratypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B082225FFD72C27E9B08E59FECEFDA5" collectionCode="SMF" pageId="17" pageNumber="41" specimenCode="SMF 219027" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBC68003FFD72C27E9B08E59FBCDFD86" box="[959,1111,582,606]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="17" pageNumber="41">SMF 219027</specimenCode>
/
<quantity id="4C98859DFFD72C27EE6C8E59FB09FD86" box="[1123,1171,582,606]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="41" unit="in" value="1.0">1 in</quantity>
addition to
</materialsCitation>
<collectionCode id="ED71B0BDFFD72C27EB528E79FE0FFDA6" box="[349,405,614,638]" pageId="17" pageNumber="41">BOE</collectionCode>
collection (Boeters personal communication) that comprises
<materialsCitation id="3B082225FFD72C27EE548E79FB87FD46" collectionCode="BOE" pageId="17" pageNumber="41" specimenCode="BOE 0222, BOE 2629" specimenCount="222" typeStatus="paratype">
<collectionCode id="ED71B0BDFFD72C27EE548E79FB09FDA6" box="[1115,1171,614,638]" pageId="17" pageNumber="41">BOE</collectionCode>
<specimenCount id="9D66E3F1FFD72C27EADC8E99FEEEFD46" box="[211,372,646,670]" pageId="17" pageNumber="41" type="generic">222 specimens</specimenCount>
either in alcohol (now
<specimenCode id="DBC68003FFD72C27E8788E99FD76FD46" box="[631,748,646,670]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 0222</specimenCode>
) or shells (now
<specimenCode id="DBC68003FFD72C27E9938E99FB8FFD46" box="[924,1045,646,670]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 2629</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD72C27EAF18EB9FD8AFC85" blockId="17.[211,1172,167,1690]" pageId="17" pageNumber="41">
-
<materialsCitation id="3B082225FFD72C27EB1E8EBAFDEFFCC5" collectionCode="BOE" collectorName="However G. Falkner" pageId="17" pageNumber="41" specimenCode="BOE 223" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBC68003FFD72C27EB1E8EBAFEE1FD65" box="[273,379,677,701]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
is the second original locality (“écoulement occidentale”).
<collectorName id="26954DAEFFD72C27EE208EB9FED4FD05" pageId="17" pageNumber="41">However G. Falkner</collectorName>
distinguished between “Acequia unterhalb der westlichen Balsa” [acequia downhill of the western balsa] (
<specimenCode id="DBC68003FFD72C27E8488EFAFD2FFD25" box="[583,693,741,765]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
a) and “Quellaustritt” [spring emergence] (
<specimenCode id="DBC68003FFD72C27EAD48F1AFED3FCC5" box="[219,329,773,797]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
b).
<typeStatus id="54DB96DAFFD72C27EB628F19FE41FCC6" box="[365,475,774,798]" pageId="17" pageNumber="41" type="paratype">Paratypes</typeStatus>
at present in
</materialsCitation>
<materialsCitation id="3B082225FFD72C27E8718F1AFD96FC85" collectionCode="BOE" pageId="17" pageNumber="41" specimenCode="BOE 223, BOE 0223, BOE 2630" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED71B0BDFFD72C27E8718F1AFD2CFCC5" box="[638,694,773,797]" pageId="17" pageNumber="41">BOE</collectionCode>
collection (Boeters personal communication) comprise animals in alcohol (from
<specimenCode id="DBC68003FFD72C27E8928F3AFC92FCE5" box="[669,776,805,829]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
a, now
<specimenCode id="DBC68003FFD72C27E9578F3AFC54FCE5" box="[856,974,805,829]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 0223</specimenCode>
) and shells (from
<specimenCode id="DBC68003FFD72C27EADC8F5AFEA6FC85" box="[211,316,837,861]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 223</specimenCode>
b, now
<specimenCode id="DBC68003FFD72C27EB838F5AFD9EFC85" box="[396,516,837,861]" collectionCode="BOE" pageId="17" pageNumber="41">BOE 2630</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD72C27EAF18F7AFBB7F942" blockId="17.[211,1172,167,1690]" pageId="17" pageNumber="41">
Neither the papers by
<bibRefCitation id="EFF15589FFD72C27EBFD8F79FD17FCA5" author="Boeters HD" box="[498,653,869,893]" pageId="17" pageNumber="41" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters (1970</bibRefCitation>
,
<bibRefCitation id="EFF15589FFD72C27E8988F7AFD57FCA5" author="Boeters HD" box="[663,717,869,893]" pageId="17" pageNumber="41" pagination="181 - 261" refId="ref22923" refString="Boeters HD. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal. Archiv Molluskenkd. 118 (4 / 6): 181 - 261." type="journal article" year="1988">1988</bibRefCitation>
) nor the labels of the collection materials mention the names of the springs, and it was therefore difficult to identify the precise location of the
<typeStatus id="54DB96DAFFD72C27EB8F8FBAFE2AFC65" box="[384,432,933,957]" pageId="17" pageNumber="41">type</typeStatus>
localities. Boeters (personal communication) provided additional data that have allowed us to locate them. The exact location of the Boeters samples is: (European Datum 1950):
<collectionCode id="ED71B0BDFFD72C27E81C8FFBFDD1FC24" box="[531,587,996,1020]" pageId="17" pageNumber="41">BOE</collectionCode>
222, UTM: 30S 543001 4175819,
<collectionCode id="ED71B0BDFFD72C27E9C78FFBFB9AFC24" box="[968,1024,996,1020]" pageId="17" pageNumber="41">BOE</collectionCode>
223a, UTM: 30S 542838 4175797,
<collectionCode id="ED71B0BDFFD72C27EBC4881BFD99FBC4" box="[459,515,1028,1052]" pageId="17" pageNumber="41">BOE</collectionCode>
223b, UTM: 30S 542840 4175779 (all explored by D.M., N.M. and E.M.
<date id="FFDE0EB8FFD72C27EB9D883BFDDCFBE4" box="[402,582,1060,1084]" pageId="17" pageNumber="41" value="2011-03-20">20 March 2011</date>
). Water flows near Cerro de la Virgen are now dry or destroyed and only a few empty shells were found at locality 223b (D. Moreno, personal communication). The area was extensively explored and
<taxonomicName id="4C6053FBFFD72C27E9E2887BFE27FB44" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="41" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD72C27E9E2887BFE27FB44" italics="true" pageId="17" pageNumber="41">Pseudamnicola (Corrosella) falkneri</emphasis>
</taxonomicName>
was found in five springs at the town of Orce close to Orce river. We accordingly based our anatomical descriptions and molecular data on these specimens. All the material is deposited at the
<collectionCode id="ED71B0BDFFD72C27E89A88DCFD74FB03" box="[661,750,1219,1243]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
except for some specimens mentioned below that have been deposited at the
<collectionCode id="ED71B0BDFFD72C27E89288FBFD67FB23" box="[669,765,1252,1275]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="41" type="Museum">NHMW</collectionCode>
. Localities and collection numbers are as follows: La Armada spring, Orce,
<collectingCountry id="F37768E8FFD72C27E8CD891CFCB0FAC3" box="[706,810,1283,1307]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
, UTM: 30S 0546272/4176139, JM.B.,
<date id="FFDE0EB8FFD72C27EB23893CFE56FAE3" box="[300,460,1315,1340]" pageId="17" pageNumber="41" value="2008-05-21">21 May 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27EBD2893CFDACFAE3" box="[477,566,1315,1339]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49161 (96% ethanol,
<figureCitation id="135B34FDFFD72C27E986893CFB99FAE3" box="[905,1027,1315,1339]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="17" pageNumber="41">Figure 8D</figureCitation>
and ESEM preparation,
<figureCitation id="135B34FDFFD72C27EB68895CFDBBFA83" box="[359,545,1347,1371]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="17" pageNumber="41">Figure 5A,DG</figureCitation>
,),
<collectionCode id="ED71B0BDFFD72C27E834895CFD0EFA83" box="[571,660,1347,1371]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
/ADN 3484734851 (96% ethanol),
<collectionCode id="ED71B0BDFFD72C27EE3C895BFB09FA83" box="[1075,1171,1348,1371]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="41" type="Museum">NHMW</collectionCode>
92420/
<specimenCount id="9D66E3F1FFD72C27EB2D897CFEE5FAA3" box="[290,383,1379,1403]" pageId="17" pageNumber="41" type="generic">10 spec.</specimenCount>
(70% ethanol); JM.B.,
<date id="FFDE0EB8FFD72C27E89E897CFCC9FAA3" box="[657,851,1379,1403]" pageId="17" pageNumber="41" value="2008-10-30">30 October 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27E96D897CFC21FAA3" box="[866,955,1379,1403]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49162 (70% ethanol); Palo spring, Orce,
<collectingCountry id="F37768E8FFD72C27E818899CFDE5FA43" box="[535,639,1411,1435]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
, UTM: 30S 0545712/4175851, JM.B.,
<date id="FFDE0EB8FFD72C27EE33899CFE91FA63" pageId="17" pageNumber="41" value="2008-05-21">21 May 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27EB1989BCFEF5FA63" box="[278,367,1443,1467]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49163 (96% ethanol,
<figureCitation id="135B34FDFFD72C27E8A889BCFC8CFA63" box="[679,790,1443,1467]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="17" pageNumber="41">Figure 8F</figureCitation>
) and
<collectionCode id="ED71B0BDFFD72C27E95989BCFC35FA63" box="[854,943,1443,1467]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
/ADN 3485234857 (96% ethanol); JM.B.,
<date id="FFDE0EB8FFD72C27EBD589DDFD01FA02" box="[474,667,1474,1498]" pageId="17" pageNumber="41" value="2008-10-30">30 October 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27E8A689DDFC98FA02" box="[681,770,1474,1498]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49164 (70% ethanol); Zarza spring, Orce,
<collectingCountry id="F37768E8FFD72C27EB6189FDFE4CFA22" box="[366,470,1506,1530]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
, UTM: 30S 0544907/4176049, JM.B.,
<date id="FFDE0EB8FFD72C27E99B89FDFBB7FA22" box="[916,1069,1506,1531]" pageId="17" pageNumber="41" value="2008-05-21">21 May 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27EE3589FDFB09FA22" box="[1082,1171,1506,1530]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49165 (96% ethanol) and
<collectionCode id="ED71B0BDFFD72C27E8418A1DFD3DF9C2" box="[590,679,1538,1562]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
/ADN 3485834859 (96% ethanol); La Pi spring, Orce,
<collectingCountry id="F37768E8FFD72C27EB618A3DFE4CF9E2" box="[366,470,1570,1594]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
, UTM: 30S 0544632/4175976, JM.B.,
<date id="FFDE0EB8FFD72C27E99B8A3DFBB7F9E2" box="[916,1069,1570,1595]" pageId="17" pageNumber="41" value="2008-05-21">21 May 2008</date>
,
<collectionCode id="ED71B0BDFFD72C27EE358A3DFB09F9E2" box="[1082,1171,1570,1594]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49166 (96% ethanol) and
<collectionCode id="ED71B0BDFFD72C27E8558A5DFD29F982" box="[602,691,1602,1626]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
/ADN 3486034861 (96% ethanol); Las Mimbreras spring, Orce,
<collectingCountry id="F37768E8FFD72C27E80E8A7DFDF3F9A2" box="[513,617,1634,1658]" name="Grenada" pageId="17" pageNumber="41">Granada</collectingCountry>
, UTM: 30S 0545875/4175182, JM.B.,
<collectionCode id="ED71B0BDFFD72C27EE358A7DFB09F9A2" box="[1082,1171,1634,1658]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
15.05/49167 (96% ethanol) and
<collectionCode id="ED71B0BDFFD72C27E8348A9DFD0EF942" box="[571,660,1666,1690]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="17" pageNumber="41" type="Museum">MNCN</collectionCode>
/ADN: 3486234863 (96% ethanol).
</paragraph>
<paragraph id="8BDF2878FFD42C24EA808CB7FBDBFD46" blockId="18.[143,1104,167,670]" pageId="18" pageNumber="42">
<emphasis id="B914F46AFFD42C24EA808CB7FE4AFF18" box="[143,464,168,192]" italics="true" pageId="18" pageNumber="42">Other material examined</emphasis>
.
<materialsCitation id="3B082225FFD42C24EBF98CB8FBA7FD46" accessionNumber="ADN 34877" collectingDate="1994-10-12" collectingDateMax="2009-12" collectingDateMin="1994-10-12" collectionCode="MNCN" collectorName="Dos Canos &amp; Castril &amp; Diana Delicado &amp; La Erra &amp; La Dehesa &amp; Albacete &amp; Letur &amp; Albacete, D. M. &amp; Fuente Garcia &amp; Cordovilla &amp; Oria &amp; Almeria" country="Grenada" county="Castril" location="Tubos" municipality="Castril" pageId="18" pageNumber="42" specimenCode="MNCN 15.05, ADN 34894-34898, ADN 34965-34966" specimenCount="1">
<location id="8EBF7EA3FFD42C24EBF98CB8FDA6FF67" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD42C24EBF98CB8FDA6FF67" box="[502,572,167,191]" country="Grenada" county="Castril" municipality="Castril" name="Tubos" pageId="18" pageNumber="42">Tubos</location>
spring,
<location id="8EBF7EA3FFD42C24E8CA8CB8FC8EFF67" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD42C24E8CA8CB8FC8EFF67" box="[709,788,167,191]" country="Grenada" county="Castril" municipality="Castril" name="Castril" pageId="18" pageNumber="42">Castril</location>
,
<collectingCountry id="F37768E8FFD42C24E9398CB8FC04FF67" box="[822,926,167,191]" name="Grenada" pageId="18" pageNumber="42">Granada</collectingCountry>
, UTM: 30S 0520451/4185742, JM.B.,
<date id="FFDE0EB8FFD42C24EBCC8CD8FDFEFF07" box="[451,612,199,224]" pageId="18" pageNumber="42" value="2008-05-21">
<collectingDate id="EF9AF750FFD42C24EBCC8CD8FDFEFF07" box="[451,612,199,224]" pageId="18" pageNumber="42" value="2008-05-21">21 May 2008</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E87A8CD8FC8DFF07" box="[629,791,199,223]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49168 (96% ethanol) and
<collectionCode id="ED71B0BDFFD42C24EA808CF8FF72FF27" box="[143,232,231,255]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/ADN: 3486434873 (96% ethanol); spring in
<location id="8EBF7EA3FFD42C24E8F38CF8FCE0FF27" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD42C24E8F38CF8FCE0FF27" box="[764,890,231,255]" country="Grenada" county="Castril" municipality="Castril" name="Pontezuela" pageId="18" pageNumber="42">Pontezuela</location>
,
<collectingMunicipality id="6BBBB202FFD42C24E9888CF8FC4CFF27" box="[903,982,231,255]" pageId="18" pageNumber="42">Castril</collectingMunicipality>
,
<collectingCountry id="F37768E8FFD42C24E9EC8CF8FBD1FF27" box="[995,1099,231,255]" name="Grenada" pageId="18" pageNumber="42">Granada</collectingCountry>
, UTM: 30S 0519553/4184431, JM.B.,
<date id="FFDE0EB8FFD42C24E8568D18FD65FEC7" box="[601,767,263,288]" pageId="18" pageNumber="42" value="2008-05-21">
<collectingDate id="EF9AF750FFD42C24E8568D18FD65FEC7" box="[601,767,263,288]" pageId="18" pageNumber="42" value="2008-05-21">21 May 2008</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E91C8D18FC2DFEC7" box="[787,951,263,287]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49169 (96% ethanol) and
<collectionCode id="ED71B0BDFFD42C24EB298D38FEE5FEE7" box="[294,383,295,319]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/ADN: 3487434876 (96% ethanol); spring in
<collectingCounty id="62BE50F4FFD42C24E9868D38FC42FEE7" box="[905,984,295,319]" pageId="18" pageNumber="42">Castril</collectingCounty>
,
<collectingCountry id="F37768E8FFD42C24E9EC8D38FBD1FEE7" box="[995,1099,295,319]" name="Grenada" pageId="18" pageNumber="42">Granada</collectingCountry>
, UTM: 30SWH2087, JM.B.,
<date id="FFDE0EB8FFD42C24EBDF8D58FD1BFE87" box="[464,641,327,351]" pageId="18" pageNumber="42" value="2006-08-26">
<collectingDate id="EF9AF750FFD42C24EBDF8D58FD1BFE87" box="[464,641,327,351]" pageId="18" pageNumber="42" value="2006-08-26">26 August 2006</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E8828D58FCB3FE87" box="[653,809,327,351]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49170 (96% ethanol) and
<collectionCode id="ED71B0BDFFD42C24EA808D78FF72FEA7" box="[143,232,359,383]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/ADN 34877 (96% ethanol);
<collectorName id="26954DAEFFD42C24E8498D78FD53FEA7" box="[582,713,359,383]" pageId="18" pageNumber="42">Dos Caños</collectorName>
spring,
<collectorName id="26954DAEFFD42C24E9218D78FCE7FEA7" box="[814,893,359,383]" pageId="18" pageNumber="42">Castril</collectorName>
,
<collectingCountry id="F37768E8FFD42C24E9818D78FC6CFEA7" box="[910,1014,359,383]" name="Grenada" pageId="18" pageNumber="42">Granada</collectingCountry>
, UTM: 30S 0520478/4185772,
<collectorName id="26954DAEFFD42C24EB948D98FE4EFE46" box="[411,468,391,414]" name="Diana Delicado" pageId="18" pageNumber="42">D.D.</collectorName>
,
<date id="FFDE0EB8FFD42C24EBEB8D99FD3DFE46" box="[484,679,390,414]" pageId="18" pageNumber="42" value="2008-10-13">
<collectingDate id="EF9AF750FFD42C24EBEB8D99FD3DFE46" box="[484,679,390,414]" pageId="18" pageNumber="42" value="2008-10-13">13 October 2008</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E8B98D99FCCCFE46" box="[694,854,390,414]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49171 (70% ethanol,
<figureCitation id="135B34FDFFD42C24EA808DB9FE90FE66" box="[143,266,422,446]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="18" pageNumber="42">Figure 8A</figureCitation>
, and ESEM preparation,
<figureCitation id="135B34FDFFD42C24E84D8DB9FD23FE66" box="[578,697,422,446]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figure 5B</figureCitation>
) and
<collectionCode id="ED71B0BDFFD42C24E90B8DB9FCC7FE66" box="[772,861,422,446]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/ADN: 3487834893 (frozen material);
<collectorName id="26954DAEFFD42C24EB568DD8FE2BFE06" box="[345,433,454,478]" pageId="18" pageNumber="42">La Errá</collectorName>
spring,
<collectorName id="26954DAEFFD42C24E8038DD8FD12FE06" box="[524,648,454,478]" pageId="18" pageNumber="42">La Dehesa</collectorName>
,
<collectorName id="26954DAEFFD42C24E89C8DD9FD62FE06" box="[659,760,454,478]" pageId="18" pageNumber="42">Albacete</collectorName>
, UTM: 30S 0572400/4242458,
<collectorName id="26954DAEFFD42C24EA808DF8FF52FE26" box="[143,200,487,510]" name="Diana Delicado" pageId="18" pageNumber="42">D.D.</collectorName>
,
<date id="FFDE0EB8FFD42C24EAD78DF9FE16FE26" box="[216,396,486,510]" pageId="18" pageNumber="42" value="2008-03-30">
<collectingDate id="EF9AF750FFD42C24EAD78DF9FE16FE26" box="[216,396,486,510]" pageId="18" pageNumber="42" value="2008-03-30">30 March 2008</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24EB948DF9FDA0FE26" box="[411,570,486,510]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49172 (70% ethanol and ESEM preparation,
<figureCitation id="135B34FDFFD42C24EA808E19FE90FDC6" box="[143,266,518,542]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figure 5C</figureCitation>
) and
<collectionCode id="ED71B0BDFFD42C24EB548E19FE2EFDC6" box="[347,436,518,542]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/
<specimenCode id="DBC68003FFD42C24EBCF8E19FD38FDC6" box="[448,674,518,542]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">ADN 3489434898</specimenCode>
(frozen material); stream in
<collectorName id="26954DAEFFD42C24EE038E18FBD6FDC6" box="[1036,1100,519,542]" pageId="18" pageNumber="42">Letur</collectorName>
,
<collectorName id="26954DAEFFD42C24EA808E39FEA4FDE6" box="[143,318,550,574]" pageId="18" pageNumber="42">Albacete, D.M.</collectorName>
,
<date id="FFDE0EB8FFD42C24EB458E39FD9FFDE6" box="[330,517,550,574]" pageId="18" pageNumber="42" value="1994-10-12">
<collectingDate id="EF9AF750FFD42C24EB458E39FD9FFDE6" box="[330,517,550,574]" pageId="18" pageNumber="42" value="1994-10-12">12 October 1994</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E81F8E39FD36FDE6" box="[528,684,550,574]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49173 (70% ethanol);
<collectorName id="26954DAEFFD42C24E9A28E38FBD5FDE6" box="[941,1103,550,574]" pageId="18" pageNumber="42">Fuente García</collectorName>
stream in
<collectorName id="26954DAEFFD42C24EB038E59FE1DFD86" box="[268,391,582,606]" pageId="18" pageNumber="42">Cordovilla</collectorName>
,
<collectorName id="26954DAEFFD42C24EB968E59FDCAFD86" box="[409,592,582,606]" pageId="18" pageNumber="42">Albacete, D.M.</collectorName>
,
<date id="FFDE0EB8FFD42C24E86D8E59FC9EFD86" box="[610,772,582,607]" pageId="18" pageNumber="42" value="1997-05-18">
<collectingDate id="EF9AF750FFD42C24E86D8E59FC9EFD86" box="[610,772,582,607]" pageId="18" pageNumber="42" value="1997-05-18">18 May 1997</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24E9198E59FC23FD86" box="[790,953,582,606]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49174 (70% ethanol);
<collectorName id="26954DAEFFD42C24EB0C8E79FEA3FDA6" box="[259,313,614,638]" pageId="18" pageNumber="42">Oria</collectorName>
,
<collectorName id="26954DAEFFD42C24EB448E79FE30FDA6" box="[331,426,614,638]" pageId="18" pageNumber="42">Almería</collectorName>
, UTM: 30S 0570250/4150420, JM. B.,
<date id="FFDE0EB8FFD42C24E99E8E79FBD1FDA6" box="[913,1099,614,638]" pageId="18" pageNumber="42" value="2009-12">
<collectingDate id="EF9AF750FFD42C24E99E8E79FBD1FDA6" box="[913,1099,614,638]" pageId="18" pageNumber="42" value="2009-12">December 2009</collectingDate>
</date>
,
<specimenCode id="DBC68003FFD42C24EA808E99FEB1FD46" box="[143,299,646,670]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN 15.05</specimenCode>
/49198 (96% ethanol) and
<collectionCode id="ED71B0BDFFD42C24E8588E99FD2AFD46" box="[599,688,646,670]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">MNCN</collectionCode>
/
<specimenCode id="DBC68003FFD42C24E8B38E99FC0FFD46" box="[700,917,646,670]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="42" type="Museum">ADN 3496534966</specimenCode>
(96% ethanol)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD42C24EA808ED7FDB8FD38" blockId="18.[143,546,712,736]" box="[143,546,712,736]" pageId="18" pageNumber="42">
<emphasis id="B914F46AFFD42C24EA808ED7FDB8FD38" box="[143,546,712,736]" italics="true" pageId="18" pageNumber="42">Material examined for morphometry</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD42C24EA808EEEFDEAFC3F" blockId="18.[143,1104,752,1000]" pageId="18" pageNumber="42">
Measurements of the
<typeStatus id="54DB96DAFFD42C24EB8E8EEFFE7FFCD0" box="[385,485,752,776]" pageId="18" pageNumber="42" type="holotype">holotype</typeStatus>
based on published photographs (
<bibRefCitation id="EFF15589FFD42C24E96A8EEFFC60FCD0" author="Boeters HD" box="[869,1018,752,776]" pageId="18" pageNumber="42" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
, fig. 10;
<bibRefCitation id="EFF15589FFD42C24EA808F0EFEB9FCF0" author="Boeters HD" box="[143,291,784,808]" pageId="18" pageNumber="42" pagination="77 - 81" refId="ref22950" refString="Boeters HD. 1999. Alzionella navarrensis n sp., Pseudamnicola (Corrosella) hydrobiopsis n. sp. and the type especies of Pseudamnicola Paulucci, 1878 unknown West European Prosobranchia. Basteria 63: 77 - 81." type="journal article" year="1999">Boeters 1999</bibRefCitation>
, fig. 3) (
<figureCitation id="135B34FDFFD42C24EB728F0FFE68FCF0" box="[381,498,784,808]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="18" pageNumber="42">Figure 8A</figureCitation>
) and the available
<typeStatus id="54DB96DAFFD42C24E8CB8F0EFCABFCF1" box="[708,817,785,809]" pageId="18" pageNumber="42" type="paratype">paratypes</typeStatus>
(
<figureCitation id="135B34FDFFD42C24E94F8F0FFC56FCF0" box="[832,972,784,808]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="18" pageNumber="42">Figure 8B,C</figureCitation>
) were compared with measurements of specimens of all the new localities where the species has been found. As explained below, La Armada spring specimens (Orce) were selected for morphometric comparisons with the other species here examined because they were the least eroded and most had a preserved apex. Shell, anatomical, operculum and radular measurements for La Armada (Appendix
<tableCitation id="C6E21DC3FFD42C24E8D88FB0FCCAFC1F" box="[727,848,943,967]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStart-4="Table 5" captionStart-5="Table 6" captionStart-6="Table 7" captionStart-7="Table 8" captionStartId-0="57.[58,113,547,569]" captionStartId-1="58.[58,113,526,548]" captionStartId-2="59.[58,113,630,652]" captionStartId-3="60.[58,113,646,668]" captionStartId-4="61.[58,113,600,622]" captionStartId-5="62.[58,113,483,505]" captionStartId-6="63.[58,113,585,607]" captionStartId-7="64.[58,113,439,461]" captionText-0="Table 1. Shell measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén" captionText-1="Table 2. Operculum measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-2="Table 3. Ctenidium and osphradium measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-3="Table 4. Radula formulae and measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-4="Table 5. Digestive system measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-5="Table 6. Female genitalia measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-6="Table 7. Male genitalia measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." captionText-7="Table 8. Nervous system measurements recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="18" pageNumber="42">Tables 18</tableCitation>
) correspond to males and females collected in May and October.
</paragraph>
<paragraph id="8BDF2878FFD42C24EA80880DFF61FBF1" blockId="18.[143,251,1042,1065]" box="[143,251,1042,1065]" pageId="18" pageNumber="42">
<emphasis id="B914F46AFFD42C24EA80880DFF61FBF1" box="[143,251,1042,1065]" italics="true" pageId="18" pageNumber="42">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD42C24EA808826FC3AFAC8" blockId="18.[143,1104,1081,1296]" pageId="18" pageNumber="42">Shell small with yellowish periostracum; radial sculpture on teleoconch; outer peristome with a slight sinuosity in lateral view; plain central radular tooth weakly serrated and lateral teeth with six lateral cusps at each side of median cusp; bursa copulatrix long cylindrical; seminal receptacle elongated; renal oviduct black pigmented until loop; prostate gland three or four times longer than wide; penis slender tubular with a small black patch of pigment in distal region; nervous system black pigmented with a supraoesophageal connective eight times longer than suboesophageal.</paragraph>
<paragraph id="8BDF2878FFD42C24EA808923FE94FA8B" blockId="18.[143,270,1340,1363]" box="[143,270,1340,1363]" pageId="18" pageNumber="42">
<emphasis id="B914F46AFFD42C24EA808923FE94FA8B" box="[143,270,1340,1363]" italics="true" pageId="18" pageNumber="42">Description</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD42C24EA80897CFBD5F942" blockId="18.[143,1103,1379,1690]" pageId="18" pageNumber="42">
Shell ovate-conic, yellowish periostracum with 4.55 spire whorls, height
<quantity id="4C98859DFFD42C24E9CB897CFBA9FAA3" box="[964,1075,1379,1403]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="18" pageNumber="42" unit="mm" value="2.3" valueMax="2.6" valueMin="2.0">22.6 mm</quantity>
in non-eroded specimens (
<figureCitation id="135B34FDFFD42C24EB94899CFDA4FA43" box="[411,574,1411,1435]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figures 5AC</figureCitation>
,
<figureCitation id="135B34FDFFD42C24E844899CFDF5FA43" box="[587,623,1411,1435]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="18" pageNumber="42">8D</figureCitation>
, Appendix
<tableCitation id="C6E21DC3FFD42C24E8FB899CFCDCFA43" box="[756,838,1411,1435]" captionStart="Table 1" captionStartId="57.[58,113,547,569]" captionText="Table 1. Shell measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén" pageId="18" pageNumber="42">Table 1</tableCitation>
). In some populations, shells are very eroded at their apex or apex is even lost in different measure showing white colour in this zone; protoconch with around 1.25 whorls, 350 µm total length and nucleus width approximately 100 µm (
<figureCitation id="135B34FDFFD42C24E86589FDFD6CFA22" box="[618,758,1506,1530]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figure 5E,F</figureCitation>
); granular microsculpture with granules more strongly patched at the protoconch nucleus (
<figureCitation id="135B34FDFFD42C24E94C8A1DFC24F9C2" box="[835,958,1538,1562]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figure 5G</figureCitation>
); teleoconch with radial sculpture marks on its surface, more marked on body whorl which occupies more than half total shell length; peristome orthocline, oval, complete, with a thin outer lip and a thicker inner lip which is in contact with body whorl hiding the umbilicus; outer peristome simple showing slight sinuosity in lateral view (
<figureCitation id="135B34FDFFD42C24E9C58A9DFBD9F942" box="[970,1091,1666,1690]" captionStart="Figure 5" captionStartId="13.[211,278,1505,1527]" captionTargetBox="[211,1170,169,1472]" captionTargetId="figure-5@13.[211,1170,169,1472]" captionTargetPageId="13" captionText="Figure 5. Shells of Pseudamnicola (C.) falkneri. (A, DG) Shells from La Armada spring, Orce, Granada (MNCN 15.05/49161); (B) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171; (C) shell from La Errá spring, La Dehesa, Albacete (MNCN 15.05/49172). (EG) protoconch and microsculpture." figureDoi="http://doi.org/10.5281/zenodo.5199557" httpUri="https://zenodo.org/record/5199557/files/figure.png" pageId="18" pageNumber="42">Figure 5D</figureCitation>
).
</paragraph>
<paragraph id="8BDF2878FFD52C25EAF18CB8FC72FF07" blockId="19.[211,1172,167,478]" pageId="19" pageNumber="43">
Operculum with around 2.5 spire whorls on internal side; oval muscular attachment mark appears near the nucleus (
<figureCitation id="135B34FDFFD52C25E8778CD8FC9CFF07" box="[632,774,199,223]" captionStart="Figure 6" captionStartId="14.[143,210,1417,1439]" captionTargetBox="[143,1102,168,1384]" captionTargetId="figure-6@14.[143,1102,167,1384]" captionTargetPageId="14" captionText="Figure 6. Operculum and radula of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49161). (A) Internal side of the operculum; (B): external side of the operculum; (C): radula; (D) rows of teeth of the radula; (E) central tooth; (F) outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.5199559" httpUri="https://zenodo.org/record/5199559/files/figure.png" pageId="19" pageNumber="43">Figure 6A,B</figureCitation>
, Appendix
<tableCitation id="C6E21DC3FFD52C25E9888CD8FC41FF07" box="[903,987,199,223]" captionStart="Table 2" captionStartId="58.[58,113,526,548]" captionText="Table 2. Operculum measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BDF2878FFD52C25EAF18CF8FCB8FE06" blockId="19.[211,1172,167,478]" pageId="19" pageNumber="43">
Radula approximately eight times longer than wide (
<figureCitation id="135B34FDFFD52C25E9498CF8FC20FF27" box="[838,954,231,255]" captionStart="Figure 6" captionStartId="14.[143,210,1417,1439]" captionTargetBox="[143,1102,168,1384]" captionTargetId="figure-6@14.[143,1102,167,1384]" captionTargetPageId="14" captionText="Figure 6. Operculum and radula of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49161). (A) Internal side of the operculum; (B): external side of the operculum; (C): radula; (D) rows of teeth of the radula; (E) central tooth; (F) outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.5199559" httpUri="https://zenodo.org/record/5199559/files/figure.png" pageId="19" pageNumber="43">Figure 6C</figureCitation>
, Appendix
<tableCitation id="C6E21DC3FFD52C25EE358CF8FB11FF27" box="[1082,1163,231,255]" captionStart="Table 4" captionStartId="60.[58,113,646,668]" captionText="Table 4. Radula formulae and measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 4</tableCitation>
) of intermediate size (representing 30% of total shell length); contains around 62 rows of teeth; central tooth flattened, without cusps, border only weakly serrate (
<figureCitation id="135B34FDFFD52C25E9F28D38FB10FEE7" box="[1021,1162,295,319]" captionStart="Figure 6" captionStartId="14.[143,210,1417,1439]" captionTargetBox="[143,1102,168,1384]" captionTargetId="figure-6@14.[143,1102,167,1384]" captionTargetPageId="14" captionText="Figure 6. Operculum and radula of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49161). (A) Internal side of the operculum; (B): external side of the operculum; (C): radula; (D) rows of teeth of the radula; (E) central tooth; (F) outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.5199559" httpUri="https://zenodo.org/record/5199559/files/figure.png" pageId="19" pageNumber="43">Figure 6D,E</figureCitation>
) with small basal cusp on each side; lateral teeth contain tongue-shaped central cusps and six laterals (external ones longer than internal); inner marginal teeth have 33 cusps which are smaller towards the tooth base, but the fourth cusp from the tooth base is wider and longer than those surrounding it; outer marginal teeth with 25 cusps occupying over 25% of tooth surface (
<figureCitation id="135B34FDFFD52C25E8728DD9FC8FFE06" box="[637,789,454,478]" captionStart="Figure 6" captionStartId="14.[143,210,1417,1439]" captionTargetBox="[143,1102,168,1384]" captionTargetId="figure-6@14.[143,1102,167,1384]" captionTargetPageId="14" captionText="Figure 6. Operculum and radula of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49161). (A) Internal side of the operculum; (B): external side of the operculum; (C): radula; (D) rows of teeth of the radula; (E) central tooth; (F) outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.5199559" httpUri="https://zenodo.org/record/5199559/files/figure.png" pageId="19" pageNumber="43">Figure 6DF</figureCitation>
).
</paragraph>
<paragraph id="8BDF2878FFD52C25EADC8DE8FCB6FD15" blockId="19.[211,1172,502,1323]" pageId="19" pageNumber="43">
<emphasis id="B914F46AFFD52C25EADC8DE8FD99FDD6" box="[211,515,502,526]" italics="true" pageId="19" pageNumber="43">Pigmentation and anatomy.</emphasis>
Head and tentacles dark brown, but pigment absent from ocular lobes (
<figureCitation id="135B34FDFFD52C25EB628E09FE79FDF6" box="[365,483,534,558]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7E</figureCitation>
); foot intermediate in size with dark brown pigmentation on its dorsal side. Ctenidium in the middle of the pallial cavity; contains 1213 gill filaments taller than long; osphradium opposite middle section of ctenidium (
<figureCitation id="135B34FDFFD52C25EE168E49FB14FDB6" box="[1049,1166,598,622]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7C</figureCitation>
, Appendix
<tableCitation id="C6E21DC3FFD52C25EB418E69FE3CFD56" box="[334,422,630,654]" captionStart="Table 3" captionStartId="59.[58,113,630,652]" captionText="Table 3. Ctenidium and osphradium measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 3</tableCitation>
). Stomach and style sac longer than wide (Appendix
<tableCitation id="C6E21DC3FFD52C25EE228E69FB1FFD56" box="[1069,1157,630,654]" captionStart="Table 5" captionStartId="61.[58,113,600,622]" captionText="Table 5. Digestive system measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 5</tableCitation>
); its anterior region being surrounded by intestine, pigmented in some specimens (
<figureCitation id="135B34FDFFD52C25EAD48EAAFED4FD15" box="[219,334,693,717]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7F</figureCitation>
); rectum slightly S-shaped in pallial cavity.
</paragraph>
<paragraph id="8BDF2878FFD52C25EAF18EC9FC63FC14" blockId="19.[211,1172,502,1323]" pageId="19" pageNumber="43">
Two morphologies of female genitalia observed within the same population: one with both glands in the pallial oviduct approximately equal in size (Appendix
<tableCitation id="C6E21DC3FFD52C25EE358EEAFB11FCD5" box="[1082,1163,757,781]" captionStart="Table 6" captionStartId="62.[58,113,483,505]" captionText="Table 6. Female genitalia measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 6</tableCitation>
) and a long cylindrical bursa folded into a J-shape (
<figureCitation id="135B34FDFFD52C25E9018F0AFC39FCF5" box="[782,931,789,813]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7G,H</figureCitation>
), sometimes its distal end also folds; and the other with a capsule gland longer than albumen gland and cylindrical U-shaped bursa copulatrix (
<figureCitation id="135B34FDFFD52C25E8528F4AFD44FCB5" box="[605,734,853,877]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7I,J</figureCitation>
); in both, the renal oviduct is straight and lacks pigment from its joining point with the bursal duct to oviduct loop, thereafter it becomes blackish and undulates; small elongated seminal receptacle without duct lying on renal oviduct above insertion of the bursa copulatrix duct.
</paragraph>
<paragraph id="8BDF2878FFD52C25EAF18FCBFEFEFB54" blockId="19.[211,1172,502,1323]" pageId="19" pageNumber="43">
Male genitalia with a three or four times longer than wide prostate gland (Appendix
<tableCitation id="C6E21DC3FFD52C25EB578FEBFE2BFBD4" box="[344,433,1012,1036]" captionStart="Table 7" captionStartId="63.[58,113,585,607]" captionText="Table 7. Male genitalia measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 7</tableCitation>
), and efferent duct entering into the ventral middle-posterior section and deferent duct emerging in the anterior section (
<figureCitation id="135B34FDFFD52C25E98E880BFC61FBF4" box="[897,1019,1044,1068]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7D</figureCitation>
); penis intermediate size with base wider than distal region, flattened with a black small patch of pigmentation in distal region (
<figureCitation id="135B34FDFFD52C25E828884BFD06FBB4" box="[551,668,1108,1132]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7E</figureCitation>
); straight penial duct runs along the external side of penis.
</paragraph>
<paragraph id="8BDF2878FFD52C25EAF1888BFC41FAF3" blockId="19.[211,1172,502,1323]" pageId="19" pageNumber="43">
Nervous system black pigmented, with darker pigment on ganglia; cerebral ganglion equal in size approximately joined by
<quantity id="4C98859DFFD52C25E8BC88ACFC8EFB13" box="[691,788,1203,1227]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="43" unit="mm" value="0.11">0.11 mm</quantity>
of commissure; supraoesophageal connective is eight times longer than suboesophageal, which usually consists of a simple strangulation; left pleural ganglion usually larger than right (
<figureCitation id="135B34FDFFD52C25E9F488ECFB15FAD3" box="[1019,1167,1267,1291]" captionStart="Figure 7" captionStartId="15.[211,278,1576,1598]" captionTargetBox="[213,1167,164,1543]" captionTargetId="figure-5@15.[211,1170,163,1543]" captionTargetPageId="15" captionText="Figure 7. Anatomy of Pseudamnicola (C.) falkneri from La Armada spring, Orce, Granada (MNCN 15.05/49162). (A,B) Partial nervous system; (C) ctenidium and osphradium; (D) prostate gland; (E) head of a male and penis; (F) stomach; (G) first morphology of female genitalia; (H) bursa copulatrix and seminal receptacle of first morphology; (I) second morphology of female genitalia; (J) bursa copulatrix and seminal receptacle of second morphology." figureDoi="http://doi.org/10.5281/zenodo.5199561" httpUri="https://zenodo.org/record/5199561/files/figure.png" pageId="19" pageNumber="43">Figure 7A,B</figureCitation>
, Appendix
<tableCitation id="C6E21DC3FFD52C25EB46890CFE01FAF3" box="[329,411,1299,1323]" captionStart="Table 8" captionStartId="64.[58,113,439,461]" captionText="Table 8. Nervous system measurements recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="19" pageNumber="43">Table 8</tableCitation>
). Mean RPG ratio 0.49 (moderately concentrated).
</paragraph>
<paragraph id="8BDF2878FFD52C25EADC8974FEAFFA5B" blockId="19.[211,309,1387,1411]" box="[211,309,1387,1411]" pageId="19" pageNumber="43">
<emphasis id="B914F46AFFD52C25EADC8974FEAFFA5B" box="[211,309,1387,1411]" italics="true" pageId="19" pageNumber="43">Remarks</emphasis>
</paragraph>
<paragraph id="8BDF2878FFD52C22EADC898CFBBEFF27" blockId="19.[211,1172,1427,1706]" lastBlockId="20.[143,1104,167,1467]" lastPageId="20" lastPageNumber="44" pageId="19" pageNumber="43">
<materialsCitation id="3B082225FFD52C25EADC898CFBFBF9D2" collectionCode="BOE, SMF" collectorName="Falkner" county="Orce" location="After" municipality="Galera" pageId="19" pageNumber="43" specimenCode="BOE 222, 223, SMF 219026" specimenCount="1" typeStatus="holotype">
<location id="8EBF7EA3FFD52C25EADC898CFE95FA73" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD52C25EADC898CFE95FA73" box="[211,271,1427,1451]" county="Orce" municipality="Galera" name="After" pageId="19" pageNumber="43">After</location>
examining the available type material, the labels and the original description (
<bibRefCitation id="EFF15589FFD52C25EAD489ADFEF7FA12" author="Boeters HD" box="[219,365,1458,1482]" pageId="19" pageNumber="43" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
and personal communication 2010) one can conclude that
<location id="8EBF7EA3FFD52C25EE0989ACFBC1FA12" LSID="urn:lsid:plazi:treatment:03C9996EFFCA2C22EB4389BDFDAAFA63:8EBF7EA3FFD52C25EE0989ACFBC1FA12" box="[1030,1115,1459,1482]" county="Orce" municipality="Galera" name="Boeters" pageId="19" pageNumber="43">Boeters</location>
considered that: (1) all the material collected by
<collectorName id="26954DAEFFD52C25E8C389CDFCBEFA32" box="[716,804,1490,1514]" pageId="19" pageNumber="43">Falkner</collectorName>
between
<collectingMunicipality id="6BBBB202FFD52C25E98189CDFC41FA32" box="[910,987,1490,1514]" pageId="19" pageNumber="43">Galera</collectingMunicipality>
and
<collectingCounty id="62BE50F4FFD52C25EE1A89CDFBD1FA32" box="[1045,1099,1490,1514]" pageId="19" pageNumber="43">Orce</collectingCounty>
(three samples
<specimenCode id="DBC68003FFD52C25EB3A89EDFE3AF9D2" box="[309,416,1522,1546]" collectionCode="BOE" pageId="19" pageNumber="43">BOE 222</specimenCode>
,
<specimenCode id="DBC68003FFD52C25EBA389EDFE4FF9D2" box="[428,469,1522,1546]" collectionCode="BOE" pageId="19" pageNumber="43">223</specimenCode>
a and 223b) was type material, (2)
<typeStatus id="54DB96DAFFD52C25E96D89EDFC5CF9D2" box="[866,966,1522,1546]" pageId="19" pageNumber="43" type="holotype">holotype</typeStatus>
<specimenCode id="DBC68003FFD52C25E9C189EDFBFBF9D2" box="[974,1121,1522,1546]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="43">SMF 219026</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD52C25EE6689EDFE47F9F2" collectionCode="SMF" pageId="19" pageNumber="43" specimenCode="SMF 219027" specimenCount="1" typeStatus="paratype">
and
<typeStatus id="54DB96DAFFD52C25EADC8A0CFEDAF9F3" box="[211,320,1555,1579]" pageId="19" pageNumber="43" type="paratype">paratypes</typeStatus>
<specimenCode id="DBC68003FFD52C25EB468A0DFE47F9F2" box="[329,477,1554,1578]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="43">SMF 219027</specimenCode>
</materialsCitation>
/
<materialsCitation id="3B082225FFD52C25EBE68A0DFC17F9F2" box="[489,909,1554,1578]" collectionCode="BOE" pageId="19" pageNumber="43" specimenCode="BOE 222, BOE 0222" specimenCount="1" typeStatus="paratype">
1 (ex.
<specimenCode id="DBC68003FFD52C25E83E8A0DFD01F9F2" box="[561,667,1554,1578]" collectionCode="BOE" pageId="19" pageNumber="43">BOE 222</specimenCode>
) plus
<specimenCode id="DBC68003FFD52C25E8EC8A0DFCC3F9F2" box="[739,857,1554,1578]" collectionCode="BOE" pageId="19" pageNumber="43">BOE 0222</specimenCode>
and
</materialsCitation>
<materialsCitation id="3B082225FFD52C25E99A8A0DFB91F9F2" box="[917,1035,1554,1578]" collectionCode="BOE" isTailEnumeration="true" pageId="19" pageNumber="43" specimenCode="BOE 2629" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBC68003FFD52C25E99A8A0DFB91F9F2" box="[917,1035,1554,1578]" collectionCode="BOE" pageId="19" pageNumber="43">BOE 2629</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD52C25EE1B8A0DFEB9F952" collectionCode="SMF" pageId="19" pageNumber="43" specimenCode="SMF 219028" specimenCount="1" typeStatus="paratype">
correspond to the type locality “écoulement oriental”, (3) the other two samples (223a and 223b) are from the second original locality “écoulement occidentale”, (4) specimens
<specimenCode id="DBC68003FFD52C25EE568A4DFEB9F952" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="19" pageNumber="43">SMF 219028</specimenCode>
</materialsCitation>
/
<materialsCitation id="3B082225FFD52C25EB208A6DFC91F952" box="[303,779,1650,1674]" collectionCode="NHMW" pageId="19" pageNumber="43" specimenCode="NHMW 107331" specimenCount="1" typeStatus="paratype">
10 and the
<specimenCount id="9D66E3F1FFD52C25EBA68A6DFDCAF952" box="[425,592,1650,1674]" pageId="19" pageNumber="43" type="generic">four specimens</specimenCount>
<specimenCode id="DBC68003FFD52C25E85A8A6CFC91F952" box="[597,779,1650,1674]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="43" type="Museum">NHMW 107331</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD52C22E91E8A6CFF45FF67" collectionCode="BOE, NHMW" lastPageId="20" lastPageNumber="44" pageId="19" pageNumber="43" specimenCode="BOE 223, NHMW 107332" specimenCount="1" typeStatus="paratype">
are subsamples of the same sample (ex.
<specimenCode id="DBC68003FFD52C25EB0C8A8DFEF6F972" box="[259,364,1682,1706]" collectionCode="BOE" pageId="19" pageNumber="43">BOE 223</specimenCode>
b) (“Westl. Quellenaustritt”), whereas the provenance of the 10
<specimenCode id="DBC68003FFD52C22EE3C8A8CFF45FF67" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" lastPageId="20" lastPageNumber="44" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="43" type="Museum">NHMW 107332</specimenCode>
</materialsCitation>
<materialsCitation id="3B082225FFD22C22EAE68CB7FBBAFF27" collectionCode="BOE" collectorName="W. Balsa" pageId="20" pageNumber="44" specimenCode="BOE 223" specimenCount="1" typeStatus="paratype">
<typeStatus id="54DB96DAFFD22C22EAE68CB7FECCFF18" box="[233,342,168,192]" pageId="20" pageNumber="44" type="paratype">paratypes</typeStatus>
is still doubtful because of the contradictory information in their original labels; that is, the location “Quellenaustritt der
<collectorName id="26954DAEFFD22C22E8F78CD7FCC6FF07" box="[760,860,199,223]" pageId="20" pageNumber="44">W. Balsa</collectorName>
” seems to correspond to the third sample 223a, although the writing on the label reads “ex.
<specimenCode id="DBC68003FFD22C22E9958CF8FB9FFF27" box="[922,1029,231,255]" collectionCode="BOE" pageId="20" pageNumber="44">BOE 223</specimenCode>
b”
</materialsCitation>
.
</paragraph>
<paragraph id="8BDF2878FFD22C22EAB58D18FCBDFE66" blockId="20.[143,1104,167,1467]" pageId="20" pageNumber="44">
<figureCitation id="135B34FDFFD22C22EAB58D18FE80FEC7" box="[186,282,263,287]" captionStart="Figure 8" captionStartId="16.[143,210,1025,1047]" captionTargetBox="[143,1102,169,992]" captionTargetId="figure-6@16.[143,1102,169,992]" captionTargetPageId="16" captionText="Figure 8. Shells of several populations of Pseudamnicola (C.) falkneri. (A) Holotype (SMF 219026); (B,C) paratypes (SMF 219027/1, 219028/10); (D) shell from La Armada spring, Orce, Granada (MNCN 15.05/49161); (E) shell from Dos Caños spring, Castril, Granada (MNCN 15.05/49171); (F) shell from Fuente Palo, Orce, Granada (MNCN 15.05/49163)." figureDoi="http://doi.org/10.5281/zenodo.5199563" httpUri="https://zenodo.org/record/5199563/files/figure.png" pageId="20" pageNumber="44">Figure 8</figureCitation>
shows the apex erosion that can be observed in both the
<typeStatus id="54DB96DAFFD22C22E9A08D17FB86FEF8" box="[943,1052,264,288]" pageId="20" pageNumber="44" type="paratype">paratypes</typeStatus>
and shells of recently collected specimens. Erosion frequently produces loss of first whorls and therefore most length measurements are useless, the remaining variables being too few for a comparative statistical analysis. The population from La Armada is the most similar to the
<typeStatus id="54DB96DAFFD22C22EB3A8D99FE03FE46" box="[309,409,390,414]" pageId="20" pageNumber="44" type="holotype">holotype</typeStatus>
and accordingly it was used for conchological comparisons with the other species in this study (see Statistical analysis).
</paragraph>
<paragraph id="8BDF2878FFD22C22EAB58DD9FD29FCC5" blockId="20.[143,1104,167,1467]" pageId="20" pageNumber="44">
Populations from
<collectingCountry id="F37768E8FFD22C22EB8E8DD9FE7CFE06" box="[385,486,454,478]" name="Grenada" pageId="20" pageNumber="44">Granada</collectingCountry>
exhibit two morphologies of female genitalia within the same population, though genetic studies (unpublished data) indicate no divergence between the two morphotypes (0.00%) that would support two different taxonomic entities (see Discussion). However, all females from Albacete have a large bursa copulatrix and smaller pallial oviduct, the bursa copulatrix being even longer in populations from Albacete than
<collectingCountry id="F37768E8FFD22C22EB768E79FE78FDA6" box="[377,482,614,638]" name="Grenada" pageId="20" pageNumber="44">Granada</collectingCountry>
. These morphological inter-population differences are reflected in molecular sequences (0.5% for
<collectionCode id="ED71B0BDFFD22C22E8748E99FD34FD46" box="[635,686,646,670]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="20" pageNumber="44" type="Herbarium">COI</collectionCode>
) but are relatively low and can only be taken to indicate inter-population variability. Boeters described an unpigmented penis both in the original description (
<bibRefCitation id="EFF15589FFD22C22E8388EDAFD53FD05" author="Boeters HD" box="[567,713,709,733]" pageId="20" pageNumber="44" pagination="63 - 69" refId="ref22773" refString="Boeters HD 1970. Corrosella n. gen. (Prosobranchia: Hydrobiidae). J Conchiliol. 108 (3): 63 - 69." type="journal article" year="1970">Boeters 1970</bibRefCitation>
) and in his revision (
<bibRefCitation id="EFF15589FFD22C22E9A18EDAFBDEFD05" author="Boeters HD" box="[942,1092,709,733]" pageId="20" pageNumber="44" pagination="181 - 261" refId="ref22923" refString="Boeters HD. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal. Archiv Molluskenkd. 118 (4 / 6): 181 - 261." type="journal article" year="1988">Boeters 1988</bibRefCitation>
). However, we observed that the distal region of the penis contains a small black patch of pigmentation in all the populations examined.
</paragraph>
<paragraph id="8BDF2878FFD22C22EAB58F3AFD24FBA4" blockId="20.[143,1104,167,1467]" pageId="20" pageNumber="44">
Molecularly,
<taxonomicName id="4C6053FBFFD22C22EB5D8F39FD9FFCE5" box="[338,517,805,829]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EB5D8F39FD9FFCE5" box="[338,517,805,829]" italics="true" pageId="20" pageNumber="44">P. (C.) falkneri</emphasis>
</taxonomicName>
represents an independent lineage within the subgenus
<taxonomicName id="4C6053FBFFD22C22EAD68F5AFED6FC85" authorityName="Boeters" authorityYear="1970" box="[217,332,837,861]" class="Gastropoda" family="Hydrobiidae" genus="Corrosella" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="genus">
<emphasis id="B914F46AFFD22C22EAD68F5AFED6FC85" box="[217,332,837,861]" italics="true" pageId="20" pageNumber="44">Corrosella</emphasis>
</taxonomicName>
, and its relationship with the other two clades is not well defined (
<figureCitation id="135B34FDFFD22C22EA988F7AFE91FCA5" box="[151,267,869,893]" captionStart="Figure 26" captionStartId="50.[143,210,1490,1512]" captionTargetBox="[143,1102,710,1456]" captionTargetId="figure-158@50.[143,1102,710,1457]" captionTargetPageId="50" captionText="Figure 26. Bayesian 50% majority rule consensus tree inferred employing cytochrome c oxidase subunit I (COI) mitochondrial gene partial sequence. The numbers above branches represent Bayesian posterior probabilities. Drawings of synapomorphies are referred to: (1) Shell shape, (2) Penis shape, (3) Seminal receptacle and pigmentation on renal oviduct, (4) Bursa copulatrix shape, (5) Lateral teeth of the radula, (6) Length of gastric caecum, (7) Length of supraoesophageal connective. () and () symbolize two and three specimens with identical haplotype respectively. Branch lengths calculated as the mean branch length of all trees in the posterior distribution in which this branch is present. Scale bar: number of estimated changes per site." figureDoi="http://doi.org/10.5281/zenodo.5199599" httpUri="https://zenodo.org/record/5199599/files/figure.png" pageId="20" pageNumber="44">Figure 26</figureCitation>
). Morphological similarities with the clade comprising
<taxonomicName id="4C6053FBFFD22C22E9878F79FBD5FCA5" authority="Delicado &amp; Machordom &amp; Ramos, 2011" authorityName="Delicado &amp; Machordom &amp; Ramos" authorityYear="2011" box="[904,1103,869,893]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="marisolae" status="sp. nov." subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22E9878F79FBD5FCA5" box="[904,1103,869,893]" italics="true" pageId="20" pageNumber="44">P. (C.) marisolae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2274911FFD22C22EA808F9AFF7CFC44" box="[143,230,901,924]" pageId="20" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C6053FBFFD22C22EAFE8F99FEECFC45" authorityName="Boeters" authorityYear="1984" box="[241,374,901,925]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="luisi" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EAFE8F99FEECFC45" box="[241,374,901,925]" italics="true" pageId="20" pageNumber="44">P. (C.) luisi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6053FBFFD22C22EB8B8F99FDB3FC45" authority="Delicado &amp; Machordom &amp; Ramos, 2011" authorityName="Delicado &amp; Machordom &amp; Ramos" authorityYear="2011" box="[388,553,901,925]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="iruritai" status="sp. nov." subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EB8B8F99FDB3FC45" box="[388,553,901,925]" italics="true" pageId="20" pageNumber="44">P. (C.) iruritai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2274911FFD22C22E83F8F9AFD1FFC44" box="[560,645,901,924]" pageId="20" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6053FBFFD22C22E8B28F99FC12FC45" authority="Delicado &amp; Machordom &amp; Ramos, 2011" authorityName="Delicado &amp; Machordom &amp; Ramos" authorityYear="2011" box="[701,904,901,925]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="andalusica" status="sp. nov." subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22E8B28F99FC12FC45" box="[701,904,901,925]" italics="true" pageId="20" pageNumber="44">P. (C.) andalusica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2274911FFD22C22E9808F9AFC7EFC44" box="[911,996,901,924]" pageId="20" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
are: slen- der shell with same SL/SW ratio (about 1.80, Appendix
<tableCitation id="C6E21DC3FFD22C22E9078FBBFCC2FC64" box="[776,856,932,956]" captionStart="Table 1" captionStartId="57.[58,113,547,569]" captionText="Table 1. Shell measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén" pageId="20" pageNumber="44">Table 1</tableCitation>
), long penis also with the same PL/Head length ratio (about 1.30, Appendix
<tableCitation id="C6E21DC3FFD22C22E9028FDBFCF8FC04" box="[781,866,964,988]" captionStart="Table 7" captionStartId="63.[58,113,585,607]" captionText="Table 7. Male genitalia measurements (in mm) recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="20" pageNumber="44">Table 7</tableCitation>
) and pattern of pigmentation in renal oviduct, starting at the joining point with the bursa duct. Shared characters with
<taxonomicName id="4C6053FBFFD22C22EB4E881AFE76FBC4" authority="Delicado &amp; Machordom &amp; Ramos, 2011" authorityName="Delicado &amp; Machordom &amp; Ramos" authorityYear="2011" box="[321,492,1028,1052]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="manueli" status="sp. nov." subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EB4E881AFE76FBC4" box="[321,492,1028,1052]" italics="true" pageId="20" pageNumber="44">P. (C.) manueli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2274911FFD22C22EBFC881AFDDDFBC4" box="[499,583,1029,1052]" pageId="20" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C6053FBFFD22C22E872881AFC8DFBC4" authority="Delicado &amp; Machordom &amp; Ramos, 2011" authorityName="Delicado &amp; Machordom &amp; Ramos" authorityYear="2011" box="[637,791,1028,1052]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="bareai" status="sp. nov." subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22E872881AFC8DFBC4" box="[637,791,1028,1052]" italics="true" pageId="20" pageNumber="44">P. (C.) bareai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2274911FFD22C22E911881AFCE8FBC4" box="[798,882,1029,1052]" pageId="20" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
include: small trace of pigment on distal region of penis and shorter seminal receptacle compared with the rest of the studied species. Further, the shape and size of the shell aperture in
<taxonomicName id="4C6053FBFFD22C22E9F1885AFF7EFBA4" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22E9F1885AFF7EFBA4" italics="true" pageId="20" pageNumber="44">P. (C.) falkneri</emphasis>
</taxonomicName>
are similar to that of
<taxonomicName id="4C6053FBFFD22C22EBD7887AFD2DFBA4" authorityName="Boeters" authorityYear="1999" box="[472,695,1124,1148]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="hydrobiopsis" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EBD7887AFD2DFBA4" box="[472,695,1124,1148]" italics="true" pageId="20" pageNumber="44">P. (C.) hydrobiopsis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF2878FFD22C22EAB5889BFE0DFAE3" blockId="20.[143,1104,167,1467]" pageId="20" pageNumber="44">
<taxonomicName id="4C6053FBFFD22C22EAB5889BFE62FB44" box="[186,504,1156,1180]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EAB5889BFE62FB44" box="[186,504,1156,1180]" italics="true" pageId="20" pageNumber="44">Pseudamnicola (C.) falkneri</emphasis>
</taxonomicName>
has numerous autapomorphies because it is the only species of
<emphasis id="B914F46AFFD22C22EB0888BBFEEAFB64" box="[263,368,1188,1212]" italics="true" pageId="20" pageNumber="44">Corosella</emphasis>
that shows: a shell as small as
<quantity id="4C98859DFFD22C22E8D288BCFCCBFB63" box="[733,849,1187,1211]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="20" pageNumber="44" unit="mm" value="2.3" valueMax="2.6" valueMin="2.0">22.6 mm</quantity>
, central radular tooth without cusps, six lateral cusps in radular lateral teeth, as few as 1213 gill filaments,
<specimenCount id="9D66E3F1FFD22C22EA8088FBFE8AFB23" box="[143,272,1251,1275]" pageId="20" pageNumber="44" type="female">two female</specimenCount>
genitalia morphologies in some populations, short seminal receptacle (around
<quantity id="4C98859DFFD22C22EAE0891CFECEFAC3" box="[239,340,1283,1307]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="44" unit="mm" value="0.15">0.15 mm</quantity>
), prostate gland as long as almost
<quantity id="4C98859DFFD22C22E8DE891CFC8AFAC3" box="[721,784,1283,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="44" unit="mm" value="2.0">2 mm</quantity>
and RPG ratio approaching 0.5 (Appendix
<tableCitation id="C6E21DC3FFD22C22EB39893CFE13FAE3" box="[310,393,1315,1339]" captionStart="Table 8" captionStartId="64.[58,113,439,461]" captionText="Table 8. Nervous system measurements recorded from Pseudamnicola species: 1, P. (C.) luisi from La Gitana spring in La Peza, Granada; 2, P. (C.) falkneri from La Armada spring, Orce, Granada; 3, P. (C.) manueli sp. nov. from La Garganta stream in Nava de San Pedro, Jaén; 4, P. (C.) bareai sp. nov. from Spring in Ermita de las Santas, Collados de la Sagra, Granada; 5, P. (C.) marisolae sp. nov. from Pilar del Mono spring in Dúrcal, Granada; 6, P. (C.) iruritai sp. nov. from Don Pedro spring in Loja, Granada; 7, P. (C.) andalusica sp. nov. from La Salud spring in Albanchez de Mágina, Jaén." pageId="20" pageNumber="44">Table 8</tableCitation>
).
</paragraph>
<paragraph id="8BDF2878FFD22C22EAB5895BFDAAFA63" blockId="20.[143,1104,167,1467]" pageId="20" pageNumber="44">
In this study, we extend the distribution area both in
<collectingCountry id="F37768E8FFD22C22E90B895CFCF3FA83" box="[772,873,1347,1371]" name="Grenada" pageId="20" pageNumber="44">Granada</collectingCountry>
and in Albacete and the species is cited for the first time from the Almería province. Despite this expansion,
<taxonomicName id="4C6053FBFFD22C22EA80899BFEA6FA43" box="[143,316,1411,1435]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="44" phylum="Mollusca" rank="species" species="falkneri" subGenus="Corrosella">
<emphasis id="B914F46AFFD22C22EA80899BFEA6FA43" box="[143,316,1411,1435]" italics="true" pageId="20" pageNumber="44">P. (C.) falkneri</emphasis>
</taxonomicName>
was included as “Vulnerable” in the Red Book of the Invertebrates of Andalusia (
<bibRefCitation id="EFF15589FFD22C22EB1E89BCFDBFFA63" author="Barea-Azcon JM &amp; Ballesteros-Duperon E &amp; Moreno D" box="[273,549,1443,1467]" pageId="20" pageNumber="44" refId="ref22735" refString="Barea-Azcon JM, Ballesteros-Duperon E, Moreno D. (ed.). 2008. Libro Rojo de los Invertebrados de Andalucia. Consejeria de Medio Ambiente, Junta de Andalucia, Sevilla, 1430 pp." type="book" year="2008">Barea-Azcón et al. 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>