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<document id="6C0B0CB5CCC78A008958178C0431C727" ID-DOI="10.5281/zenodo.174977" ID-GBIF-Dataset="525ace52-9dae-40a6-8392-8a5f22b4ed97" ID-ISSN="1175-5326" ID-Zenodo-Dep="174977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459793812464" checkinUser="plazi" docAuthor="Capa, María &amp; Hutchings, Pat A." docDate="2006" docId="F84A87E37A56BD15B61CFEF3FB13FDC8" docLanguage="en" docName="zt01375p029.pdf" docOrigin="Zootaxa 1375" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Eupolymnia regnans Chamberlin 1919" docType="treatment" docVersion="15" lastPageNumber="13" masterDocId="0473FF9B7A5DBD19B714FFEFFF82FF86" masterDocTitle="Terebellidae (Polychaeta) from Coiba National Park, Panamanian Pacific, including description of four new species and synonymy of the genus Paraeupolymnia with Lanicola" masterLastPageNumber="29" masterPageNumber="1" pageNumber="12" updateTime="1698219408529" updateUser="plazi">
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<mods:title id="FEB8A43366F330EDE47C0E1B2B079E80">Terebellidae (Polychaeta) from Coiba National Park, Panamanian Pacific, including description of four new species and synonymy of the genus Paraeupolymnia with Lanicola</mods:title>
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<mods:namePart id="ED77415F4D7808F692C301B6712937AF">Capa, María</mods:namePart>
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<mods:namePart id="ACCA5BE149B3CD7830C046B2AFCE52DE">Hutchings, Pat A.</mods:namePart>
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<mods:date id="4C61494CBB5065300607278D4C0B9592">2006</mods:date>
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<mods:number id="B2FB0B9B6BEB005DEA0C8491FA75E52A">1375</mods:number>
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<treatment id="F84A87E37A56BD15B61CFEF3FB13FDC8" ID-DOI="http://doi.org/10.5281/zenodo.5657461" ID-GBIF-Taxon="119350764" ID-Zenodo-Dep="5657461" LSID="urn:lsid:plazi:treatment:F84A87E37A56BD15B61CFEF3FB13FDC8" httpUri="http://treatment.plazi.org/id/F84A87E37A56BD15B61CFEF3FB13FDC8" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="38F9657E7A56BD12B61CFEF3FE7CFEDF" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="705C36F57A56BD12B61CFEF3FD6CFEB3" blockId="11.[264,750,283,345]" box="[264,750,283,310]" pageId="11" pageNumber="12">
<heading id="2B1481997A56BD12B61CFEF3FD6CFEB3" bold="true" box="[264,750,283,310]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="B7E34D767A56BD12B61CFEF3FD6CFEB3" ID-CoL="3CWXL" authority="Chamberlin, 1919" authorityName="Chamberlin" authorityYear="1919" box="[264,750,283,310]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="regnans">
<emphasis id="4297EAE77A56BD12B61CFEF3FD6CFEB3" bold="true" box="[264,750,283,310]" pageId="11" pageNumber="12">
<emphasis id="4297EAE77A56BD12B61CFEF3FD84FEB0" bold="true" box="[264,518,284,310]" italics="true" pageId="11" pageNumber="12">Eupolymnia regnans</emphasis>
Chamberlin, 1919
</emphasis>
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<paragraph id="705C36F57A56BD12B61CFED0FE7CFEDF" blockId="11.[264,750,283,345]" box="[264,510,319,345]" pageId="11" pageNumber="12">
<figureCitation id="E8D82A707A56BD12B61CFED0FEF4FEDF" box="[264,374,319,345]" captionStart="FIGURE 2" captionStartId="7.[264,367,1630,1654]" captionTargetBox="[370,1204,317,1615]" captionTargetId="figure@7.[370,1237,316,1615]" captionTargetPageId="7" captionText="FIGURE 2. Line drawings of lateral views of uncini. Eupolymnia joaoi sp. nov (AM W 29686): A. Anterior segment; B. Mid-body segment; C. posterior segment; E. regnans Chamberlin, 1919 (MCZ 2395): D. Segment 13; E. Segment 40. Lanicola guillermoi sp. nov. (MNCN 16.01 / 10525): F. Segment 10; G. Segment 17; H. Mid-body segment; I. Posterior segment. Lanicola eduardoi sp. nov. (MNCN 16.01 / 10527): J. Segment 11; K, L. Mid-body segment; M. Posterior segment. &quot; Terebella &quot; cf. T. gorgonae Monro, 1933 (MNCN 16.01 / 10528): N. Segments 7; O. Mid-body segment; P. Posterior segment. Thelepus fraggleorum sp. nov. (MNCN 16.01 / 10530): Q. Segment 8; R. Segment 20; S. Segment 50; T. Posterior segment. Scale: 20 µm." httpUri="https://zenodo.org/record/174979/files/figure.png" pageId="11" pageNumber="12">Figures 2</figureCitation>
D, E, 3A–E
</paragraph>
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<paragraph id="705C36F57A56BD12B61CFE60FB93FE23" blockId="11.[264,1106,398,454]" box="[264,1041,398,422]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="50F311DB7A56BD12B61CFE60FB93FE23" box="[264,1041,398,422]" pageId="11" pageNumber="12">
<taxonomicName id="B7E34D767A56BD12B61CFE60FD6BFE23" ID-CoL="3CWXL" authority="Chamberlin,1919a: 433" authorityName="Chamberlin" authorityPageNumber="433" authorityYear="1919" box="[264,745,398,422]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="regnans">
<emphasis id="4297EAE77A56BD12B61CFE60FE66FE20" box="[264,484,399,422]" italics="true" pageId="11" pageNumber="12">Eupolymnia regnans</emphasis>
<treatmentCitation id="F14210E47A56BD12B6FFFE61FD6BFE23" author="Chamberlin" box="[491,745,398,421]" page="433" pageId="11" pageNumber="12" year="1919">
<bibRefCitation id="14724B047A56BD12B6FFFE61FD34FE23" author="Chamberlin" box="[491,694,398,421]" pageId="11" pageNumber="27" refString="Chamberlin, R. V. (1919 a) The Annelida Polychaeta. Memoirs of the Museum of Comparative Zoology, 48, 1 - 514." type="journal article" year="1919" yearSuffix="a">Chamberlin,1919a</bibRefCitation>
: 433
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</taxonomicName>
, 434, plate 43,
<figureCitation id="E8D82A707A56BD12B498FE61FB8FFE23" box="[908,1037,398,421]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="6.[264,367,1619,1643]" captionStartId-1="7.[264,367,1630,1654]" captionStartId-2="12.[264,367,1645,1669]" captionTargetBox-0="[268,1317,490,1590]" captionTargetBox-1="[370,1204,317,1615]" captionTargetBox-2="[269,1318,653,1618]" captionTargetId-0="figure@6.[265,1322,486,1594]" captionTargetId-1="figure@7.[370,1237,316,1615]" captionTargetId-2="figure@12.[264,1323,646,1621]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="12" captionText-0="FIGURE 1. Eupolymnia joaoi sp. nov. (AM W 29704): A. Anterior end, dorsal view, right branchiae removed; B. Lateral view; C. Ventral view; D. Notopodium, segment 9; E. Uncini in situ, segment 13; F. Notopodia, segment 13 (top) and 14; G. Uncini in situ, fourth posterior segment with one row of uncini. Annotations: 1 – 16: number of segment, br 1, br 2: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, P: prostomium, ul: upper lip. Scale: see images." captionText-1="FIGURE 2. Line drawings of lateral views of uncini. Eupolymnia joaoi sp. nov (AM W 29686): A. Anterior segment; B. Mid-body segment; C. posterior segment; E. regnans Chamberlin, 1919 (MCZ 2395): D. Segment 13; E. Segment 40. Lanicola guillermoi sp. nov. (MNCN 16.01 / 10525): F. Segment 10; G. Segment 17; H. Mid-body segment; I. Posterior segment. Lanicola eduardoi sp. nov. (MNCN 16.01 / 10527): J. Segment 11; K, L. Mid-body segment; M. Posterior segment. &quot; Terebella &quot; cf. T. gorgonae Monro, 1933 (MNCN 16.01 / 10528): N. Segments 7; O. Mid-body segment; P. Posterior segment. Thelepus fraggleorum sp. nov. (MNCN 16.01 / 10530): Q. Segment 8; R. Segment 20; S. Segment 50; T. Posterior segment. Scale: 20 µm." captionText-2="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri-0="https://zenodo.org/record/174978/files/figure.png" httpUri-1="https://zenodo.org/record/174979/files/figure.png" httpUri-2="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figures 1–3</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="705C36F57A56BD12B61CFE40FBD0FE43" blockId="11.[264,1106,398,454]" box="[264,1106,430,454]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="50F311DB7A56BD12B61CFE40FBD0FE43" box="[264,1106,430,454]" pageId="11" pageNumber="12">
<taxonomicName id="B7E34D767A56BD12B61CFE40FD67FE43" authority="Monro 1933: 1072" authorityName="Monro" authorityPageNumber="1072" authorityYear="1933" box="[264,741,430,454]" class="Magnoliopsida" family="Asteraceae" genus="Polymnia" kingdom="Animalia" order="Asterales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="regnans">
<emphasis id="4297EAE77A56BD12B61CFE40FE48FE40" box="[264,458,431,454]" italics="true" pageId="11" pageNumber="12">Polymnia regnans</emphasis>
:
<treatmentCitation id="F14210E47A56BD12B6CCFE41FD21FE43" author="Monro" box="[472,675,430,453]" page="1072" pageId="11" pageNumber="12" year="1933">
<bibRefCitation id="14724B047A56BD12B6CCFE41FDE2FE43" author="Monro" box="[472,608,430,453]" pageId="11" pageNumber="29" refString="Monro, C. C. A. (1933) The polychaeta sedentaria collected by Dr. C. Crossland at Colon, in the Panama region, and the Galapagos Islands during the expedition of the S. Y. ' St. George'. Proceedings of the Zoological Society of London, 4, 1039 - 1092." type="journal article" year="1933">Monro 1933</bibRefCitation>
: 1072
</treatmentCitation>
, 1073
</taxonomicName>
, Figure 19a, b;
<treatmentCitation id="F14210E47A56BD12B49FFE41FBCCFE43" author="Fauchald" box="[907,1102,430,453]" page="56" pageId="11" pageNumber="12" year="1977">
<bibRefCitation id="14724B047A56BD12B49FFE41FBACFE43" author="Fauchald" box="[907,1070,430,453]" pageId="11" pageNumber="27" refString="Fauchald, K. (1977) Polychaetes from intertidal areas in Panama, with a review of previous shallow-water records. Smithsonian Contributions to Zoology, 221, 1 - 81." type="journal article" year="1977">Fauchald 1977</bibRefCitation>
:56
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="38F9657E7A56BD12B61CFE18FD50FDBF" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="705C36F57A56BD12B61CFE18FD50FDBF" blockId="11.[264,1323,503,1892]" pageId="11" pageNumber="12">
<emphasis id="4297EAE77A56BD12B61CFE18FE64FD96" box="[264,486,503,528]" italics="true" pageId="11" pageNumber="12">Material examined</emphasis>
.
<typeStatus id="AF5888577A56BD12B6E1FE18FDDAFD97" box="[501,600,503,529]" pageId="11" pageNumber="12" type="syntype">Syntype</typeStatus>
,
<collectionCode id="16F2AE307A56BD12B570FE18FD21FD97" LSID="urn:lsid:biocol.org:col:33791" box="[612,675,503,529]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="11" pageNumber="12">MCZ</collectionCode>
2395,
<collectingCountry id="08F476657A56BD12B5E1FE18FCD0FD97" box="[757,850,503,529]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
Expedition,
<collectionCode id="16F2AE307A56BD12B4FCFE18FBB3FD97" box="[1000,1073,503,529]" httpUri="http://grbio.org/cool/30w3-609t" name="U. S. Fish Commission" pageId="11" pageNumber="12">USFC</collectionCode>
Albatross,
<date id="045D10357A56BD12B3ACFE18FEC7FDBF" pageId="11" pageNumber="12" value="1891-03-13">13 March 1891</date>
, coll. Agassiz, location unknown.
</paragraph>
</subSubSection>
<subSubSection id="38F9657E7A56BD12B62CFDA8FB7EF9CA" pageId="11" pageNumber="12" type="description">
<paragraph id="705C36F57A56BD12B62CFDA8FB7EF9CA" blockId="11.[264,1323,503,1892]" pageId="11" pageNumber="12">
<emphasis id="4297EAE77A56BD12B62CFDA8FE40FDE6" box="[312,450,583,608]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
.
<typeStatus id="AF5888577A56BD12B6C7FDA8FDB1FDE7" box="[467,563,583,609]" pageId="11" pageNumber="12" type="syntype">Syntype</typeStatus>
broken in five pieces, measuring altogether around
<quantity id="B71B9B107A56BD12B3B3FDA8FA88FDE7" box="[1191,1290,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="mm" value="160.0">160 mm</quantity>
in length, with about 157 segments. No pigmentation present after fixation; eyespots not seen. Broad tentacular ridge with several rows of long tentacles. Lips well developed; upper lip large and surrounding most of mouth, lower lip longer, folded, and restricted to ventral side of mouth (
<figureCitation id="E8D82A707A56BD12B535FD08FD08FC87" box="[545,650,743,769]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
B, D). Peristomium forming dorsal ring inserted below prostomium. Three pairs of branchiae on segments 2–4 (now partly detached from body), with first pair largest and decreasing slightly in size from 1st to 3rd pair (
<figureCitation id="E8D82A707A56BD12B37AFCD5FB56FCD2" box="[1134,1236,826,852]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
A, C); branchiae arranged in a longitudinal row (not evident in
<figureCitation id="E8D82A707A56BD12B4B5FC8DFB81FCFA" box="[929,1027,866,892]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
C), with each main stalk, long, broad, about half length of entire branchia, and with numerous short compact distal branches. Three pairs of small, pointed lateral lobes on segments 2–4, all similar in size, arranged obliquely ventro-dorsally (
<figureCitation id="E8D82A707A56BD12B5D5FC35FCA9FC72" box="[705,811,986,1012]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
E). Eighteen glandular ventral pads from segment 2 onwards, divided horizontally into 2; pads becoming gradually narrower posteriorly (
<figureCitation id="E8D82A707A56BD12B688FBC5FD85FBC2" box="[412,519,1066,1092]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
B, D). Rounded nephridial papillae located ventral to notopodia, visible on segments 3–5 only (
<figureCitation id="E8D82A707A56BD12B56FFBBDFD63FBEA" box="[635,737,1106,1132]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
E). Notopodia cylindrical and long, on segments 4–21. Notochaetae smooth narrow-winged capillaries arranged in 2 irregular rows, showing dorso-ventral increase in length, graded in length within fascicle. Neuropodia from segment 5 (
<figureCitation id="E8D82A707A56BD12B6E5FB25FDE2FB62" box="[497,608,1226,1252]" captionStart="FIGURE 3" captionStartId="12.[264,367,1645,1669]" captionTargetBox="[269,1318,653,1618]" captionTargetId="figure@12.[264,1323,646,1621]" captionTargetPageId="12" captionText="FIGURE 3. Eupolymnia regnans Chamberlin, 1919 (MCZ 2395): A. Anterior end, lateral view; B. Ventral view; C. Dorso-lateral view; D. Prostomium and anterior segments, ventral view; E. First segments, ventro-lateral view. Scale: A-C: 5 mm, D, E: 1 mm. Annotations: 1 – 14: number of segment, br 1 – br 3: number of pair of branchiae, ll 1 – ll 3: number of pair of lateral lobes, lol: lower lip, ul: upper lip, white arrows: nephridial papillae." httpUri="https://zenodo.org/record/174980/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
E); uncini arranged in single rows on segments 5–10, subsequently in two rows, arranged face to face, on all remaining chaetigers with notopodia. Uncini decreasing in size posteriorly, with one row of teeth above main fang, small or inconspicuous in some uncini (dental formula MF:1:1–0), and dorsal button situated mid way along between prow and base of main fang pointing towards prow (
<figureCitation id="E8D82A707A56BD12B604FA7DFEF9FA2A" box="[272,379,1426,1452]" captionStart="FIGURE 2" captionStartId="7.[264,367,1630,1654]" captionTargetBox="[370,1204,317,1615]" captionTargetId="figure@7.[370,1237,316,1615]" captionTargetPageId="7" captionText="FIGURE 2. Line drawings of lateral views of uncini. Eupolymnia joaoi sp. nov (AM W 29686): A. Anterior segment; B. Mid-body segment; C. posterior segment; E. regnans Chamberlin, 1919 (MCZ 2395): D. Segment 13; E. Segment 40. Lanicola guillermoi sp. nov. (MNCN 16.01 / 10525): F. Segment 10; G. Segment 17; H. Mid-body segment; I. Posterior segment. Lanicola eduardoi sp. nov. (MNCN 16.01 / 10527): J. Segment 11; K, L. Mid-body segment; M. Posterior segment. &quot; Terebella &quot; cf. T. gorgonae Monro, 1933 (MNCN 16.01 / 10528): N. Segments 7; O. Mid-body segment; P. Posterior segment. Thelepus fraggleorum sp. nov. (MNCN 16.01 / 10530): Q. Segment 8; R. Segment 20; S. Segment 50; T. Posterior segment. Scale: 20 µm." httpUri="https://zenodo.org/record/174979/files/figure.png" pageId="11" pageNumber="12">Figure 2</figureCitation>
D). Neuropodia from segment 22 with short tori and dorsal rounded papilla; avicular uncini arranged in single rows to pygidium, with pointed dorsal button mid way along between prow and base of main fang pointing towards prow; dental formula MF:1, with the secondary tooth clearly separated from main fang (
<figureCitation id="E8D82A707A56BD12B4D3F9E5FBACF9A2" box="[967,1070,1546,1572]" captionStart="FIGURE 2" captionStartId="7.[264,367,1630,1654]" captionTargetBox="[370,1204,317,1615]" captionTargetId="figure@7.[370,1237,316,1615]" captionTargetPageId="7" captionText="FIGURE 2. Line drawings of lateral views of uncini. Eupolymnia joaoi sp. nov (AM W 29686): A. Anterior segment; B. Mid-body segment; C. posterior segment; E. regnans Chamberlin, 1919 (MCZ 2395): D. Segment 13; E. Segment 40. Lanicola guillermoi sp. nov. (MNCN 16.01 / 10525): F. Segment 10; G. Segment 17; H. Mid-body segment; I. Posterior segment. Lanicola eduardoi sp. nov. (MNCN 16.01 / 10527): J. Segment 11; K, L. Mid-body segment; M. Posterior segment. &quot; Terebella &quot; cf. T. gorgonae Monro, 1933 (MNCN 16.01 / 10528): N. Segments 7; O. Mid-body segment; P. Posterior segment. Thelepus fraggleorum sp. nov. (MNCN 16.01 / 10530): Q. Segment 8; R. Segment 20; S. Segment 50; T. Posterior segment. Scale: 20 µm." httpUri="https://zenodo.org/record/174979/files/figure.png" pageId="11" pageNumber="12">Figure 2</figureCitation>
E). Some uncini have a knob on this secondary tooth (
<figureCitation id="E8D82A707A56BD12B56AF9DDFD61F9CA" box="[638,739,1586,1612]" captionStart="FIGURE 2" captionStartId="7.[264,367,1630,1654]" captionTargetBox="[370,1204,317,1615]" captionTargetId="figure@7.[370,1237,316,1615]" captionTargetPageId="7" captionText="FIGURE 2. Line drawings of lateral views of uncini. Eupolymnia joaoi sp. nov (AM W 29686): A. Anterior segment; B. Mid-body segment; C. posterior segment; E. regnans Chamberlin, 1919 (MCZ 2395): D. Segment 13; E. Segment 40. Lanicola guillermoi sp. nov. (MNCN 16.01 / 10525): F. Segment 10; G. Segment 17; H. Mid-body segment; I. Posterior segment. Lanicola eduardoi sp. nov. (MNCN 16.01 / 10527): J. Segment 11; K, L. Mid-body segment; M. Posterior segment. &quot; Terebella &quot; cf. T. gorgonae Monro, 1933 (MNCN 16.01 / 10528): N. Segments 7; O. Mid-body segment; P. Posterior segment. Thelepus fraggleorum sp. nov. (MNCN 16.01 / 10530): Q. Segment 8; R. Segment 20; S. Segment 50; T. Posterior segment. Scale: 20 µm." httpUri="https://zenodo.org/record/174979/files/figure.png" pageId="11" pageNumber="12">Figure 2</figureCitation>
E). Pygidium as a wide rim surrounding anus.
</paragraph>
</subSubSection>
<subSubSection id="38F9657E7A56BD15B62CF9B5FB13FDC8" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
<paragraph id="705C36F57A56BD15B62CF9B5FDC5FE28" blockId="11.[264,1323,503,1892]" lastBlockId="12.[264,1323,284,590]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="4297EAE77A56BD12B62CF9B5FE1CF9F5" box="[312,414,1626,1651]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
. The original description is brief and inadequately illustrated so the only known
<typeStatus id="AF5888577A56BD12B674F96DFE39F91A" box="[352,443,1666,1692]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
is redescribed here. Some slight differences between this and the original description can be noticed. Eyespots mentioned by
<bibRefCitation id="14724B047A56BD12B473F945FBD7F942" author="Chamberlin" box="[871,1109,1706,1732]" pageId="11" pageNumber="27" refString="Chamberlin, R. V. (1919 a) The Annelida Polychaeta. Memoirs of the Museum of Comparative Zoology, 48, 1 - 514." type="journal article" year="1919" yearSuffix="a">Chamberlin (1919a)</bibRefCitation>
are not visible on the
<typeStatus id="AF5888577A56BD12B620F93DFE10F96A" box="[308,402,1746,1772]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
, probably due to the effect of the fixation or long term storage. Lateral lobes were not described in the original description although they are present on the
<typeStatus id="AF5888577A56BD12B3BAF915FA8BF892" box="[1198,1289,1786,1812]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
as small, pointed lobes, similar in size on all three segments and arranged obliquely. Nephridial papillae are only visible on segments 3–5, although
<bibRefCitation id="14724B047A56BD12B321F8A5FAA8F8E2" author="Chamberlin" box="[1077,1322,1866,1892]" pageId="11" pageNumber="27" refString="Chamberlin, R. V. (1919 a) The Annelida Polychaeta. Memoirs of the Museum of Comparative Zoology, 48, 1 - 514." type="journal article" year="1919" yearSuffix="a">Chamberlin (1919a)</bibRefCitation>
described them as present on segments 3–8; this difference is probably due to the poor condition of the
<typeStatus id="AF5888577A51BD15B6D8FEABFDA8FED8" box="[460,554,324,350]" pageId="12" pageNumber="13" type="syntype">syntype</typeStatus>
, especially on the tegument of these segments. The length of the handle of the posterior uncini resembles that described by Chamberlin but is not as long as recorded by
<bibRefCitation id="14724B047A51BD15B68EFE7BFDC1FE28" author="Monro" box="[410,579,404,430]" pageId="12" pageNumber="29" refString="Monro, C. C. A. (1933) The polychaeta sedentaria collected by Dr. C. Crossland at Colon, in the Panama region, and the Galapagos Islands during the expedition of the S. Y. ' St. George'. Proceedings of the Zoological Society of London, 4, 1039 - 1092." type="journal article" year="1933">Monro (1933)</bibRefCitation>
.
</paragraph>
<paragraph id="705C36F57A51BD15B62CFE53FB13FDC8" blockId="12.[264,1323,284,590]" pageId="12" pageNumber="13">
<taxonomicName id="B7E34D767A51BD15B62CFE53FDB1FE53" box="[312,563,444,469]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="12" pageNumber="13" phylum="Annelida" rank="species" species="regnans">
<emphasis id="4297EAE77A51BD15B62CFE53FDB1FE53" box="[312,563,444,469]" italics="true" pageId="12" pageNumber="13">Eupolymnia regnans</emphasis>
</taxonomicName>
is characterised by having 18 ventral pads whereas other
<taxonomicName id="B7E34D767A51BD15B61CFE0BFE17FE7B" box="[264,405,484,509]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="12" pageNumber="13" phylum="Annelida" rank="genus">
<emphasis id="4297EAE77A51BD15B61CFE0BFE17FE7B" box="[264,405,484,509]" italics="true" pageId="12" pageNumber="13">Eupolymnia</emphasis>
</taxonomicName>
species have 14 or 15 ventral pads. In addition, the uncini have only one secondary tooth per row above the main fang with two rows in anterior and one in posterior segments, while all other species have more than one tooth per row.
</paragraph>
</subSubSection>
</treatment>
</document>
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