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<document id="BA586B1A3EDC0586AF41593FFC10EF44" ID-CLB-Dataset="8277" ID-DOI="10.11646/zootaxa.4858.3.1" ID-GBIF-Dataset="f8abea37-7992-482c-8a49-982d08cf9f66" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412321" ID-ZooBank="7BA52F1C-4084-4915-A7D9-8DA99379086B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609688599536" checkinUser="plazi" docAuthor="Uusitalo, Matti, Ueckermann, Edward A. &amp; Theron, Pieter D." docDate="2020" docId="AC6C87BAC473EB7BFF1C62E4138DD4DB" docLanguage="en" docName="zootaxa.4858.3.1.pdf" docOrigin="Zootaxa 4858 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Proteromichaeliini Uusitalo &amp; Ueckermann &amp; Theron 2020, trib. nov." docType="treatment" docVersion="8" lastPageNumber="324" masterDocId="5055FFC2C464EB6CFF8B62731073D77B" masterDocTitle="A review of the family Alycidae (Acari, Acariformes) from South Africa" masterLastPageNumber="340" masterPageNumber="301" pageNumber="324" updateTime="1698851566087" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7FB09E207F0BBAB90F513C895E58D622">A review of the family Alycidae (Acari, Acariformes) from South Africa</mods:title>
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<mods:namePart id="32CDE0FE8FAD0CD8ABE9C902EE2D20A3">Uusitalo, Matti</mods:namePart>
<mods:affiliation id="E54BA2D8F117BB2E0EB53EFE45B6B628">Zoological Museum, Centre for Biodiversity, University of Turku, FIN- 20014 Turku, Finland</mods:affiliation>
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<mods:namePart id="0D0ABFFBE8A2B7BEB1F9B52FE2A199E3">Ueckermann, Edward A.</mods:namePart>
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<mods:affiliation id="BE5E745390238DABF1CAD73F6FB69A4D">Unit for Environmental Sciences and Management, Potchefstroom Campus, North-West University, Private Bag X 6001, Potchefstroom, 2520, South Africa &amp; https: // orcid. org / 0000 - 0003 - 4213 - 4309</mods:affiliation>
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<mods:namePart id="D73858C97451889502FE13F3868BB94B">Theron, Pieter D.</mods:namePart>
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<mods:affiliation id="217C88F5408835FA9E75A6BD22192D6A">Unit for Environmental Sciences and Management, Potchefstroom Campus, North-West University, Private Bag X 6001, Potchefstroom, 2520, South Africa &amp; https: // orcid. org / 0000 - 0003 - 4213 - 4309</mods:affiliation>
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<mods:date id="4A6C528AC5E4236152E7E65A8B489EA4">2020</mods:date>
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<treatment id="AC6C87BAC473EB7BFF1C62E4138DD4DB" ID-DOI="http://doi.org/10.5281/zenodo.4535989" ID-GBIF-Taxon="177319602" ID-Zenodo-Dep="4535989" LSID="urn:lsid:plazi:treatment:AC6C87BAC473EB7BFF1C62E4138DD4DB" httpUri="http://treatment.plazi.org/id/AC6C87BAC473EB7BFF1C62E4138DD4DB" lastPageNumber="324" pageId="23" pageNumber="324">
<subSubSection id="6CDF6527C473EB7BFF1C62E4118AD7CA" box="[151,505,151,177]" pageId="23" pageNumber="324" type="nomenclature">
<paragraph id="247A36ACC473EB7BFF1C62E4118AD7CA" blockId="23.[151,505,151,177]" box="[151,505,151,177]" pageId="23" pageNumber="324">
<heading id="7F3281C0C473EB7BFF1C62E4118AD7CA" bold="true" box="[151,505,151,177]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<emphasis id="16B1EABEC473EB7BFF1C62E4118AD7CA" bold="true" box="[151,505,151,177]" pageId="23" pageNumber="324">
<taxonomicName id="E3C54D2FC473EB7BFF1C62E411F2D7CA" authority="Uusitalo &amp; Ueckermann &amp; Theron, 2020" authorityName="Uusitalo &amp; Ueckermann &amp; Theron" authorityYear="2020" box="[151,385,151,177]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
<taxonomicNameLabel id="0D8257C5C473EB7BFE0362E4118AD7CA" box="[392,505,151,177]" pageId="23" pageNumber="324" rank="tribe">trib. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6CDF6527C473EB7BFF1C62AF125FD788" box="[151,556,220,244]" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="247A36ACC473EB7BFF1C62AF125FD788" blockId="23.[151,556,220,244]" box="[151,556,220,244]" pageId="23" pageNumber="324">
<typeStatus id="FB7E880EC473EB7BFF1C62AF10BAD78F" box="[151,201,220,244]" pageId="23" pageNumber="324">Type</typeStatus>
genus:
<taxonomicName id="E3C54D2FC473EB7BFE9362AF11BAD78F" authority="Uusitalo &amp; Ueckermann &amp; Theron, 2020" authorityName="Uusitalo &amp; Ueckermann &amp; Theron" authorityYear="2020" box="[280,457,220,244]" class="Arachnida" family="Alycidae" genus="Proteromichaelia" higherTaxonomySource="CoL" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="16B1EABEC473EB7BFE9362AF11BAD78F" box="[280,457,220,244]" italics="true" pageId="23" pageNumber="324">Proteromichaelia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0D8257C5C473EB7BFE4462AF125FD788" box="[463,556,220,244]" pageId="23" pageNumber="324" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6CDF6527C473EB7BFF1C635214DFD68B" pageId="23" pageNumber="324" type="diagnosis">
<paragraph id="247A36ACC473EB7BFF1C635214DFD68B" blockId="23.[151,1437,289,928]" pageId="23" pageNumber="324">
<emphasis id="16B1EABEC473EB7BFF1C635211EDD640" bold="true" box="[151,414,289,315]" pageId="23" pageNumber="324">Differential diagnosis.</emphasis>
The sole member of this new tribe has several plesiomorphic character states distinctly different from other members of the subfamily
<taxonomicName id="E3C54D2FC473EB7BFD0A6335135DD61B" authorityName="Womersley" authorityYear="1944" box="[641,814,326,352]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="subFamily" subFamily="Bimichaeliinae">Bimichaeliinae</taxonomicName>
. The new tribe can easily be differentiated from its sister group by having large lamellae (the primary cuticular pattern) organized
<emphasis id="16B1EABEC473EB7BFC3463191386D6FF" bold="true" box="[959,1013,362,388]" pageId="23" pageNumber="324">both</emphasis>
in cells
<emphasis id="16B1EABEC473EB7BFBDA6319140DD6FF" bold="true" box="[1105,1150,362,388]" pageId="23" pageNumber="324">and</emphasis>
in clumps inside the cells (
<figureCitation id="BCFE2A29C473EB7BFF1463FD108ED6D3" box="[159,253,398,424]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 92</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFE8C63FD1114D6D3" box="[263,359,398,424]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134A, B</figureCitation>
), secondary pattern by glabrous ridges (small granules/lamellae absent,
<figureCitation id="BCFE2A29C473EB7BFB0C63FD1490D6D3" box="[1159,1251,398,424]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 92</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFB6563FD155ED6D3" box="[1262,1325,398,424]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134B</figureCitation>
), a pair of eyes present (
<figureCitation id="BCFE2A29C473EB7BFEA563C111C1D6B7" box="[302,434,434,460]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 90, 91</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFE3663C11199D6B7" box="[445,490,434,460]" captionStart="FIGURES 101104" captionStartId="26.[151,264,1888,1912]" captionTargetBox="[160,1426,188,1854]" captionTargetId="figure-29@26.[151,1436,181,1864]" captionTargetPageId="26" captionText="FIGURES 101104. Proteromichaelia sila sp. nov. 101—Deutonymph, venter, 4 genital setae per valve; 102—Protonymph, venter, 1 genital seta per valve; 103—Larva, ventral habitus, holotrichy, genital valves absent; 104—Larva, dorsal habitus, neotrichy. 90104 from RSA, near Potchefstroom." figureDoi="http://doi.org/10.5281/zenodo.4412367" httpUri="https://zenodo.org/record/4412367/files/figure.png" pageId="23" pageNumber="324">104</figureCitation>
), a pair of cheliceral setae present (
<figureCitation id="BCFE2A29C473EB7BFCE663C113CED6B7" box="[877,957,434,460]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Fig. 94</figureCitation>
) and divided femora IV (
<figureCitation id="BCFE2A29C473EB7BFB5863C11540D6B7" box="[1235,1331,434,460]" captionStart="FIGURES 101104" captionStartId="26.[151,264,1888,1912]" captionTargetBox="[160,1426,188,1854]" captionTargetId="figure-29@26.[151,1436,181,1864]" captionTargetPageId="26" captionText="FIGURES 101104. Proteromichaelia sila sp. nov. 101—Deutonymph, venter, 4 genital setae per valve; 102—Protonymph, venter, 1 genital seta per valve; 103—Larva, ventral habitus, holotrichy, genital valves absent; 104—Larva, dorsal habitus, neotrichy. 90104 from RSA, near Potchefstroom." figureDoi="http://doi.org/10.5281/zenodo.4412367" httpUri="https://zenodo.org/record/4412367/files/figure.png" pageId="23" pageNumber="324">Fig. 101</figureCitation>
;
<bibRefCitation id="40544B5DC473EB7BFAB663C110A0D68B" author="Uusitalo, M." pageId="23" pageNumber="324" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: Appendix 3). In other respects, the species is the same as the rest of the bimichaeliines.
</paragraph>
</subSubSection>
<subSubSection id="6CDF6527C473EB7BFF4C638A142DD44F" pageId="23" pageNumber="324" type="description">
<paragraph id="247A36ACC473EB7BFF4C638A142DD44F" blockId="23.[151,1437,289,928]" pageId="23" pageNumber="324">
<emphasis id="16B1EABEC473EB7BFF4C638A112AD568" bold="true" box="[199,345,505,531]" pageId="23" pageNumber="324">Description.</emphasis>
Hysterosoma soft, segmented (
<figureCitation id="BCFE2A29C473EB7BFD4B63891365D56F" box="[704,790,506,532]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Fig. 91</figureCitation>
); ornamented idiosomal cuticle (
<figureCitation id="BCFE2A29C473EB7BFB076389149FD56F" box="[1164,1260,506,532]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 92</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFB726389152ED56F" box="[1273,1373,506,532]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134A, B</figureCitation>
): primary pattern formed by large lamellae of polygons and separate clumps, secondary pattern by fine ridges (
<figureCitation id="BCFE2A29C473EB7BFAEF606D10C5D527" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 92</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFF4A60311151D527" box="[193,290,578,604]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134A, B</figureCitation>
); adults neotrichous, dorsal and ventral setae ciliated, genital, anal and pedal setae elongated, plumose (
<figureCitation id="BCFE2A29C473EB7BFF1460151109D5FB" box="[159,378,614,640]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 92, 93, 97, 98</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFE0D601511B5D5FB" box="[390,454,614,640]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134B</figureCitation>
); prodorsum (
<figureCitation id="BCFE2A29C473EB7BFDE860151299D5FB" box="[611,746,614,640]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 90, 91</figureCitation>
,
<figureCitation id="BCFE2A29C473EB7BFD7D60151349D5FB" box="[758,826,614,640]" captionStart="FIGURE 134" captionStartId="37.[151,250,1716,1740]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-18@37.[151,1436,181,1693]" captionTargetPageId="37" captionText="FIGURE 134. Primary pattern of cuticle in the subfamilies Alycinae and Bimichaeliinae.A, B—primary pattern by clumps and rosettes of large lamellae, secondary pattern absent in Proteromichaelia sp. (Bimichaeliinae: Proteromichaeliini), C, D—striate integument with small lamellae in Amphialycus sp., E, F—striate integument with small lamellae in Petralycus sp. (Alycinae: Petralycini)." figureDoi="http://doi.org/10.5281/zenodo.4412387" httpUri="https://zenodo.org/record/4412387/files/figure.png" pageId="23" pageNumber="324">134A</figureCitation>
): naso button-like, a pair of lateral eyes present; posterior sensilla
<emphasis id="16B1EABEC473EB7BFEAA60F81133D5DF" box="[289,320,651,676]" italics="true" pageId="23" pageNumber="324">sci</emphasis>
globular, barbed, anterior pair of sensilla
<emphasis id="16B1EABEC473EB7BFC9860F8135FD5DF" box="[787,812,651,676]" italics="true" pageId="23" pageNumber="324">ve</emphasis>
filamentous; chelate chelicera elongated into stylet-like elements, a pair of cheliceral setae present (
<figureCitation id="BCFE2A29C473EB7BFD0960DD12A4D5B3" box="[642,727,686,712]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Fig. 94</figureCitation>
); subcapitulum elongated, rutella missing, adoral setae missing (
<figureCitation id="BCFE2A29C473EB7BFF2B60A11080D596" box="[160,243,722,749]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Fig. 95</figureCitation>
); palpi with an apical solenidion and two eupathids (
<figureCitation id="BCFE2A29C473EB7BFCB560A113E1D596" box="[830,914,722,749]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Fig. 96</figureCitation>
); supracoxal setae
<emphasis id="16B1EABEC473EB7BFBEE60A0140CD597" box="[1125,1151,723,748]" italics="true" pageId="23" pageNumber="324">ep</emphasis>
present, supracoxal setae
<emphasis id="16B1EABEC473EB7BFF1C608410DED46B" box="[151,173,759,784]" italics="true" pageId="23" pageNumber="324">eI</emphasis>
present, femora IV divided, eupathids apically and ventrally on tarsi I and II, solenidia baculiform and ceratiform, symmetrically paired claws with fine barbs, setulate empodia clawless (
<figureCitation id="BCFE2A29C473EB7BFC3161691420D44F" box="[954,1107,794,820]" captionStart="FIGURES 90100" captionStartId="25.[151,264,1800,1824]" captionTargetBox="[197,1386,188,1767]" captionTargetId="figure-18@25.[189,1397,181,1776]" captionTargetPageId="25" captionText="FIGURES 90100. Proteromichaelia sila sp. nov. 90—Prodorsum, naso button-like, distance between counterparts of setae vi larger than between sensilla ve, setae in with some additional setae; 91—Dorsal habitus, neotrichy; 92—Dorsal integument; 93—Ventral habitus, 910 genital setae per valve, 46 anal setae per valve (cf. Fig. 97); 94—Chelicera, ciliated cheliceral seta; 95—Subcapitulum; 96—Palp tarsus; 97—Genital valve of female, 9 genital setae, 2 pairs of eugenital setae; 98—Tarsus I (2B = 2 baculiform solenidia), famulus I (arrow), Tibia I (1B2C = 1 baculiform solenidion and 2 ceratiform solenidia); 99—Tarsus II (2B), Tibia II (1B2C), Genu II (1B2C); 100—2 slightly bent claws and a setulated empodium." figureDoi="http://doi.org/10.5281/zenodo.4412363" httpUri="https://zenodo.org/record/4412363/files/figure.png" pageId="23" pageNumber="324">Figs. 98100</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6CDF6527C473EB7BFF4C614D138DD4DB" pageId="23" pageNumber="324" type="discussion">
<paragraph id="247A36ACC473EB7BFF4C614D138DD4DB" blockId="23.[151,1437,289,928]" pageId="23" pageNumber="324">
<emphasis id="16B1EABEC473EB7BFF4C614D1143D423" bold="true" box="[199,304,830,856]" pageId="23" pageNumber="324">Remark.</emphasis>
This monotypic sister group of
<taxonomicName id="E3C54D2FC473EB7BFD13614D1343D423" authorityName="Womersley" authorityYear="1944" box="[664,816,830,856]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
was obtained from the analysis of the family
<taxonomicName id="E3C54D2FC473EB7BFABB614D15EAD423" authorityName="G.Canestrini &amp; Fanzago" authorityYear="1877" box="[1328,1433,830,856]" class="Arachnida" family="Alycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinedida" pageId="23" pageNumber="324" phylum="Arthropoda" rank="family">Alycidae</taxonomicName>
. Some drawings of the undescribed species
<emphasis id="16B1EABEC473EB7BFD07611112D5D407" box="[652,678,866,892]" italics="true" pageId="23" pageNumber="324">sil</emphasis>
, representing the new tribe, were included already in the analysis paper to illustrate the character state differences (
<bibRefCitation id="40544B5DC473EB7BFD3561F51312D4DB" author="Uusitalo, M." box="[702,865,902,928]" pageId="23" pageNumber="324" refId="ref18765" refString="Uusitalo, M. (2010) Revision of the family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, Helsinki University Print, University of Helsinki, Helsinki, 143 pp. Available from: http: // ethesis. helsinki. fi (accessed 20 January 2020)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: Appendix 3).
</paragraph>
</subSubSection>
</treatment>
</document>