133 lines
16 KiB
XML
133 lines
16 KiB
XML
<document id="5F3632DB570593D3BEB0314D1AD24D6C" ID-CLB-Dataset="2464" ID-DOI="10.11646/zootaxa.5092.3.1" ID-GBIF-Dataset="2d69d5b2-eeeb-4f2d-b2d0-cd86c462a613" ID-ISSN="1175-5326" ID-Zenodo-Dep="5881128" ID-ZooBank="91B1F45E-F3DF-4FF4-873A-DD3442ABD12A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642665425317" checkinUser="plazi" docAuthor="Marshall, Stephen A." docDate="2022" docId="8A1D87FBFFAAFFF2FF4BFB744007FE7E" docLanguage="en" docName="zootaxa.5092.3.1.pdf" docOrigin="Zootaxa 5092 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nigripeza spangleri Marshall 2022, new species" docType="treatment" docVersion="6" lastPageNumber="271" masterDocId="7624FF83FFB9FFE6FFDCFFF5446CFFAE" masterDocTitle="Amapeza and Nigripeza, new genera of Neotropical micropezid flies (Diptera, Micropezidae, Taeniapterinae)" masterLastPageNumber="272" masterPageNumber="251" pageNumber="270" updateTime="1699302907151" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="608CA4C43A158C2E2B927B9452FA8B2E">Amapeza and Nigripeza, new genera of Neotropical micropezid flies (Diptera, Micropezidae, Taeniapterinae)</mods:title>
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<mods:name id="59A689B7FC22E63CE45F75806D958BCF" type="personal">
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<mods:roleTerm id="B9505AF9B12AFBCAB8689754E188CCB2">Author</mods:roleTerm>
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<mods:namePart id="F8E8310C3F492BE33924FE7F2F14C063">Marshall, Stephen A.</mods:namePart>
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<mods:typeOfResource id="D4946A516A2974DDBA5834BF601B7E06">text</mods:typeOfResource>
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<mods:title id="63CB8DFFF839CF5079BE151907B71D34">Zootaxa</mods:title>
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<mods:part id="57B215561D265C2D17D3D9717D9DFA01">
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<mods:date id="A64852BA46B1D335ED8FD4AFB5CF08A9">2022</mods:date>
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<mods:number id="F733AD4C8116DF640FABF0A9E3D071DB">2022-01-20</mods:number>
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<mods:number id="F8066B0BB24F1087E88EE6982C0DDDD1">5092</mods:number>
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<mods:number id="3D4AC95F982E8D023D41C00602B6D67D">3</mods:number>
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<mods:start id="899FC9B31590847E7AB73EE6A33B977F">251</mods:start>
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<mods:identifier id="9B13EEA56309665E4B88C7079043CCD9" type="DOI">10.11646/zootaxa.5092.3.1</mods:identifier>
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<mods:identifier id="04136B638EC1DD9AC2D94D4F4CF92510" type="ISSN">1175-5326</mods:identifier>
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<treatment id="8A1D87FBFFAAFFF2FF4BFB744007FE7E" ID-DOI="http://doi.org/10.5281/zenodo.5881176" ID-GBIF-Taxon="192501054" ID-Zenodo-Dep="5881176" LSID="urn:lsid:plazi:treatment:8A1D87FBFFAAFFF2FF4BFB744007FE7E" httpUri="http://treatment.plazi.org/id/8A1D87FBFFAAFFF2FF4BFB744007FE7E" lastPageId="20" lastPageNumber="271" pageId="19" pageNumber="270">
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<subSubSection id="4AAE6566FFAAFFF5FF4BFB744677FB35" box="[151,539,1153,1180]" pageId="19" pageNumber="270" type="nomenclature">
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<paragraph id="020B36EDFFAAFFF5FF4BFB744677FB35" blockId="19.[151,539,1153,1215]" box="[151,539,1153,1180]" pageId="19" pageNumber="270">
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<heading id="59438181FFAAFFF5FF4BFB744677FB35" bold="true" box="[151,539,1153,1180]" fontSize="11" level="1" pageId="19" pageNumber="270" reason="1">
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<emphasis id="30C0EAFFFFAAFFF5FF4BFB744677FB35" bold="true" box="[151,539,1153,1180]" pageId="19" pageNumber="270">
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<taxonomicName id="C5B44D6EFFAAFFF5FF4BFB7445E8FB32" ID-CoL="9HZ5W" authority="Marshall, 2022" authorityName="Marshall" authorityYear="2022" box="[151,388,1153,1180]" class="Insecta" family="Micropezidae" genus="Nigripeza" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="270" phylum="Arthropoda" rank="species" species="spangleri" status="sp. nov.">
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<emphasis id="30C0EAFFFFAAFFF5FF4BFB7445E8FB32" bold="true" box="[151,388,1153,1180]" italics="true" pageId="19" pageNumber="270">Nigripeza spangleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2BF35784FFAAFFF5FE50FB744677FB35" box="[396,539,1153,1179]" pageId="19" pageNumber="270" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAE6566FFAAFFF5FF4BFB504049F858" pageId="19" pageNumber="270" type="description">
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<paragraph id="020B36EDFFAAFFF5FF4BFB50450DFB11" blockId="19.[151,539,1153,1215]" box="[151,353,1189,1215]" pageId="19" pageNumber="270">
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<figureCitation id="9A8F2A68FFAAFFF5FF4BFB50450DFB11" box="[151,353,1189,1215]" captionStart="FIGURES 10" captionStartId="20.[151,264,1927,1951]" captionTargetBox="[151,1436,489,1904]" captionTargetId="figure-151@20.[151,1436,488,1904]" captionTargetPageId="20" captionText="FIGURES 10A–10D. Nigripeza spangleri new species, Ecuador. A, spermatheca and associated structures; B, male sternite 5 and genital fork; C, male terminalia, lateral; D, female." figureDoi="http://doi.org/10.5281/zenodo.5881160" httpUri="https://zenodo.org/record/5881160/files/figure.png" pageId="19" pageNumber="270">Figures 10A–10D</figureCitation>
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</paragraph>
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<paragraph id="020B36EDFFAAFFF5FF4BFB1D459DFA50" blockId="19.[151,1437,1256,2039]" pageId="19" pageNumber="270">
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<emphasis id="30C0EAFFFFAAFFF5FF4BFB1D4547FAAC" bold="true" box="[151,299,1256,1282]" pageId="19" pageNumber="270">Description:</emphasis>
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Colour: Body mostly black, head dark brown dorsally, frontal vitta orange anteriorly but phasing to brown posteriorly. Orbital plate pruinose orange, epicephalon dark shiny brown, ocellar plate small and black. Clypeus dark brown. Fore femur brown basally, yellow distally. Fore tibia dark brown, fore tarsomeres 1–2 white, 3–5 black. Other legs yellowish brown; hind femur with a strong and elongate dark ring in basal third; distomedian and distal rings very faint. Vertical row of katepisternal bristles black. Wing with complete but weak broad discal band. Abdominal pleuron (discoloured on available specimens) apparently dirty white except for the male pleural sac, which is large, brownish with dense black microsetulosity. Female oviscape pruinose basally but otherwise shining dark brown, apex pale.
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</paragraph>
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<paragraph id="020B36EDFFAAFFF5FF1BF9FD4089F9C4" blockId="19.[151,1437,1256,2039]" pageId="19" pageNumber="270">Head: Frontal vitta almost parallel sided behind ocelli, expanded and slightly elevated in front of ocelli, hardly tapered anteriorly. Orbital plate dulled with very fine striae; epicephalon shiny and bare; paracephalon mostly bare, microsetulose only between postocellar and inner vertical bristles. Clypeus entirely microsetulose.</paragraph>
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<paragraph id="020B36EDFFAAFFF5FF1BF981458EF91C" blockId="19.[151,1437,1256,2039]" pageId="19" pageNumber="270">Thorax: Postpronotal lobe distinctly elevated posteriorly, sparsely microsetulose. One dorsocentral bristle. Suprahumeral bristles absent.</paragraph>
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<paragraph id="020B36EDFFAAFFF5FF1BF949455BF8C8" blockId="19.[151,1437,1256,2039]" pageId="19" pageNumber="270">Female abdomen: Major (paired) spermathecal duct elongate and almost parallel-sided, gradually expanded to a bulb-like swollen apex from which short, distally expanded transversely striate, convoluted stems lead to 2 funnel-shaped spermathecae. Spermathecae with distinct outpocketings at base and broad distal invaginations. Minor (single) duct shorter and slightly narrower but similar in shape, single spermatheca large, similar to paired spermathecae.</paragraph>
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<paragraph id="020B36EDFFAAFFF5FF1BF8854049F858" blockId="19.[151,1437,1256,2039]" pageId="19" pageNumber="270">Male abdomen: Genital fork with very short, broad, parallel-sided and straight arms; 5 short stout bristles on mesal surface, all in apical third, and 3 long outer preapical or apical bristles as long as or longer than length of arm. Distiphallus short and broad, ending in a broad but tapered and distally pointed phallic bulb. Ejaculatory apodeme not clearly visible in only available male, but apparently smaller than epandrium.</paragraph>
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</subSubSection>
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<subSubSection id="4AAE6566FFADFFF2FF1BFF63402DFF7A" pageId="20" pageNumber="271" type="materials_examined">
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<paragraph id="020B36EDFFADFFF2FF1BFF63402DFF7A" blockId="20.[151,1437,150,465]" pageId="20" pageNumber="271">
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<emphasis id="30C0EAFFFFADFFF2FF1BFF634514FF1E" bold="true" box="[199,376,150,176]" pageId="20" pageNumber="271">Type material:</emphasis>
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<materialsCitation id="B2DC3CB0FFADFFF2FE5CFF63402DFF7A" ID-GBIF-Occurrence="3450193331" collectingDate="1976-01-31" collectionCode="USNM" country="Ecuador" county="Corps Smithsonian Institution Aquatic Insect" location="Spangler" municipality="Peace Corps Smithsonian Institution Aquatic" pageId="20" pageNumber="271" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="DD0F884FFFADFFF2FE5CFF634584FF1E" box="[384,488,150,176]" pageId="20" pageNumber="271" type="holotype">Holotype</typeStatus>
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(
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<specimenCount id="14B2FD64FFADFFF2FE20FF634663FF1E" box="[508,527,150,176]" pageId="20" pageNumber="271" type="female">♀</specimenCount>
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,
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<collectionCode id="64A5AE28FFADFFF2FDC1FF63461CFF1E" box="[541,624,150,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="271" type="Museum">USNM</collectionCode>
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) and
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<typeStatus id="DD0F884FFFADFFF2FD6CFF624777FF1F" box="[688,795,151,177]" pageId="20" pageNumber="271" type="paratype">paratypes</typeStatus>
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(
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<specimenCount id="14B2FD64FFADFFF2FCF2FF634739FF1E" box="[814,853,150,176]" pageId="20" pageNumber="271" type="male">1♂</specimenCount>
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,
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<specimenCount id="14B2FD64FFADFFF2FC87FF634716FF1E" box="[859,890,150,176]" pageId="20" pageNumber="271" type="female">1♀</specimenCount>
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,
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<collectionCode id="64A5AE28FFADFFF2FC54FF6347B3FF1E" box="[904,991,150,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="271" type="Museum">USNM</collectionCode>
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):
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<collectingCountry id="7AA3767DFFADFFF2FC2EFF634011FF1E" box="[1010,1149,150,176]" name="Ecuador" pageId="20" pageNumber="271">ECUADOR</collectingCountry>
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. Past. Puyo,
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<date id="760A102DFFADFFF2FAD2FF6341F4FF1E" box="[1294,1432,150,176]" pageId="20" pageNumber="271" value="1976-01-31">
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<collectingDate id="664EE9C5FFADFFF2FAD2FF6341F4FF1E" box="[1294,1432,150,176]" pageId="20" pageNumber="271" value="1976-01-31">31.Jan.1976</collectingDate>
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</date>
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,
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<location id="076B6036FFADFFF2FF4BFF4F4497FF7A" LSID="urn:lsid:plazi:treatment:8A1D87FBFFAAFFF2FF4BFB744007FE7E:076B6036FFADFFF2FF4BFF4F4497FF7A" box="[151,251,186,212]" country="Ecuador" county="Corps Smithsonian Institution Aquatic Insect" municipality="Peace Corps Smithsonian Institution Aquatic" name="Spangler" pageId="20" pageNumber="271">Spangler</location>
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<emphasis id="30C0EAFFFFADFFF2FEDEFF4E455FFF7A" box="[258,307,186,212]" italics="true" pageId="20" pageNumber="271">et al</emphasis>
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,
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<location id="076B6036FFADFFF2FE9DFF4F4751FF7A" LSID="urn:lsid:plazi:treatment:8A1D87FBFFAAFFF2FF4BFB744007FE7E:076B6036FFADFFF2FE9DFF4F4751FF7A" box="[321,829,186,213]" country="Ecuador" county="Corps Smithsonian Institution Aquatic Insect" municipality="Peace Corps Smithsonian Institution Aquatic" name="Ecuador Peace Corps Smithsonian Institution" pageId="20" pageNumber="271">
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Ecuador
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<collectingMunicipality id="E26FAC97FFADFFF2FE79FF4E4751FF7A" box="[421,829,186,213]" pageId="20" pageNumber="271">
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Peace
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<collectingCounty id="EB6A4E61FFADFFF2FE31FF4F4751FF7A" box="[493,829,186,212]" pageId="20" pageNumber="271">Corps Smithsonian Institution</collectingCounty>
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</collectingMunicipality>
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</location>
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Aquatic Insect Survey.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAE6566FFADFFF2FF1BFF2B4502FEB2" pageId="20" pageNumber="271" type="etymology">
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<paragraph id="020B36EDFFADFFF2FF1BFF2B4502FEB2" blockId="20.[151,1437,150,465]" pageId="20" pageNumber="271">
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<emphasis id="30C0EAFFFFADFFF2FF1BFF2B453EFF56" bold="true" box="[199,338,222,248]" pageId="20" pageNumber="271">Etymology:</emphasis>
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<taxonomicName id="C5B44D6EFFADFFF2FE84FF2A465BFF56" authorityName="Marshall" authorityYear="2022" box="[344,567,222,248]" class="Insecta" family="Micropezidae" genus="Nigripeza" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="spangleri">
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<emphasis id="30C0EAFFFFADFFF2FE84FF2A465BFF56" box="[344,567,222,248]" italics="true" pageId="20" pageNumber="271">Nigripeza spangleri</emphasis>
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</taxonomicName>
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is named after coleopterist Paul Spangler, the collector of this and many other new insect species.
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</paragraph>
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</subSubSection>
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<subSubSection id="4AAE6566FFADFFF2FF1BFED34007FE7E" pageId="20" pageNumber="271" type="discussion">
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<paragraph id="020B36EDFFADFFF2FF1BFED34007FE7E" blockId="20.[151,1437,150,465]" pageId="20" pageNumber="271">
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<emphasis id="30C0EAFFFFADFFF2FF1BFED3453EFEEE" bold="true" box="[199,338,294,320]" pageId="20" pageNumber="271">Comments:</emphasis>
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<taxonomicName id="C5B44D6EFFADFFF2FE85FED24655FEEE" authorityName="Marshall" authorityYear="2022" box="[345,569,294,320]" class="Insecta" family="Micropezidae" genus="Nigripeza" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="spangleri">
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<emphasis id="30C0EAFFFFADFFF2FE85FED24655FEEE" box="[345,569,294,320]" italics="true" pageId="20" pageNumber="271">Nigripeza spangleri</emphasis>
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</taxonomicName>
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is very similar to the closely related
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<taxonomicName id="C5B44D6EFFADFFF2FC04FED2403FFEEE" authorityName="Marshall" authorityYear="2022" baseAuthorityName="Marshall" baseAuthorityYear="2022" box="[984,1107,294,320]" class="Insecta" family="Micropezidae" genus="Nigripeza" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="obscura">
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<emphasis id="30C0EAFFFFADFFF2FC04FED2403FFEEE" box="[984,1107,294,320]" italics="true" pageId="20" pageNumber="271">N. obscura</emphasis>
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</taxonomicName>
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, the only dark congener with a plumose arista. Both species have a swollen cervical sclerite and a single dorsocentral bristle. Male and female terminalia of these species reflect the close relationship but differ clearly in details such as the shape, dimensions and chaetotaxy of the genital fork and the shape and dimensions of the spermathecae and associated ducts.
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<taxonomicName id="C5B44D6EFFADFFF2FAF2FE66456DFE7E" authorityName="Marshall" authorityYear="2022" class="Insecta" family="Micropezidae" genus="Nigripeza" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="spangleri">
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<emphasis id="30C0EAFFFFADFFF2FAF2FE66456DFE7E" italics="true" pageId="20" pageNumber="271">Nigripeza spangleri</emphasis>
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</taxonomicName>
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is distinctive for its very short genital fork and very large single spermatheca.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |