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<document id="139CCB003A760D75246F96ADE4CFD621" ID="10.11646/zootaxa.4269.2.3SLASH11107" ID-CLB-Dataset="32999" ID-DOI="10.11646/zootaxa.4269.2.3" ID-GBIF-Dataset="38f686ec-d435-40e6-8d43-58d7e7b8ae70" ID-ISSN="1175-5326" ID-Zenodo-Dep="582011" ID-ZooBank="01E8B213-8F9D-45E4-86CD-893716D3A295" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495439440136" checkinUser="plazi" docAuthor="Gharali, Babak &amp; Evenhuis, Neal" docDate="2017" docId="834EA522FFE4FFDFDD81F94C2846F81F" docLanguage="en" docName="zootaxa.4269.2.3SLASH11107.pdf" docOrigin="Zootaxa 4269 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtosia bispermatheca Gharali &amp; Evenhuis, sp. nov." docType="treatment" docUuid="7C4A47F5-CE81-4058-AC3B-A13771E3C3C0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="238" masterDocId="7F77DD5AFFE1FFD7DD16FF812920FFD7" masterDocTitle="Review of the genus Cyrtosia Perris (Diptera: Mythicomyiidae: Platypyginae) in Iran, with description of three new species" masterLastPageNumber="244" masterPageNumber="230" pageNumber="235" updateTime="1698439808658" updateUser="ExternalLinkService">
<mods:mods id="9F407A44F3ADB562AFA66555A750CF7A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9A9740A1A9B9684E2479D6390FB9B048">
<mods:title id="45C71CE46A4C58B310A1E2FFB9E67456">Review of the genus Cyrtosia Perris (Diptera: Mythicomyiidae: Platypyginae) in Iran, with description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="6E78890B0E8B35D8062FDE76466C719C" type="personal">
<mods:role id="E44AE9D7679C7C5AA88A4D385A3C8138">
<mods:roleTerm id="0B035D8B8DE1B9DB8634EDBC38EDE687">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="289C312A2953E531CE9E21BC764C916D">Gharali, Babak</mods:namePart>
</mods:name>
<mods:name id="E1A422B617DBFDB0CBAA7E246E073DF9" type="personal">
<mods:role id="72B11E3EEC851D5DA6A201E56AAC40D8">
<mods:roleTerm id="99C552C660F50B4068BB5761A74A856B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEDF5356F884240C57728F57A15E5F56">Evenhuis, Neal</mods:namePart>
</mods:name>
<mods:typeOfResource id="5F3177AAC2D720D4C55EF77A82F156E9">text</mods:typeOfResource>
<mods:relatedItem id="3A9CD76761E7DC9730410A1CC70204B6" type="host">
<mods:titleInfo id="C77809C735CBBABD0D68F7919CA46634">
<mods:title id="DCC64B14D6B0B9EDF5B3F53FF770D245">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1753C875D3D98FC5FAD65183AE28D9F9">
<mods:date id="1A6999E004986357CFA446C982B7EEEE">2017</mods:date>
<mods:detail id="F113612A9684DD84156AAB9370EB9E1A" type="volume">
<mods:number id="B1149C966A7D4F8F90C7E8452B7F8AED">4269</mods:number>
</mods:detail>
<mods:detail id="B5117A57402AA17D407A0EFA41781C0B" type="issue">
<mods:number id="3FA061E1A5FE932950FE8CACFFC9F817">2</mods:number>
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<mods:extent id="1D1E7702A20D9C14B015AE75C5CF0645" unit="page">
<mods:start id="E690121D8E275F56937820C62D1DA3B6">230</mods:start>
<mods:end id="DBFC8808AFFE3A539E5EA1BC179F23CC">244</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="E8E3A356AC59EDF271925B0571147EFA">journal article</mods:classification>
<mods:identifier id="49DCE01F12A5C38F5A36BAF8336348C8" type="CLB-Dataset">32999</mods:identifier>
<mods:identifier id="7675D283E442D7ED83822FCF074CA2FC" type="DOI">10.11646/zootaxa.4269.2.3</mods:identifier>
<mods:identifier id="66DA1F82297A67CE881CDB28AD6CE933" type="GBIF-Dataset">38f686ec-d435-40e6-8d43-58d7e7b8ae70</mods:identifier>
<mods:identifier id="08E216E95C0313057FBE2F3D35B83D53" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BC844C5AAD590C18AE636A78EDBEF29A" type="Zenodo-Dep">582011</mods:identifier>
<mods:identifier id="FBE584C3F57A13D8CA2A114ED8601244" type="ZooBank">01E8B213-8F9D-45E4-86CD-893716D3A295</mods:identifier>
</mods:mods>
<treatment id="834EA522FFE4FFDFDD81F94C2846F81F" ID-DOI="http://doi.org/10.5281/zenodo.6029913" ID-GBIF-Taxon="130810618" ID-Zenodo-Dep="6029913" LSID="urn:lsid:zoobank.org:act:7C4A47F5-CE81-4058-AC3B-A13771E3C3C0" httpUri="http://treatment.plazi.org/id/834EA522FFE4FFDFDD81F94C2846F81F" lastPageId="8" lastPageNumber="238" pageId="5" pageNumber="235">
<subSubSection id="43FD47BFFFE4FFD2DD81F94C2832F8DE" pageId="5" pageNumber="235" type="nomenclature">
<paragraph id="0B581434FFE4FFD2DD81F94C2A03F930" blockId="5.[151,803,1741,1801]" box="[151,803,1741,1767]" pageId="5" pageNumber="235">
<heading id="5010A358FFE4FFD2DD81F94C2A03F930" bold="true" box="[151,803,1741,1767]" fontSize="11" level="1" pageId="5" pageNumber="235" reason="1">
<emphasis id="3993C826FFE4FFD2DD81F94C2A03F930" bold="true" box="[151,803,1741,1767]" pageId="5" pageNumber="235">
<taxonomicName id="CCE76FB7FFE4FFD2DD81F94C2B98F930" authority="Gharali &amp; Evenhuis" authorityName="Gharali &amp; Evenhuis" box="[151,696,1741,1767]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="235" phylum="Arthropoda" rank="species" species="bispermatheca" status="sp. nov.">
<emphasis id="3993C826FFE4FFD2DD81F94C2893F930" bold="true" box="[151,435,1741,1767]" italics="true" pageId="5" pageNumber="235">Cyrtosia bispermatheca</emphasis>
Gharali &amp; Evenhuis
</taxonomicName>
,
<taxonomicNameLabel id="22A0755DFFE4FFD2DFD3F94C2A03F930" box="[709,803,1741,1767]" pageId="5" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B581434FFE4FFD2DD81F9712832F8DE" blockId="5.[151,803,1741,1801]" box="[151,274,1776,1801]" pageId="5" pageNumber="235">
(
<figureCitation id="93DC08B1FFE4FFD2DD89F971282AF8DE" box="[159,266,1776,1801]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1947,1969]" captionStartId-1="7.[151,250,1947,1969]" captionTargetBox-0="[174,1384,193,1900]" captionTargetBox-1="[186,1396,195,1905]" captionTargetId-0="figure@6.[174,1384,193,1905]" captionTargetId-1="figure@7.[186,1396,193,1905]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. C. bispermatheca sp. nov., male holotype: a. habitus in lateral view; b. mesonotum dorsal view; c. head dorsal view; d. abdomen in laterodorsal view (arrow points to black brown spot on halter knob); e. male abdomen dorsal view; f. wing." captionText-1="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri-0="https://zenodo.org/record/582015/files/figure.png" httpUri-1="https://zenodo.org/record/582016/files/figure.png" pageId="5" pageNumber="235">Figs. 4–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43FD47BFFFE4FFD2DD81F8B928F5F86A" pageId="5" pageNumber="235" type="materials_examined">
<paragraph id="0B581434FFE4FFD2DD81F8B928F5F86A" blockId="5.[151,1436,1848,1981]" pageId="5" pageNumber="235">
<materialsCitation id="BB8F1E69FFE4FFD2DD81F8B92C06F8A2" ID-GBIF-Occurrence="1503410535" collectingDate="2009-07-01" collectionCode="HMDS, ZMHB" collectorName="Leg. B." elevation="2400" latitude="36.35" location="Juladak village" longLatPrecision="1188" longitude="50.533333" pageId="5" pageNumber="235" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Qazvin province" typeStatus="holotype">
<emphasis id="3993C826FFE4FFD2DD81F8B92868F886" bold="true" box="[151,328,1848,1873]" pageId="5" pageNumber="235">
<typeStatus id="D45CAA96FFE4FFD2DD81F8B929F2F886" box="[151,210,1848,1873]" pageId="5" pageNumber="235">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D45CAA96FFE4FFD2DC45F8B9289BF886" box="[339,443,1848,1873]" pageId="5" pageNumber="235" type="holotype">Holotype</typeStatus>
male,
<specimenCount id="1DE1DFBDFFE4FFD2DF1BF8B92B4AF886" box="[525,618,1848,1873]" pageId="5" pageNumber="235" type="male">2 males</specimenCount>
,
<specimenCount id="1DE1DFBDFFE4FFD2DF6FF8B92BFDF886" box="[633,733,1848,1873]" pageId="5" pageNumber="235" type="female">1 female</specimenCount>
<typeStatus id="D45CAA96FFE4FFD2DFFEF8B82A68F886" box="[744,840,1849,1873]" pageId="5" pageNumber="235" type="paratype">paratype</typeStatus>
(dried by
<collectionCode id="6DF68CF1FFE4FFD2DED1F8B92D3CF887" box="[967,1052,1848,1872]" pageId="5" pageNumber="235">HMDS</collectionCode>
), Juladak rangeland,
<geoCoordinate id="6ED372F3FFE4FFD2D804F8B82C5DF886" box="[1298,1405,1848,1873]" direction="north" orientation="latitude" pageId="5" pageNumber="235" precision="925" value="36.35">N 36°21'</geoCoordinate>
,
<geoCoordinate id="6ED372F3FFE4FFD2D89DF8B829C3F8A2" direction="east" orientation="longitude" pageId="5" pageNumber="235" precision="925" value="50.533333">E 50°32'</geoCoordinate>
,
<elevation id="80CAF307FFE4FFD2DDF8F8DC2864F8A3" box="[238,324,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="235" unit="m" value="2400.0">
<quantity id="CC1FB9D1FFE4FFD2DDF8F8DC2864F8A3" box="[238,324,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="235" unit="m" value="2400.0">2400 m</quantity>
</elevation>
asl.,
<location id="0E3842EFFFE4FFD2DC97F8DD2B0BF8A2" LSID="urn:lsid:plazi:treatment:834EA522FFE4FFDFDD81F94C2846F81F:0E3842EFFFE4FFD2DC97F8DD2B0BF8A2" box="[385,555,1884,1909]" latitude="36.35" longLatPrecision="1188" longitude="50.533333" name="Juladak village" pageId="5" pageNumber="235" stateProvince="Qazvin province">Juladak village</location>
(Alamot),
<collectingDate id="6F1DCB1CFFE4FFD2DFBFF8DC2A11F8A2" box="[681,817,1884,1909]" pageId="5" pageNumber="235" value="2009-07-01">
<date id="7F5932F4FFE4FFD2DFBFF8DC2A11F8A2" box="[681,817,1884,1909]" pageId="5" pageNumber="235" value="2009-07-01">1 July 2009</date>
</collectingDate>
,
<collectingRegion id="C923DAD6FFE4FFD2DE2BF8DD2ADBF8A2" box="[829,1019,1884,1909]" country="Iran" name="Qazvin" pageId="5" pageNumber="235">Qazvin province</collectingRegion>
,
<collectorName id="A61271E2FFE4FFD2D911F8DC2D7BF8A3" box="[1031,1115,1885,1909]" pageId="5" pageNumber="235">Leg. B.</collectorName>
Gharali (
<collectionCode id="6DF68CF1FFE4FFD2D9D1F8DC2C3EF8A2" box="[1223,1310,1885,1909]" pageId="5" pageNumber="235">ZMHB</collectionCode>
)
</materialsCitation>
; 1 male, 1 female paratypes,
<materialsCitation id="BB8F1E69FFE4FFD2DC7EF8002CB8F84E" ID-GBIF-Occurrence="1503410541" box="[360,1432,1920,1945]" collectingDate="2009-07-01" collectionCode="BPBM" collectorName="Leg. B." elevation="2400" latitude="36.35" location="Juladak village" longLatPrecision="1188" longitude="50.533333" pageId="5" pageNumber="235" specimenCount="1" specimenCount-male="1" stateProvince="Qazvin province" typeStatus="paratype">
same data as holotype (in alcohol) (
<collectionCode id="6DF68CF1FFE4FFD2DE10F8012A79F84F" box="[774,857,1920,1944]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="235">BPBM</collectionCode>
),
<specimenCount id="1DE1DFBDFFE4FFD2DE79F8012A9CF84E" box="[879,956,1920,1945]" pageId="5" pageNumber="235" type="male">1 male</specimenCount>
<typeStatus id="D45CAA96FFE4FFD2DED0F8002D06F84E" box="[966,1062,1921,1945]" pageId="5" pageNumber="235" type="paratype">paratype</typeStatus>
(genitalia dissected, in alcohol)
</materialsCitation>
,
<materialsCitation id="BB8F1E69FFE4FFD2DD81F82428F5F86A" ID-GBIF-Occurrence="1503410532" box="[151,469,1956,1981]" collectingDate="2009-07-01" collectionCode="HMDS, ZMHB" collectorName="Leg. B." elevation="2400" latitude="36.35" location="Juladak village" longLatPrecision="1188" longitude="50.533333" pageId="5" pageNumber="235" specimenCount="1" stateProvince="Qazvin province" typeStatus="paratype">same data as holotype (BG).</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5F9844BCFFE7FFD1DD81F81A29F1F839" httpUri="https://zenodo.org/record/582015/files/figure.png" pageId="6" pageNumber="236" targetBox="[174,1384,193,1900]" targetPageId="6">
<paragraph id="0B581434FFE7FFD1DD81F81A29F1F839" blockId="6.[151,1436,1947,2030]" pageId="6" pageNumber="236">
<emphasis id="3993C826FFE7FFD1DD81F81A282DF866" bold="true" box="[151,269,1947,1969]" pageId="6" pageNumber="236">FIGURE 4</emphasis>
.
<taxonomicName id="CCE76FB7FFE7FFD1DC0AF81D28F0F866" box="[284,464,1947,1969]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="236" phylum="Arthropoda" rank="species" species="bispermatheca" status="sp. nov.">
<emphasis id="3993C826FFE7FFD1DC0AF81D28F0F866" box="[284,464,1947,1969]" italics="true" pageId="6" pageNumber="236">C. bispermatheca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22A0755DFFE7FFD1DCCFF81A2B0BF866" box="[473,555,1947,1969]" pageId="6" pageNumber="236" rank="species">
<emphasis id="3993C826FFE7FFD1DCCFF81A2B0BF866" bold="true" box="[473,555,1947,1969]" pageId="6" pageNumber="236">sp. nov.</emphasis>
</taxonomicNameLabel>
, male holotype:
<emphasis id="3993C826FFE7FFD1DFCDF81A2BC8F866" bold="true" box="[731,744,1947,1969]" pageId="6" pageNumber="236">a</emphasis>
. habitus in lateral view;
<emphasis id="3993C826FFE7FFD1DEFDF81A2AD9F866" bold="true" box="[1003,1017,1947,1969]" pageId="6" pageNumber="236">b</emphasis>
. mesonotum dorsal view;
<emphasis id="3993C826FFE7FFD1D81AF81A2C3EF866" bold="true" box="[1292,1310,1947,1969]" pageId="6" pageNumber="236">c.</emphasis>
head dorsal view;
<emphasis id="3993C826FFE7FFD1DDCEF83829C6F818" bold="true" box="[216,230,1977,1999]" pageId="6" pageNumber="236">d</emphasis>
. abdomen in laterodorsal view (arrow points to black brown spot on halter knob);
<emphasis id="3993C826FFE7FFD1D95AF8382D77F818" bold="true" box="[1100,1111,1977,1999]" pageId="6" pageNumber="236">e</emphasis>
. male abdomen dorsal view;
<emphasis id="3993C826FFE7FFD1D89BF8382CB5F818" bold="true" box="[1421,1429,1977,1999]" pageId="6" pageNumber="236">f</emphasis>
. wing.
</paragraph>
</caption>
<caption id="5F9844BCFFE6FFD0DD81F81A287FF839" httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="7" pageNumber="237" targetBox="[186,1396,195,1905]" targetPageId="7">
<paragraph id="0B581434FFE6FFD0DD81F81A287FF839" blockId="7.[151,1436,1947,2030]" pageId="7" pageNumber="237">
<emphasis id="3993C826FFE6FFD0DD81F81A282EF866" bold="true" box="[151,270,1947,1969]" pageId="7" pageNumber="237">FIGURE 5</emphasis>
.
<taxonomicName id="CCE76FB7FFE6FFD0DC0BF81D2B32F866" box="[285,530,1947,1969]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="237" phylum="Arthropoda" rank="species" species="bispermatheca" status="sp. nov.">
<emphasis id="3993C826FFE6FFD0DC0BF81D2B32F866" box="[285,530,1947,1969]" italics="true" pageId="7" pageNumber="237">Cyrtosia bispermatheca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22A0755DFFE6FFD0DF0DF81A2B4EF866" box="[539,622,1947,1969]" pageId="7" pageNumber="237" rank="species">
<emphasis id="3993C826FFE6FFD0DF0DF81A2B4EF866" bold="true" box="[539,622,1947,1969]" pageId="7" pageNumber="237">sp. nov.</emphasis>
</taxonomicNameLabel>
, male and female genitalia:
<emphasis id="3993C826FFE6FFD0DE8CF81A2A87F866" bold="true" box="[922,935,1947,1969]" pageId="7" pageNumber="237">a</emphasis>
. gonocoxites dorsal view (the arrow shows less sclerotized plate);
<emphasis id="3993C826FFE6FFD0DC41F8382845F818" bold="true" box="[343,357,1977,1999]" pageId="7" pageNumber="237">b</emphasis>
. epandrium dorsal view;
<emphasis id="3993C826FFE6FFD0DF65F8382B5EF818" bold="true" box="[627,638,1977,1999]" pageId="7" pageNumber="237">c</emphasis>
. phallic complex dorsal view;
<emphasis id="3993C826FFE6FFD0DED2F8382AF2F818" bold="true" box="[964,978,1977,1999]" pageId="7" pageNumber="237">d</emphasis>
. vaginal furca;
<emphasis id="3993C826FFE6FFD0D96CF8382DA5F818" bold="true" box="[1146,1157,1977,1999]" pageId="7" pageNumber="237">e</emphasis>
. spermathecal reservoir;
<emphasis id="3993C826FFE6FFD0D89BF8382CB5F818" bold="true" box="[1421,1429,1977,1999]" pageId="7" pageNumber="237">f</emphasis>
. spermathecal ducts.
</paragraph>
</caption>
<subSubSection id="43FD47BFFFE9FFDFDDD1FF162C18FE97" pageId="8" pageNumber="238" type="diagnosis">
<paragraph id="0B581434FFE9FFDFDDD1FF162C18FE97" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1FF162861FF67" bold="true" box="[199,321,151,176]" pageId="8" pageNumber="238">Diagnosis.</emphasis>
Frons black except lateral and lower margins yellow; mesonotum matte brown, with three pollinose longitudinal stripes; scutellum dark brown except small spot at posterior margin yellow; halter stem yellow, knob white with large brown spot dorsally; epandrium in dorsal view subquadrate with a short process apicoventrally; gonocoxites apicolaterally with very long black sclerotized process, apicomedially with sickle-shaped processes; Vaginal furca U-shaped, two spermathecal reservoirs, cylindrical, with a cylindrical invagination apically.
</paragraph>
</subSubSection>
<subSubSection id="43FD47BFFFE9FFDFDDD1FECD2B27FA23" pageId="8" pageNumber="238" type="description">
<paragraph id="0B581434FFE9FFDFDDD1FECD2BD5FACB" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1FECD2879FEB2" bold="true" box="[199,345,332,357]" pageId="8" pageNumber="238">Description.</emphasis>
<typeStatus id="D45CAA96FFE9FFDFDC49FECD28E7FEB2" box="[351,455,332,357]" pageId="8" pageNumber="238" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="93DC08B1FFE9FFDFDF05FECD2B77FEB3" box="[531,599,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1947,1969]" captionTargetBox="[174,1384,193,1900]" captionTargetId="figure@6.[174,1384,193,1905]" captionTargetPageId="6" captionText="FIGURE 4. C. bispermatheca sp. nov., male holotype: a. habitus in lateral view; b. mesonotum dorsal view; c. head dorsal view; d. abdomen in laterodorsal view (arrow points to black brown spot on halter knob); e. male abdomen dorsal view; f. wing." httpUri="https://zenodo.org/record/582015/files/figure.png" pageId="8" pageNumber="238">Fig. 4</figureCitation>
a). Lengths. Body:
<quantity id="CC1FB9D1FFE9FFDFDE3AFECD2AA4FEB3" box="[812,900,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="238" unit="mm" value="2.1">2.1 mm</quantity>
; wing:
<quantity id="CC1FB9D1FFE9FFDFDEC0FECD2D0FFEB3" box="[982,1071,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="238" unit="mm" value="2.2">2.2 mm</quantity>
. Head (
<figureCitation id="93DC08B1FFE9FFDFD995FECD2DF1FEB3" box="[1155,1233,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1947,1969]" captionTargetBox="[174,1384,193,1900]" captionTargetId="figure@6.[174,1384,193,1905]" captionTargetPageId="6" captionText="FIGURE 4. C. bispermatheca sp. nov., male holotype: a. habitus in lateral view; b. mesonotum dorsal view; c. head dorsal view; d. abdomen in laterodorsal view (arrow points to black brown spot on halter knob); e. male abdomen dorsal view; f. wing." httpUri="https://zenodo.org/record/582015/files/figure.png" pageId="8" pageNumber="238">Figs. 4</figureCitation>
a, c). Subglobular, slightly longer than high; eyes dichoptic, separated at vertex by 1.5 times distance between lateral ocelli; occiput, mentum, and vertex much below median ocellus black; frons black except lateral and lower margins yellow giving it a U-shaped yellow pattern; face yellow, with two brown small spots next to tip of oral cavity; oral margin narrowly yellow; antennae brown, scape minute, pedicel cylindrical, slightly wider than long, first flagellomere ellipsoid, length about 1.5 times greatest width, second flagellomere about one-half length of first, with minute apical style; proboscis sclerotized, stiff, pointed apically, equal to head height; labrum sclerotized, stiff, pointed apically, two-thirds proboscis length. Thorax. Pronotum brown; mesonotum matte brown, with three pollinose longitudinal stripes; postpronotal lobe yellow; lateral margin from postpronotal lobe to postalar callus yellow, interrupted widely with brown stripe at wing base; scutellum dark brown except small spot at posterior margin yellow; halter stem yellow, knob white with large brown spot dorsally; pleura black except yellow on following: propleuron completely, large spot posteromedially on anepisternum, upper margin of katepisternum, katepimeron and upper margin of anepimeron. Legs. Coxae brown and femora brown except extreme apices yellow; tibiae yellow; two basalmost tarsal segments yellow, remainder brownish; claws black. Wing (
<figureCitation id="93DC08B1FFE9FFDFD992FCA62DE9FC97" box="[1156,1225,807,832]" captionStart="FIGURE 4" captionStartId="6.[151,250,1947,1969]" captionTargetBox="[174,1384,193,1900]" captionTargetId="figure@6.[174,1384,193,1905]" captionTargetPageId="6" captionText="FIGURE 4. C. bispermatheca sp. nov., male holotype: a. habitus in lateral view; b. mesonotum dorsal view; c. head dorsal view; d. abdomen in laterodorsal view (arrow points to black brown spot on halter knob); e. male abdomen dorsal view; f. wing." httpUri="https://zenodo.org/record/582015/files/figure.png" pageId="8" pageNumber="238">Fig. 4</figureCitation>
f). Hyaline; Sc, Rs basally, vein separating basal cells and anal vein white to translucent; costal, radial, and medial veins brown; costa ends slightly beyond end of R4+5; vein Sc incomplete, ending at level about equal to origin of vein R2+3; vein R2+3 ends in wing margin equidistant between end of vein R1 and R4+5; cell dm open, without crossvein closing it apically; vein M1+2 one half length of fork point to margin of wing; length of cell br 1.5 times that of cell bm, anal cell open at wing margin narrowly; fringe of hair on posterior margin of wing minute. Abdomen. Tergites brown, posterior margin of all tergites narrowly yellow; sternites almost all brown, posterior margins of sternites yellow. Male genitalia. Epandrium (
<figureCitation id="93DC08B1FFE9FFDFDCCBFBAA2B02FB93" box="[477,546,1067,1092]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
b) dark brown with outer margin of epandrium and basomedial processes of gonocoxites narrowly yellow; epandrium in dorsal view subquadrate with a short process apicoventrally [pseudosurstylus]; cercus narrow, sclerotized; gonocoxites (
<figureCitation id="93DC08B1FFE9FFDFDE5EFBF22AAEFB5B" box="[840,910,1139,1164]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
a) fused medially, with yellow broad process basomedially, process possessing long whitish hairs apically; gonocoxites apicolaterally with very long black sclerotized process, apicomedially with sickle-shaped processes [gonostyli]; basal aedeagal apodeme (
<figureCitation id="93DC08B1FFE9FFDFD829FB3A2CA6FB03" box="[1343,1414,1211,1236]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
c) linear in dorsal view, as long as gonocoxal apodeme; aedeagal bulb swollen, narrowing to very long and apically curved aedeagus; lateral aedeagal apodeme triangular.
</paragraph>
<paragraph id="0B581434FFE9FFDFDDD1FAA62B27FA23" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1FAA6283EFA97" bold="true" box="[199,286,1319,1344]" pageId="8" pageNumber="238">Female</emphasis>
. similar to males but larger in size. Female genitalia. Vaginal furca U-shaped (
<figureCitation id="93DC08B1FFE9FFDFD9D3FAA62C2CFA97" box="[1221,1292,1319,1344]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
d), with two quadrate plates subapically at lateral arms, tubes same width throughout, common spermathecal duct thin (
<figureCitation id="93DC08B1FFE9FFDFD856FACA2CA2FAB3" box="[1344,1410,1355,1380]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
f), transparent, short, one-half length of spermathecal reservoir, sperm pump thin, transparent, with flared apical valve, about three times common duct length, apical duct thin, transparent, about two-thirds sperm pump length, slightly swollen next to reservoir, two spermathecal reservoirs (
<figureCitation id="93DC08B1FFE9FFDFDE76FA362A83FA07" box="[864,931,1463,1488]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[186,1396,195,1905]" captionTargetId="figure@7.[186,1396,193,1905]" captionTargetPageId="7" captionText="FIGURE 5. Cyrtosia bispermatheca sp. nov., male and female genitalia: a. gonocoxites dorsal view (the arrow shows less sclerotized plate); b. epandrium dorsal view; c. phallic complex dorsal view; d. vaginal furca; e. spermathecal reservoir; f. spermathecal ducts." httpUri="https://zenodo.org/record/582016/files/figure.png" pageId="8" pageNumber="238">Fig. 5</figureCitation>
e), cylindrical, length 1.5 times width, with a cylindrical invagination apically.
</paragraph>
</subSubSection>
<subSubSection id="43FD47BFFFE9FFDFDDD1FA7E2D68F9B7" pageId="8" pageNumber="238" type="etymology">
<paragraph id="0B581434FFE9FFDFDDD1FA7E2D68F9B7" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1FA7E286EF9CF" bold="true" box="[199,334,1535,1560]" pageId="8" pageNumber="238">Etymology.</emphasis>
The species name is composed of two words:
<emphasis id="3993C826FFE9FFDFDE4CF9812A50F9CF" box="[858,880,1536,1560]" italics="true" pageId="8" pageNumber="238">bi</emphasis>
[= two] and
<emphasis id="3993C826FFE9FFDFD914F9812DB1F9CF" box="[1026,1169,1536,1560]" italics="true" pageId="8" pageNumber="238">spermatheca</emphasis>
, referring to the female genitalia of
<taxonomicName id="CCE76FB7FFE9FFDFDC0DF9A528C0F9EB" box="[283,480,1572,1596]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="bispermatheca">
<emphasis id="3993C826FFE9FFDFDC0DF9A528C0F9EB" box="[283,480,1572,1596]" italics="true" pageId="8" pageNumber="238">C. bispermatheca</emphasis>
</taxonomicName>
that have unusually two spermathecal reservoirs. The normal state of this character in species of
<taxonomicName id="CCE76FB7FFE9FFDFDC1BF9C9284EF988" box="[269,366,1608,1631]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="3993C826FFE9FFDFDC1BF9C9284EF988" box="[269,366,1608,1631]" italics="true" pageId="8" pageNumber="238">Cyrtosia</emphasis>
</taxonomicName>
that have been dissected is having three spermathecal reservoirs.
</paragraph>
</subSubSection>
<subSubSection id="43FD47BFFFE9FFDFDDD1F9EA2A4CF91B" pageId="8" pageNumber="238" type="distribution">
<paragraph id="0B581434FFE9FFDFDDD1F9EA2A4CF91B" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1F9EA2840F953" bold="true" box="[199,352,1643,1668]" pageId="8" pageNumber="238">Distribution.</emphasis>
This species is only known from Alamot region of
<collectingRegion id="C923DAD6FFE9FFDFDEB1F9EA2D41F953" box="[935,1121,1643,1668]" country="Iran" name="Qazvin" pageId="8" pageNumber="238">Qazvin province</collectingRegion>
in
<collectingCountry id="73F054A4FFE9FFDFD990F9ED2D97F954" box="[1158,1207,1644,1667]" name="Iran" pageId="8" pageNumber="238">Iran</collectingCountry>
. This area is located on
<collectingRegion id="C923DAD6FFE9FFDFDDACF90E2828F97F" box="[186,264,1679,1704]" country="Iran" name="Alborz" pageId="8" pageNumber="238">Alborz</collectingRegion>
mountain ranges and famous for its medicinal plants. Specimens were collected by sweeping where they had aggregated on the flowers of
<taxonomicName id="CCE76FB7FFE9FFDFDF1BF9352BFDF91B" box="[525,733,1716,1740]" class="Magnoliopsida" family="Asteraceae" genus="Cichorium" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="intybus">
<emphasis id="3993C826FFE9FFDFDF1BF9352BFDF91B" box="[525,733,1716,1740]" italics="true" pageId="8" pageNumber="238">Cichorium intybus</emphasis>
</taxonomicName>
L. (
<figureCitation id="93DC08B1FFE9FFDFDE1AF9322A6EF91B" box="[780,846,1715,1740]" captionStart="FIGURE 3" captionStartId="4.[151,250,1951,1973]" captionTargetBox="[411,1169,193,1905]" captionTargetId="figure@4.[411,1169,193,1905]" captionTargetPageId="4" captionText="FIGURE 3. The different localities of Qazvin province from where type material were collected a. Nodahak rangeland (C. zieglerii sp. nov.), b. Joladak mountainous rangeland (C. bispermatheca sp. nov.), c. Loshan, rocky hills (C. baccadomus sp. nov.)." httpUri="https://zenodo.org/record/582014/files/figure.png" pageId="8" pageNumber="238">Fig. 3</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="43FD47BFFFE9FFDFDDD1F9562846F81F" pageId="8" pageNumber="238" type="discussion">
<paragraph id="0B581434FFE9FFDFDDD1F9562846F81F" blockId="8.[151,1437,151,1992]" pageId="8" pageNumber="238">
<emphasis id="3993C826FFE9FFDFDDD1F956281BF927" bold="true" box="[199,315,1751,1776]" pageId="8" pageNumber="238">Remarks.</emphasis>
This new species is easily distinguished from the congeners in
<collectingCountry id="73F054A4FFE9FFDFDEE8F9582D0BF927" box="[1022,1067,1753,1776]" name="Iran" pageId="8" pageNumber="238">Iran</collectingCountry>
by the presence of a large brown spot on the halter knob. Compared to species keyed in
<bibRefCitation id="6F7669C5FFE9FFDFDE1BF97A2A83F8C3" author="Engel" box="[781,931,1787,1812]" pageId="8" pageNumber="238" refString="Engel, O. (1933) Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. IV (3). Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 97 - 144." type="journal article" year="1933">Engel (1933)</bibRefCitation>
, only
<taxonomicName id="CCE76FB7FFE9FFDFDEFDF97D2C02F8C3" authority="Engel" authorityName="Engel" box="[1003,1314,1787,1812]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="3993C826FFE9FFDFDEFDF97D2DF5F8C4" box="[1003,1237,1788,1811]" italics="true" pageId="8" pageNumber="238">Cyrtosia canariensis</emphasis>
Engel
</taxonomicName>
, originally described as a variety of
<taxonomicName id="CCE76FB7FFE9FFDFDCA7F8A12A1BF8EF" authority="Bezzi, 1926" authorityName="Bezzi" authorityYear="1926" box="[433,827,1823,1848]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="tetragramma">
<emphasis id="3993C826FFE9FFDFDCA7F8A12B8BF8EF" box="[433,683,1824,1848]" italics="true" pageId="8" pageNumber="238">Cyrtosia tetragramma</emphasis>
<bibRefCitation id="6F7669C5FFE9FFDFDFA4F89E2A1BF8EF" author="Bezzi" box="[690,827,1823,1848]" pageId="8" pageNumber="238" refString="Bezzi, M. (1926) Notes additionelles sur les Bombyliidae (Dipt.) d'Egypte. Bulletin de la Societe Entomologique d'Egypte, 9, 244 - 273." type="journal article" year="1926">Bezzi, 1926</bibRefCitation>
</taxonomicName>
, has dark spot on halter knob (the original description states: “Der Schwingerkopf ist auf seiner Mitte deutlich geschwärzt”).
<taxonomicName id="CCE76FB7FFE9FFDFD90BF8C52C16F88B" box="[1053,1334,1860,1884]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="bispermatheca">
<emphasis id="3993C826FFE9FFDFD90BF8C52C16F88B" box="[1053,1334,1860,1884]" italics="true" pageId="8" pageNumber="238">Cyrtosia bispermatheca</emphasis>
</taxonomicName>
is also distinguished from this Canary Islands species by the yellow color at the posterior margin of scutellum (scutellum completely dark brown in
<taxonomicName id="CCE76FB7FFE9FFDFDCD6F80D2B42F873" box="[448,610,1932,1956]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="3993C826FFE9FFDFDCD6F80D2B42F873" box="[448,610,1932,1956]" italics="true" pageId="8" pageNumber="238">C. canariensis</emphasis>
</taxonomicName>
) and the yellow frons with a black stripe medially (frons almost all black in
<taxonomicName id="CCE76FB7FFE9FFDFDDA2F8312875F810" box="[180,341,1968,1991]" class="Insecta" family="Bombyliidae" genus="Cyrtosia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="238" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="3993C826FFE9FFDFDDA2F8312875F810" box="[180,341,1968,1991]" italics="true" pageId="8" pageNumber="238">C. canariensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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