182 lines
30 KiB
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182 lines
30 KiB
XML
<document id="ADE92F7DCACA96009C0583EDC792922E" ID-CLB-Dataset="22415" ID-DOI="10.11646/zootaxa.4768.3.10" ID-GBIF-Dataset="8799e916-86ad-4a46-951a-885bb61c4cf8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784018" ID-ZooBank="FD25D951-D6EE-4151-B516-39D0345B6B5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576324800" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Kairišs, Kristaps & Bukejs, Andris" docDate="2020" docId="6C2487CC232D2013FF5FFC1BFC38F882" docLanguage="en" docName="zootaxa.4768.3.10.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neomida Latreille 1829" docType="treatment" docVersion="6" lastPageNumber="436" masterDocId="901DFFB4232C2012FFC8FF96FFDDFF85" masterDocTitle="The oldest fossil darkling beetle of the genus Neomida Latreille, 1829 (Coleoptera: Tenebrionidae) from Eocene Baltic amber examined with X-ray microtomography" masterLastPageNumber="442" masterPageNumber="435" pageNumber="436" updateTime="1698838317980" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="D1B2E492FA2E7C0D54D75B41F8DF8EF3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8B560E6CBE9F45A875B243AB587C052F">
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<mods:title id="9832BB3F2E443506FC0BF196E99E4B69">The oldest fossil darkling beetle of the genus Neomida Latreille, 1829 (Coleoptera: Tenebrionidae) from Eocene Baltic amber examined with X-ray microtomography</mods:title>
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<mods:name id="4AF018E3CCBDF719A74F42112CB756F1" type="personal">
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<mods:roleTerm id="ECB490EA88CA2BC1083639FA82546CCB">Author</mods:roleTerm>
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<mods:namePart id="629770A53092F297059E2AECABFC549B">Nabozhenko, Maxim V.</mods:namePart>
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<mods:affiliation id="713F0F2A721A2B533215AA3DB55F16B0">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev Str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. & Dagestan State University, 43 a M. Gadzhiev Str., Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
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<mods:name id="1AC2BD2D5D14B94F9767704B7E83FD6F" type="personal">
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<mods:roleTerm id="D18F5493F2C279DA870086AFD2A2F0C7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="64F4D96BD826E9222F25C52683925EC7">Kairišs, Kristaps</mods:namePart>
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<mods:affiliation id="B7582D78CDCEB9158A9210D1A93A3080">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas iela 13, Daugavpils, LV- 5401, Latvia. kr. kairiss @ gmail. com; https: // orcid. org / 0000 - 0003 - 2329 - 1122</mods:affiliation>
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<mods:roleTerm id="AF939785002CC067EFEB877B08B7ADA5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C6B249393AE31714C4624C43B81AE4AB">Bukejs, Andris</mods:namePart>
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<mods:affiliation id="5E5C57681D72CCA762AF9BB731740537">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas iela 13, Daugavpils, LV- 5401, Latvia. kr. kairiss @ gmail. com; https: // orcid. org / 0000 - 0003 - 2329 - 1122</mods:affiliation>
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<mods:typeOfResource id="5C034487F4EEFCF341F9EE5CCB1CFED0">text</mods:typeOfResource>
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<mods:title id="F317B79034FE3678E552CD313DF6D8BE">Zootaxa</mods:title>
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<mods:part id="4C8A85CCA97A52803C46EDDC5778B0C1">
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<mods:date id="2AB23EB492FF9DC8A11C579B49916360">2020</mods:date>
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<mods:number id="29B16DC75D9E4B44F953E48C0820609C">2020-05-04</mods:number>
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<mods:number id="32D9DCE56F4BDFF827A7981A214FBACA">4768</mods:number>
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<mods:identifier id="3031CC9439EFC8FD6108E6DC32EF3886" type="DOI">10.11646/zootaxa.4768.3.10</mods:identifier>
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<mods:identifier id="BCFD2EFEDCE68D1009639F911BC0399E" type="GBIF-Dataset">8799e916-86ad-4a46-951a-885bb61c4cf8</mods:identifier>
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<treatment id="6C2487CC232D2013FF5FFC1BFC38F882" ID-DOI="http://doi.org/10.5281/zenodo.3794860" ID-GBIF-Taxon="163885947" ID-Zenodo-Dep="3794860" LSID="urn:lsid:plazi:treatment:6C2487CC232D2013FF5FFC1BFC38F882" httpUri="http://treatment.plazi.org/id/6C2487CC232D2013FF5FFC1BFC38F882" lastPageNumber="436" pageId="1" pageNumber="436">
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<subSubSection id="AC976551232D2013FF5FFC1BFA8FFA8A" pageId="1" pageNumber="436" type="nomenclature">
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<paragraph id="E43236DA232D2013FF5FFC1BFDC5FC22" blockId="1.[151,536,909,936]" box="[151,536,909,936]" pageId="1" pageNumber="436">
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<heading id="BF7A81B6232D2013FF5FFC1BFDC5FC22" bold="true" box="[151,536,909,936]" fontSize="11" level="1" pageId="1" pageNumber="436" reason="1">
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<emphasis id="D6F9EAC8232D2013FF5FFC1BFDC5FC22" bold="true" box="[151,536,909,936]" pageId="1" pageNumber="436">
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Genus
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<taxonomicName id="238D4D59232D2013FF26FC1BFDC5FC22" authority="Latreille, 1829" authorityName="Latreille" authorityYear="1829" box="[238,536,909,936]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
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<emphasis id="D6F9EAC8232D2013FF26FC1BFE86FC2D" bold="true" box="[238,347,909,936]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
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Latreille, 1829
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E43236DA232D2013FF5FFC43FA8FFA8A" blockId="1.[151,1437,981,1799]" pageId="1" pageNumber="436">
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<emphasis id="D6F9EAC8232D2013FF5FFC43FF3CFC6A" bold="true" box="[151,225,981,1007]" pageId="1" pageNumber="436">Notes.</emphasis>
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The specimen considered here was assigned to the subfamily
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<taxonomicName id="238D4D59232D2013FC09FC43FBE1FC6A" authorityName="Latreille" authorityYear="1802" box="[961,1084,981,1007]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="subFamily" subFamily="Diaperinae">Diaperinae</taxonomicName>
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and subtribe
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<taxonomicName id="238D4D59232D2013FB28FC43FA97FC6A" authorityName="Latreille" authorityYear="1802" box="[1248,1354,981,1007]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="tribe" tribe="Diaperini">Diaperini</taxonomicName>
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within
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<taxonomicName id="238D4D59232D2013FF5FFC6FFEEAFB96" authorityName="Latreille" authorityYear="1802" box="[151,311,1017,1043]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
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on the basis of the following combination of morphological characters: tarsal formula 5-5-4; head armed between eyes (
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<figureCitation id="7CB62A5F232D2013FE44FB8BFE35FBB2" box="[396,488,1053,1079]" captionStart="FIGURE 1" captionStartId="2.[151,250,1508,1533]" captionTargetBox="[151,1436,414,1480]" captionTargetId="figure@2.[151,1436,414,1480]" captionTargetPageId="2" captionText="FIGURE 1. Neomida groehni sp. nov., holotype, photomicrograph: A – habitus, latero-dorsal view; B – details of forebody, dorsal view; C – habitus, ventral view. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784020" httpUri="https://zenodo.org/record/3784020/files/figure.png" pageId="1" pageNumber="436">Figs 1B</figureCitation>
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,
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<figureCitation id="7CB62A5F232D2013FE3EFB8BFDCEFBB2" box="[502,531,1053,1079]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">2F</figureCitation>
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,
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<figureCitation id="7CB62A5F232D2013FDE8FB8BFD9EFBB2" box="[544,579,1053,1079]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">3B</figureCitation>
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); base of antennomere 1 dorsally covered by lateral extension of gena (
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<figureCitation id="7CB62A5F232D2013FAA7FB8BFF66FBDE" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Fig. 2B</figureCitation>
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); antennae gradually widened and clubbed, with slightly asymmetrical antennomeres 5–11 (
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<figureCitation id="7CB62A5F232D2013FB7AFBD7FAD6FBDE" box="[1202,1291,1089,1115]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Fig. 3A</figureCitation>
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); mouthparts tenebrioid, open (
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<figureCitation id="7CB62A5F232D2013FEABFBF3FE01FBFA" box="[355,476,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Figs 2B,E</figureCitation>
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); mentum with median elevation (
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<figureCitation id="7CB62A5F232D2013FCA5FBF3FC38FBFA" box="[877,997,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Figs 2B,E</figureCitation>
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); labro-epistomal membrane exposed; procoxal cavities externally closed, with ventrally convex prosternal process (
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<figureCitation id="7CB62A5F232D2013FBC4FB1FFBBBFB26" box="[1036,1126,1161,1187]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Fig. 3C</figureCitation>
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); mesocoxal cavities partly closed by mesoventrite, mesepimeron and metaventrite; mesotrochantine present (
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<figureCitation id="7CB62A5F232D2013FBFBFB3BFB51FB42" box="[1075,1164,1197,1223]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Fig. 3C</figureCitation>
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); each elytron with nine striae and short striole (
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<figureCitation id="7CB62A5F232D2013FE55FB47FE39FB6E" box="[413,484,1233,1259]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Figs 2</figureCitation>
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A–G); legs not fossorial; abdominal membranes between ventrites 3–5 exposed (supporting presence of defensive glands) (
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<figureCitation id="7CB62A5F232D2013FD82FB63FD7FFA8A" box="[586,674,1269,1295]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Fig. 2E</figureCitation>
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); and apical piece of aedeagus without ventral alae (
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<figureCitation id="7CB62A5F232D2013FB23FB63FA9BFA8A" box="[1259,1350,1269,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1009,1034]" captionTargetBox="[151,1436,181,984]" captionTargetId="figure@6.[151,1436,181,984]" captionTargetPageId="6" captionText="FIGURE 4. Neomida groehni sp. nov., holotype, aedeagus, X-ray micro-CT rendering: A – ventral view; B – dorsal view; C – lateral view; D – apical piece. Abbreviations: ap – apical piece, bp – basal piece, ml – median lobe." figureDoi="http://doi.org/10.5281/zenodo.3784026" httpUri="https://zenodo.org/record/3784026/files/figure.png" pageId="1" pageNumber="436">Fig. 4A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="AC976551232D2013FF5FFA8FFC38F882" pageId="1" pageNumber="436" type="description">
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<paragraph id="E43236DA232D2013FF5FFA8FFEDEF9F2" blockId="1.[151,1437,981,1799]" pageId="1" pageNumber="436">
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The genus
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<taxonomicName id="238D4D59232D2013FEDCFA8FFEA4FAB6" authorityName="Latreille" authorityYear="1829" box="[276,377,1305,1331]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
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<emphasis id="D6F9EAC8232D2013FEDCFA8FFEA4FAB6" box="[276,377,1305,1331]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
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</taxonomicName>
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has a diverse combination of characters, and sometimes can be confused with
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<taxonomicName id="238D4D59232D2013FB3EFA8FFE58FAD2" authority="Laporte et Brulle, 1831" authorityName="Laporte et Brulle" authorityYear="1831" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
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<emphasis id="D6F9EAC8232D2013FB3EFA8FFAB3FAB6" box="[1270,1390,1305,1331]" italics="true" pageId="1" pageNumber="436">Platydema</emphasis>
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Laporte et Brullé, 1831
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</taxonomicName>
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. This is because some Neotropical and Oriental
|
||
<emphasis id="D6F9EAC8232D2013FC60FAABFBFBFAD2" box="[936,1062,1341,1367]" italics="true" pageId="1" pageNumber="436">
|
||
<taxonomicName id="238D4D59232D2013FC60FAABFBFEFAD2" authorityName="Laporte et Brulle" authorityYear="1831" box="[936,1059,1341,1367]" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">Platydema</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
such as
|
||
<taxonomicName id="238D4D59232D2013FB41FAABFEE0FAF9" authority="(Grimm 2016)" baseAuthorityName="Grimm" baseAuthorityYear="2016" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="species" species="orientaloides">
|
||
<emphasis id="D6F9EAC8232D2013FB41FAABFA41FAD2" box="[1161,1436,1341,1367]" italics="true" pageId="1" pageNumber="436">Platydema orientaloides</emphasis>
|
||
(
|
||
<bibRefCitation id="801C4B2B232D2013FF57FAF7FEE8FAF9" author="Grimm, R." box="[159,309,1377,1404]" pageId="1" pageNumber="436" pagination="185 - 190" refId="ref3760" refString="Grimm, R. (2016) New and little-known species of Tenebrionidae (Coleoptera) from Borneo (6). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 9, 185 - 190. https: // doi. org / 10.18476 / sbna. v 9. a 10" type="journal article" year="2016">Grimm 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, also have a subcylindrical body and subparallel elytra as in
|
||
<taxonomicName id="238D4D59232D2013FC20FAF7FB26FAF9" authority="(Grimm 2016)" baseAuthorityName="Grimm" baseAuthorityYear="2016" box="[1000,1275,1377,1404]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FC20FAF7FB90FAFE" box="[1000,1101,1377,1403]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
|
||
(
|
||
<bibRefCitation id="801C4B2B232D2013FB94FAF7FB2EFAF9" author="Grimm, R." box="[1116,1267,1377,1404]" pageId="1" pageNumber="436" pagination="185 - 190" refId="ref3760" refString="Grimm, R. (2016) New and little-known species of Tenebrionidae (Coleoptera) from Borneo (6). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 9, 185 - 190. https: // doi. org / 10.18476 / sbna. v 9. a 10" type="journal article" year="2016">Grimm 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, the new fossil species (male) distinctly belongs to
|
||
<taxonomicName id="238D4D59232D2013FD55FA13FCDFFA1A" baseAuthorityName="Grimm" baseAuthorityYear="2016" box="[669,770,1413,1439]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FD55FA13FCDFFA1A" box="[669,770,1413,1439]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
|
||
</taxonomicName>
|
||
based on the presence of very small paired marginal epistomal tubercles (
|
||
<figureCitation id="7CB62A5F232D2013FEF5FA3FFE69FA46" box="[317,436,1449,1475]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Figs 3A,B</figureCitation>
|
||
), and the structure of the prosternal process, which is not beaded but convex and sharply declivitous immediately posterior to the procoxae, with the apex of the prosternal process bearing a tubercle at the level of the basal part of the procoxae (
|
||
<figureCitation id="7CB62A5F232D2013FD80FA67FD7DF98E" box="[584,672,1521,1547]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Fig. 3C</figureCitation>
|
||
).
|
||
<taxonomicName id="238D4D59232D2013FD7BFA67FCF6F98E" authorityName="Laporte et Brulle" authorityYear="1831" box="[691,811,1521,1547]" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FD7BFA67FCF6F98E" box="[691,811,1521,1547]" italics="true" pageId="1" pageNumber="436">Platydema</emphasis>
|
||
</taxonomicName>
|
||
has a prosternal process that is beaded, and “prominent, prolonged caudally horizontal or deflected behind” procoxae according to
|
||
<bibRefCitation id="801C4B2B232D2013FC29F983FB73F9AA" author="Triplehorn, C. A." box="[993,1198,1556,1583]" pageId="1" pageNumber="436" pagination="349 - 457" refId="ref3979" refString="Triplehorn, C. A. (1965) Revision of Diaperini of America North of Mexico with notes on extralimital species (Coleoptera: Tenebrionidae). Proceedings of the United States National Museum, Smithonian Institution, 117, 349 - 457. https: // doi. org / 10.5479 / si. 00963801.117 - 3515.349" type="journal article" year="1965">Triplehorn (1965)</bibRefCitation>
|
||
. In addition, the new fossil species has fovea intercoxal process of mesoventrite (
|
||
<figureCitation id="7CB62A5F232D2013FCFBF9AFFC54F9D6" box="[819,905,1593,1619]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1909]" captionTargetBox="[164,1423,181,1858]" captionTargetId="figure@5.[164,1423,181,1858]" captionTargetPageId="5" captionText="FIGURE 3. Neomida groehni sp. nov., holotype, X-ray micro-CT rendering:A – details of forebody, dorso-lateral view, arrows show marginal epistomal tubercles; B – head, dorsal view, arrows show marginal epistomal tubercles; C – pro-mesothoracic junction, ventral view. Abbreviations: Pstr – prosternal process, MstrE – mesosternal Y-shaped elevation, MstrF – mesosternal intercoxal fovea, Mtrch - mesotrochantine. Not reproduced to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3784024" httpUri="https://zenodo.org/record/3784024/files/figure.png" pageId="1" pageNumber="436">Fig. 3C</figureCitation>
|
||
), which is also found in at least two Palaearctic
|
||
<taxonomicName id="238D4D59232D2013FF5FF9CBFF21F9F2" baseAuthorityName="Grimm" baseAuthorityYear="2016" box="[151,252,1629,1655]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FF5FF9CBFF21F9F2" box="[151,252,1629,1655]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E43236DA232D2013FF0FF917FC38F882" blockId="1.[151,1437,981,1799]" pageId="1" pageNumber="436">
|
||
Usually, species of
|
||
<taxonomicName id="238D4D59232D2013FE6CF917FDD4F91E" baseAuthorityName="Grimm" baseAuthorityYear="2016" box="[420,521,1665,1691]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FE6CF917FDD4F91E" box="[420,521,1665,1691]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
|
||
</taxonomicName>
|
||
have epipleura that are abruptly abbreviated at or near the last abdominal suture (between ventrites 4 and 5), while epipleura in
|
||
<taxonomicName id="238D4D59232D2013FD65F933FCF8F93A" authorityName="Laporte et Brulle" authorityYear="1831" box="[685,805,1701,1727]" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FD65F933FCF8F93A" box="[685,805,1701,1727]" italics="true" pageId="1" pageNumber="436">Platydema</emphasis>
|
||
</taxonomicName>
|
||
are extending or almost extending the elytral apex. The new Eocene species has epipleura similar to those in
|
||
<taxonomicName id="238D4D59232D2013FD36F95FFCABF966" authorityName="Laporte et Brulle" authorityYear="1831" box="[766,886,1737,1763]" class="Insecta" family="Tenebrionidae" genus="Platydema" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FD36F95FFCABF966" box="[766,886,1737,1763]" italics="true" pageId="1" pageNumber="436">Platydema</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="7CB62A5F232D2013FC4FF95FFC24F966" box="[903,1017,1737,1763]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1901]" captionTargetBox="[151,1436,237,1851]" captionTargetId="figure@4.[151,1436,237,1851]" captionTargetPageId="4" captionText="FIGURE 2. Neomida groehni sp. nov., holotype, habitus, X-ray micro-CT rendering: A – dorsal view; B – ventral view with legs; C – lateral view, without legs and antennae; D – the same, with legs an antennae; E – ventral view without legs and antennae; F – anterior view; G – caudal view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3784022" httpUri="https://zenodo.org/record/3784022/files/figure.png" pageId="1" pageNumber="436">Fig. 2B,E</figureCitation>
|
||
), but at least two extant Neotropical
|
||
<taxonomicName id="238D4D59232D2013FF5FF97BFF21F882" baseAuthorityName="Grimm" baseAuthorityYear="2016" box="[151,252,1773,1799]" class="Insecta" family="Tenebrionidae" genus="Neomida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6F9EAC8232D2013FF5FF97BFF21F882" box="[151,252,1773,1799]" italics="true" pageId="1" pageNumber="436">Neomida</emphasis>
|
||
</taxonomicName>
|
||
species also have the same epipleural structure (
|
||
<bibRefCitation id="801C4B2B232D2013FCD4F97BFC05F882" author="Triplehorn, C. A." box="[796,984,1772,1799]" pageId="1" pageNumber="436" pagination="349 - 457" refId="ref3979" refString="Triplehorn, C. A. (1965) Revision of Diaperini of America North of Mexico with notes on extralimital species (Coleoptera: Tenebrionidae). Proceedings of the United States National Museum, Smithonian Institution, 117, 349 - 457. https: // doi. org / 10.5479 / si. 00963801.117 - 3515.349" type="journal article" year="1965">Triplehorn 1965</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |