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<document id="6F588D52111219178D299B4C326A9F08" ID-CLB-Dataset="6557" ID-DOI="10.5281/zenodo.4503028" ID-GBIF-Dataset="479d8c3b-8194-4f81-bed5-d157d5aa021e" ID-ISSN="2345-7600" ID-Zenodo-Dep="4503028" ID-ZooBank="C0EDB2F2-78E4-43A0-A8C4-E4165DF6406F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612459825365" checkinUser="carolina" docAuthor="Malay, Maria Celia (Machel) D., Rahayu, Dwi Listyo &amp; Chan, Tin-Yam" docDate="2018" docId="72156531297CFF9AFC1BF92AFA4CB395" docLanguage="en" docName="RafflesBZool.66.23-65.pdf.imf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Dardanus balhibuon Celia &amp; Machel &amp; Malay &amp; Chan 2018, new species" docType="treatment" docUuid="93120EF2-BB52-4883-BF3C-10C43571F87E" docUuidSource="ZooBank" docVersion="12" lastPageNumber="50" masterDocId="8E2C1D492969FF81FFB5FF97FFE1B475" masterDocTitle="Hermit crabs of the genera Calcinus Dana, Clibanarius Dana, and Dardanus Paulson from the PANGLAO 2004 Expedition, with description of a new species and a checklist of the hermit crabs of the Philippines (Crustacea: Anomura: Paguroidea)" masterLastPageNumber="65" masterPageNumber="23" pageNumber="44" updateTime="1698926965756" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A1A61995FA171E7A40262748F80BDE6B">Hermit crabs of the genera Calcinus Dana, Clibanarius Dana, and Dardanus Paulson from the PANGLAO 2004 Expedition, with description of a new species and a checklist of the hermit crabs of the Philippines (Crustacea: Anomura: Paguroidea)</mods:title>
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<mods:namePart id="6D2571054202585D9610DCFB3DD68EB7">Malay, Maria Celia (Machel) D.</mods:namePart>
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<mods:namePart id="75D938F5CFEBAF004485329B78D52C40">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="3A15F0C54613DDBE965F04146C90184B">Chan, Tin-Yam</mods:namePart>
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<treatment id="72156531297CFF9AFC1BF92AFA4CB395" ID-DOI="http://doi.org/10.5281/zenodo.4776950" ID-GBIF-Taxon="182253627" ID-Zenodo-Dep="4776950" LSID="urn:lsid:zoobank.org:act:93120EF2-BB52-4883-BF3C-10C43571F87E" httpUri="http://treatment.plazi.org/id/72156531297CFF9AFC1BF92AFA4CB395" lastPageId="27" lastPageNumber="50" pageId="21" pageNumber="44">
<subSubSection id="B2A687AC297CFF94FC1BF92AFACBB2A0" box="[942,1322,1725,1749]" pageId="21" pageNumber="44" type="nomenclature">
<paragraph id="FA03D427297CFF94FC1BF92AFACBB2A0" blockId="21.[942,1322,1725,1781]" box="[942,1322,1725,1749]" pageId="21" pageNumber="44">
<heading id="A14B634B297CFF94FC1BF92AFACBB2A0" bold="true" box="[942,1322,1725,1749]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="44" reason="2">
<emphasis id="C8C80835297CFF94FC1BF92AFACBB2A0" bold="true" box="[942,1322,1725,1749]" pageId="21" pageNumber="44">
<taxonomicName id="3DBCAFA4297CFF94FC1BF92AFB77B2A0" ID-CoL="b5998c12-12c8-4264-a422-6dbe12547b2a" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[942,1174,1725,1749]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="44" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C80835297CFF94FC1BF92AFB77B2A0" bold="true" box="[942,1174,1725,1749]" italics="true" pageId="21" pageNumber="44">Dardanus balhibuon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D3FBB54E297CFF94FB10F92AFACBB2A0" box="[1189,1322,1725,1749]" pageId="21" pageNumber="44" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="B2A687AC297CFF94FBB4F94AFB36B280" box="[1025,1239,1757,1781]" pageId="21" pageNumber="44" type="description">
<paragraph id="FA03D427297CFF94FBB4F94AFB36B280" blockId="21.[942,1322,1725,1781]" box="[1025,1239,1757,1781]" pageId="21" pageNumber="44">
(
<figureCitation id="6287C8A2297CFF94FBBCF94AFB88B280" box="[1033,1129,1757,1781]" captionStart="Fig" captionStartId="17.[113,148,1651,1673]" captionTargetBox="[113,1451,189,1636]" captionTargetId="figure-90@17.[113,1451,189,1636]" captionTargetPageId="17" captionText="Fig. 4. A, Dardanus sanguinolentus (Quoy &amp; Gaimard, 1824). Female ovig. SL 6.3 mm, stn R6; B, Dardanus balhibuon, n. sp. Holotype female SL 14.3 mm, stn R38, NMCR 40109, preserved colouration; C, Dardanus megistos (Herbst, 1804). Male SL 11.0 mm, stn M8; D, Dardanus pedunculatus (Herbst, 1804). Male SL 13.4 mm, stn N3; E, Dardanus scutellatus. Male SL 9.8 mm, stn D12; F, Dardanus woodmasoni (Alcock, 1905). Male SL 5.8 mm, stn R76." figureDoi="http://doi.org/10.5281/zenodo.4503040" httpUri="https://zenodo.org/record/4503040/files/figure.png" pageId="21" pageNumber="44">Figs. 4B</figureCitation>
,
<figureCitation id="6287C8A2297CFF94FBC3F94AFB78B280" box="[1142,1177,1757,1781]" captionStart="Fig" captionStartId="18.[136,171,1335,1357]" captionTargetBox="[139,777,188,1321]" captionTargetId="figure-125@18.[122,790,183,1324]" captionTargetPageId="18" captionText="Fig. 5. Lateral view of left third pereopod. A, Dardanus lagopodes (Forskål, 1775). Male SL 9.8 mm, stn N2, ZRC 2015.0489; B, Dardanus lagopodes (Forskål, 1775). Female SL 7.6 mm, stn R1, ZRC 2015.0488; C, Dardanus sanguinolentus (Quoy &amp; Gaimard, 1824). Male SL 7.8 mm, stn R38, ZRC 2015.0491; D, Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503046" httpUri="https://zenodo.org/record/4503046/files/figure.png" pageId="21" pageNumber="44">5D</figureCitation>
,
<figureCitation id="6287C8A2297CFF94FB13F94AFB2EB280" box="[1190,1231,1757,1781]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="22.[135,170,2022,2044]" captionStartId-1="24.[135,170,2064,2086]" captionStartId-2="25.[113,148,1993,2015]" captionTargetBox-0="[134,1471,189,2016]" captionTargetBox-1="[149,1461,189,2049]" captionTargetBox-2="[104,1451,262,1977]" captionTargetId-0="figure-8@22.[122,1489,183,2025]" captionTargetId-1="figure-8@24.[144,1464,186,2051]" captionTargetId-2="figure-14@25.[101,1458,248,1987]" captionTargetPageId-0="22" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." captionText-1="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." captionText-2="Fig. 8. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left second pereopod, lateral view; B, Propodus of left third pereopod, lateral view. Scale bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4503049" figureDoi-1="http://doi.org/10.5281/zenodo.4503051" figureDoi-2="http://doi.org/10.5281/zenodo.4503053" httpUri-0="https://zenodo.org/record/4503049/files/figure.png" httpUri-1="https://zenodo.org/record/4503051/files/figure.png" httpUri-2="https://zenodo.org/record/4503053/files/figure.png" pageId="21" pageNumber="44">68</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2A687AC297CFF94FC98F88AFB6EB300" pageId="21" pageNumber="44" type="materials_examined">
<paragraph id="FA03D427297CFF94FC98F88AFB6EB300" blockId="21.[811,1450,1821,1909]" pageId="21" pageNumber="44">
<emphasis id="C8C80835297CFF94FC98F88AFBEBB340" bold="true" box="[813,1034,1821,1845]" pageId="21" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="4AD4DE7A297CFF94FBBBF88AFB6AB300" ID-GBIF-Occurrence="3115205340" collectingDate="2004-06-11" collectionCode="NMCR" collectorName="Pamilacan I." elevation="22" latitude="9.49" location="Bohol Prov." longLatPrecision="129" longitude="123.933334" pageId="21" pageNumber="44" specimenCode="NMCR 40109" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
Female,
<typeStatus id="25076A85297CFF94FBDDF88AFB24B340" box="[1128,1221,1821,1845]" pageId="21" pageNumber="44" type="holotype">holotype</typeStatus>
SL
<quantity id="3D4479C2297CFF94FB45F88AFAACB340" box="[1264,1357,1821,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="21" pageNumber="44" unit="mm" value="14.3">14.3 mm</quantity>
(
<specimenCode id="AA1A7C5C297CFF94FAEFF88AFC97B320" collectionCode="NMCR" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="21" pageNumber="44" type="Herbarium">NMCR 40109</specimenCode>
), stn R38,
<collectorName id="5749B1F1297CFF94FC4EF8AAFB6FB320" box="[1019,1166,1853,1877]" pageId="21" pageNumber="44">Pamilacan I.</collectorName>
,
<collectingRegion id="38781AC5297CFF94FB2AF8AAFACFB320" box="[1183,1326,1853,1877]" country="Philippines" name="Bohol" pageId="21" pageNumber="44">Bohol Prov.</collectingRegion>
,
<geoCoordinate id="9F88B2E0297CFF94FA8BF8AAFA47B320" box="[1342,1446,1852,1877]" degrees="9" direction="north" minutes="29.4" orientation="latitude" pageId="21" pageNumber="44" precision="92" value="9.49">9°29.4N</geoCoordinate>
,
<geoCoordinate id="9F88B2E0297CFF94FC98F8CAFC46B300" box="[813,935,1884,1909]" degrees="123" direction="east" minutes="56.0" orientation="longitude" pageId="21" pageNumber="44" precision="92" value="123.933334">123°56.0E</geoCoordinate>
,
<quantity id="3D4479C2297CFF94FC06F8CAFBE9B300" box="[947,1032,1885,1909]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.7" metricValueMin="0.6" pageId="21" pageNumber="44" unit="m" value="21.5" valueMax="37.0" valueMin="6.0">
<elevation id="71913314297CFF94FC06F8CAFBE9B300" box="[947,1032,1885,1909]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.7" metricValueMin="0.6" pageId="21" pageNumber="44" unit="m" value="21.5" valueMax="37.0" valueMin="6.0">637 m</elevation>
</quantity>
,
<date id="8E02F2E7297CFF94FBA0F8CAFB6AB300" box="[1045,1163,1885,1909]" pageId="21" pageNumber="44" value="2004-06-11">
<collectingDate id="9E460B0F297CFF94FBA0F8CAFB6AB300" box="[1045,1163,1885,1909]" pageId="21" pageNumber="44" value="2004-06-11">11.vi.2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2A687AC297CFF9AFC98F80AFD92B2B5" lastPageId="27" lastPageNumber="50" pageId="21" pageNumber="44" type="description">
<paragraph id="FA03D427297CFF96FC98F80AFD11B540" blockId="21.[813,1452,1949,2101]" lastBlockId="23.[112,752,189,309]" lastPageId="23" lastPageNumber="46" pageId="21" pageNumber="44">
<emphasis id="C8C80835297CFF94FC98F80AFC59B3C0" bold="true" box="[813,952,1949,1973]" pageId="21" pageNumber="44">Description.</emphasis>
Shield (
<figureCitation id="6287C8A2297CFF94FBA3F80AFB8CB3C0" box="[1046,1133,1949,1973]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="21" pageNumber="44">Fig. 6A</figureCitation>
) 0.9 times longer than wide; anterior margin between rostrum and lateral projections shallowly concave; lateral margins slightly convex, slightly irregular, with dense tufts of long setae. Posterior margin rounded. Dorsal surface of shield somewhat inflated, with tufts of long setae, strongly calcified; Y-shaped line present posteriorly. Rostral lobe weakly produced. Lateral projections large, bluntly triangular, produced. Posterior carapace lateral elements well calcified, unarmed. Branchiostegites unarmed.
</paragraph>
<caption id="AEC384AF297FFF97FF32F871FCC4BC40" ID-DOI="http://doi.org/10.5281/zenodo.4503049" ID-Zenodo-Dep="4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="22" pageNumber="45" startId="22.[135,170,2022,2044]" targetBox="[134,1471,189,2016]" targetPageId="22">
<paragraph id="FA03D427297FFF97FF32F871FCC4BC40" blockId="22.[135,1474,2022,2102]" pageId="22" pageNumber="45">
Fig. 6.
<taxonomicName id="3DBCAFA4297FFF97FF64F871FE7BB38E" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[209,410,2022,2043]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="45" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C80835297FFF97FF64F871FE7BB38E" box="[209,410,2022,2043]" italics="true" pageId="22" pageNumber="45">Dardanus balhibuon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D3FBB54E297FFF97FE1CF870FDC3B389" box="[425,546,2022,2044]" pageId="22" pageNumber="45" rank="species">new species</taxonomicNameLabel>
, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="FA03D427297EFF96FFC4FECAFDC2B660" blockId="23.[113,753,349,533]" pageId="23" pageNumber="46">
Ocular peduncles (
<figureCitation id="6287C8A2297EFF96FEF8FECAFE49B500" box="[333,424,349,373]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6A</figureCitation>
) 0.8 times length of shield, subcylindrical, somewhat inflated distally, diameter of corneas slightly less than 0.2 of ocular peduncles length. Ocular acicles (
<figureCitation id="6287C8A2297EFF96FEA8FE2AFE90B5A0" box="[285,369,445,469]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6B</figureCitation>
) broad, distal margins each with 3 spines, with fringe of setae along distal margins. Interocular plate (
<figureCitation id="6287C8A2297EFF96FF03FE6AFEE8B660" box="[182,265,509,533]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6B</figureCitation>
) with pair of protrusions.
</paragraph>
<paragraph id="FA03D427297EFF96FFC4FDAAFE64B6C0" blockId="23.[112,753,573,693]" pageId="23" pageNumber="46">
Antennular peduncles (
<figureCitation id="6287C8A2297EFF96FED7FDAAFE52B620" box="[354,435,573,597]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6A</figureCitation>
) slender, when fully extended, ultimate segments reaching middle of corneas; ultimate and penultimate segments unarmed, basal segment distal margin with fringe of long setae.
</paragraph>
<paragraph id="FA03D427297EFF96FFC4FD4AFDCCB0C0" blockId="23.[112,753,733,1205]" pageId="23" pageNumber="46">
Antennal peduncles (
<figureCitation id="6287C8A2297EFF96FEDDFD4AFE0BB680" box="[360,490,733,757]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6A, C</figureCitation>
), when fully extended reaching 0.6 length of ocular peduncles; fifth segment unarmed, with 34 scattered setae on dorsal and ventral surfaces; fourth segment with tufts of setae on mesiodistal margin; third segment with tufts of long setae on ventral surface; second segment with dorsolateral distal angle produced, terminating in strong simple or bifid spine, lateral margin unarmed, dorsomesial distal angle unarmed or bearing 1 spine, mesial margin, lateral margin, and dorsolateral distal angle with dense tufts of setae; first segment with ventrolateral distal angle with 2 spines, ventromesial distal angle produced, rounded. Antennal acicle (
<figureCitation id="6287C8A2297EFF96FE2FFBAAFE10B020" box="[410,497,1085,1109]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="23" pageNumber="46">Fig. 6C</figureCitation>
) terminating in strong bifid spine; dorsomesial margin with 3 spines, dorsolateral margin unarmed or with single small spine. Antennal flagella bearing minute seta on each articulation.
</paragraph>
<paragraph id="FA03D427297EFF96FFC4FB4AFEDDB140" blockId="23.[112,752,1245,1333]" pageId="23" pageNumber="46">Third maxilliped with well developed crista dentata; basis with 1 or 2 small corneous spines, spines concealed by dense tufts of long setae.</paragraph>
<paragraph id="FA03D427297EFF96FFC4FACAFB93B560" blockId="23.[112,754,1373,2101]" lastBlockId="23.[813,1452,189,277]" pageId="23" pageNumber="46">
Chelipeds vastly unequal, left larger than right. Left cheliped (
<figureCitation id="6287C8A2297EFF96FFCCFAEAFF11B1E0" box="[121,240,1405,1430]" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7A, B</figureCitation>
) very stout, 1.7 times longer than wide. Outer face of palm, dactyl, and fixed finger (
<figureCitation id="6287C8A2297EFF96FD9BFA0AFD66B1C0" box="[558,647,1437,1461]" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7A</figureCitation>
) convex; bearing irregular rows of corneous-tipped conical spines; and covered with tufts of mixed short and long setae, setae not arising singly but rather in dense tufts of 3 or more. Dactyl and fixed finger each terminating in strong corneous claw, cutting edges each with 5 (dactyl) or 7 (fixed finger) low, molar-like calcareous teeth. Upper face of palm and dactyl with 2 or 3 irregular rows of corneous-tipped conical spines, spines larger and more hooked in upper proximal angle; lower margin bearing row of conical corneous tipped spines. Inner face of palm, dactyl, and fixed finger (
<figureCitation id="6287C8A2297EFF96FD71F94AFF70B360" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7B</figureCitation>
) slightly convex; fixed finger and disto-lateral corner of palm with 34 irregular rows of low corneous-tipped spines; inner face with scattered tufts of setae particularly on upper and lower margins. Carpus with upper margin bearing 4 prominent corneous-tipped spines; outer face slightly convex with irregular rows of corneous-tipped spines, lower outer margin with 4 small corneous-tipped spines; inner face with scattered tufts of setae particularly on inner distal margin. Merus with disto-dorsal and disto-ventral margins of lateral face each bearing 45 corneous-tipped spines; entire lateral face with tufts of long setae; ventromesial margin crested with 7 calcareous teeth, proximal larger. Ischium with 4 teeth on ventromesial margin.
</paragraph>
<paragraph id="FA03D427297EFF96FC98FEAAFBD5B000" blockId="23.[811,1453,317,1141]" pageId="23" pageNumber="46">
Right cheliped (
<figureCitation id="6287C8A2297EFF96FC63FEAAFBA9B520" box="[982,1096,317,341]" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7C, D</figureCitation>
) more slender than left, 1.7 times longer than wide. Outer face of palm, dactyl, and fixed finger (
<figureCitation id="6287C8A2297EFF96FC80FEEAFC69B5E0" box="[821,904,381,405]" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7C</figureCitation>
) bearing irregular rows of corneous-tipped spines, spines absent from lower proximal area of palm; covered with dense tufts of long setae, setae not arising singly, but rather in dense tufts of 3 or more. Dactyl terminating in strong corneous claw; cutting edge with 6 low molar-like calcareous teeth; upper margin with 3 irregular rows of strong corneous-tipped conical spines. Fixed finger terminating in strong corneous claw; cutting edge with 6 low molar-like calcareous teeth; outer, lower, and inner faces with irregular rows of low corneous-tipped spines. Inner face of palm, dactyl, and fixed finger (
<figureCitation id="6287C8A2297EFF96FBFCFD2AFB43B6A0" box="[1097,1186,701,725]" captionStart="Fig" captionStartId="24.[135,170,2064,2086]" captionTargetBox="[149,1461,189,2049]" captionTargetId="figure-8@24.[144,1464,186,2051]" captionTargetPageId="24" captionText="Fig. 7. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="23" pageNumber="46">Fig. 7D</figureCitation>
) slightly convex; fixed finger and disto-lateral corner of palm with low corneoustipped spines; inner face of palm with scattered tufts of long setae. Carpus with upper margin bearing 3 strong corneous-tipped conical spines; outer face with few strong spines; distal margin of outer face with 2 strong corneoustipped corneous spines near midline; outer face covered with tufts of long setae. Merus with distal margin bearing 3 corneous-tipped spines, two of which prominent and located at dorso-distal and ventro-distal angles; upper margin with 1 corneous-tipped spine sub-distally; outer face covered with tufts of long setae; ventromesial margin crested with 3 large calcareous teeth proximally. Ischium with 5 calcareous teeth on ventromesial margin.
</paragraph>
<paragraph id="FA03D427297EFF96FC98FB0AFA6AB3E0" blockId="23.[811,1454,1181,1941]" pageId="23" pageNumber="46">
Second pereopods (
<figureCitation id="6287C8A2297EFF96FBA3FB0AFB92B0C0" box="[1046,1139,1181,1205]" captionStart="Fig" captionStartId="25.[113,148,1993,2015]" captionTargetBox="[104,1451,262,1977]" captionTargetId="figure-14@25.[101,1458,248,1987]" captionTargetPageId="25" captionText="Fig. 8. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left second pereopod, lateral view; B, Propodus of left third pereopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503053" httpUri="https://zenodo.org/record/4503053/files/figure.png" pageId="23" pageNumber="46">Fig. 8A</figureCitation>
) and right third pereopod generally similar, but setation and armament slightly different between second pair and right third; second pair more slender than right third; of second pair, left slightly shorter than right. Dactyls all 1.31.4 length of propodi; each terminating in strong corneous claw; subcylindrical; ventral margins each with 510 small corneous spines distally; dactyls otherwise unarmed but densely covered with dense tufts of setae, setae especially dense on dorsal surfaces. Propodi 1.4 length of carpi; lateral faces convex, unarmed, and uniformly covered with dense tufts of setae (second) or setae absent from the lower 0.6 of lateral face (right third); mesial face flattened, smooth, unarmed, with sparse (second) or moderate (right third) tufts of setae. Carpi 0.70.8 length of meri; lateral faces flat with tufts of setae on dorsal and ventral margins; dorsal faces flat, covered with tufts of long setae, bearing one prominent corneous-tipped conical spine on the disto-mesial angle and 813 small to moderate corneous-tipped spines on distal 0.3. Meri lateral faces flat (second) or slightly convex (right third), with tufts of setae on dorsal and ventral margins; ventrolateral distal angle bearing 1 strong corneous-tipped spine (second) or unarmed (right third); ventral margins crested with row of tubercles bearing dense tufts of long setae and 02 calcareous teeth distally. Ischia unarmed.
</paragraph>
<paragraph id="FA03D427297EFF9BFC98F82AFD96BC6C" blockId="23.[813,1452,1981,2101]" lastBlockId="26.[135,776,1185,2073]" lastPageId="26" lastPageNumber="49" pageId="23" pageNumber="46">
Left third pereopod (
<figureCitation id="6287C8A2297EFF96FBADF82AFB91B3A0" box="[1048,1136,1981,2005]" captionStart="Fig" captionStartId="18.[136,171,1335,1357]" captionTargetBox="[139,777,188,1321]" captionTargetId="figure-125@18.[122,790,183,1324]" captionTargetPageId="18" captionText="Fig. 5. Lateral view of left third pereopod. A, Dardanus lagopodes (Forskål, 1775). Male SL 9.8 mm, stn N2, ZRC 2015.0489; B, Dardanus lagopodes (Forskål, 1775). Female SL 7.6 mm, stn R1, ZRC 2015.0488; C, Dardanus sanguinolentus (Quoy &amp; Gaimard, 1824). Male SL 7.8 mm, stn R38, ZRC 2015.0491; D, Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503046" httpUri="https://zenodo.org/record/4503046/files/figure.png" pageId="23" pageNumber="46">Fig. 5D</figureCitation>
) stout. Dactyl 1.4 length of propodus, terminating in large corneous claw; lateral face flattened and covered with tufts of short and long setae, setae especially dense along dorsal and ventral margins and not arising singly but rather in dense tufts of 3 or more; dorsal and ventral areas of lateral face with 1 (dorsal) or 23 (ventral) rows of sharp corneous spines on proximal 0.5; ventral margin with 5 small corneous spines distally; mesial face convex, with tufts of long setae along median line. Propodus (
<figureCitation id="6287C8A22973FF9BFE82FAD6FE6AB12C" box="[311,395,1345,1369]" captionStart="Fig" captionStartId="25.[113,148,1993,2015]" captionTargetBox="[104,1451,262,1977]" captionTargetId="figure-14@25.[101,1458,248,1987]" captionTargetPageId="25" captionText="Fig. 8. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left second pereopod, lateral view; B, Propodus of left third pereopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503053" httpUri="https://zenodo.org/record/4503053/files/figure.png" pageId="26" pageNumber="49">Fig. 8B</figureCitation>
) 1.4 length of carpus, very broad, 2.1 as long as broad; lateral face flattened, with nearly indistinct longitudinal lobe along midline (not visible in figure), covered with tufts of short and long setae, setae especially dense along dorsal and ventral margins; dorsal margin of lateral face with 2 irregular rows of sharp corneous spines; subventral 0.3 of lateral face with irregular rows of sharp corneous spines; distal margin of lateral face with 3 small corneous spines; mesial face convex, smooth, bearing 5 small corneous spines on ventral half of distal margin, with sparse tufts of long setae forming 3 longitudinal rows (2 rows on dorsal half, 1 row subventrally). Carpus 0.8 length of merus; lateral face flat; dorsal margin with row of corneous-tipped spines; dorsolateral angle with several strong corneous-tipped spines covered by dense tufts of long setae; dorsal, distal, and ventral margins covered by dense tufts of setae; mesial face slightly convex, smooth, unarmed, distal margin bearing tufts of long setae. Merus lateral face strongly convex, with tufts of long setae on the dorsal, ventral, and subdistal margins; mesial face flattened with tufts of setae on subdorsal margin; ventral margin crested with row of tubercles bearing dense tufts of long setae and 2 calcareous teeth distally. Ischium unarmed.
</paragraph>
<caption id="AEC384AF2971FF99FF32F787FB60BC37" ID-DOI="http://doi.org/10.5281/zenodo.4503051" ID-Zenodo-Dep="4503051" httpUri="https://zenodo.org/record/4503051/files/figure.png" pageId="24" pageNumber="47" startId="24.[135,170,2064,2086]" targetBox="[149,1461,189,2049]" targetPageId="24">
<paragraph id="FA03D4272971FF99FF32F787FB60BC37" blockId="24.[135,1474,2064,2115]" pageId="24" pageNumber="47">
Fig. 7.
<taxonomicName id="3DBCAFA42971FF99FF64F787FE79BC50" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[209,408,2064,2085]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="47" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C808352971FF99FF64F787FE79BC50" box="[209,408,2064,2085]" italics="true" pageId="24" pageNumber="47">Dardanus balhibuon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D3FBB54E2971FF99FE12F787FDFFBC53" box="[423,542,2064,2086]" pageId="24" pageNumber="47" rank="species">new species</taxonomicNameLabel>
, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left cheliped, outer view; B, Left cheliped, inner view; C, Right cheliped, outer view; D, Right cheliped, inner view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="AEC384AF2970FF98FFC4F85EFCF5B38E" ID-DOI="http://doi.org/10.5281/zenodo.4503053" ID-Zenodo-Dep="4503053" httpUri="https://zenodo.org/record/4503053/files/figure.png" pageId="25" pageNumber="48" startId="25.[113,148,1993,2015]" targetBox="[104,1451,262,1977]" targetPageId="25">
<paragraph id="FA03D4272970FF98FFC4F85EFCF5B38E" blockId="25.[113,1452,1993,2044]" pageId="25" pageNumber="48">
Fig. 8.
<taxonomicName id="3DBCAFA42970FF98FF0CF85EFE61B3AB" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[185,384,1993,2014]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C808352970FF98FF0CF85EFE61B3AB" box="[185,384,1993,2014]" italics="true" pageId="25" pageNumber="48">Dardanus balhibuon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D3FBB54E2970FF98FE38F85EFDE5B3AA" box="[397,516,1993,2015]" pageId="25" pageNumber="48" rank="species">new species</taxonomicNameLabel>
, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left second pereopod, lateral view; B, Propodus of left third pereopod, lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="AEC384AF2973FF9BFF3DFBA0FA25B01F" ID-DOI="http://doi.org/10.5281/zenodo.4503057" ID-Zenodo-Dep="4503057" httpUri="https://zenodo.org/record/4503057/files/figure.png" pageId="26" pageNumber="49" startId="26.[136,171,1079,1101]" targetBox="[134,1473,191,1070]" targetPageId="26">
<paragraph id="FA03D4272973FF9BFF3DFBA0FA25B01F" blockId="26.[136,1476,1079,1130]" pageId="26" pageNumber="49">Fig. 9. Maximum likelihood phylogram based on COI mitochondrial DNA sequences. Branch support for the major clades are indicated above (= likelihood bootstrap values) and below (= Bayesian posterior probabilities) the branches. Bootstrap values &lt;70% were not reported.</paragraph>
</caption>
<paragraph id="FA03D4272973FF9BFCF6FB36FC11B08C" blockId="26.[835,1474,1185,1273]" pageId="26" pageNumber="49">
Sternite of third pereopods (
<figureCitation id="6287C8A22973FF9BFB3CFB36FB02B0CC" box="[1161,1251,1185,1209]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="26" pageNumber="49">Fig. 6D</figureCitation>
) with anterior lobe rectangular, with 2 protrusions each bearing tuft of long setae anteriorly.
</paragraph>
<paragraph id="FA03D4272973FF9BFCF6FAB6FA20B1EC" blockId="26.[834,1474,1313,1433]" pageId="26" pageNumber="49">Fourth pereopods semichelate; dactyls each with 5 corneous ventral spines on lateral face; propodal rasps well developed; carpi each with sharp dorsodistal spine covered by dense tuft of long setae (spine missing/broken in right fourth pereopod).</paragraph>
<paragraph id="FA03D4272973FF9BFCF6FA56FC5BB18C" blockId="26.[835,1474,1473,1529]" pageId="26" pageNumber="49">Fifth pereopods chelate; rasps of dactyl and propodus well developed.</paragraph>
<paragraph id="FA03D4272973FF9BFCF6F9B6FA5EB2EC" blockId="26.[835,1474,1569,1689]" pageId="26" pageNumber="49">Female pleon with second to fifth left pleopods fringed with long setae; second to fourth triramous; fifth damaged; single, very large, elongate, triangular fleshy membrane fringed with long setae present between fourth and fifth pleopods.</paragraph>
<paragraph id="FA03D4272973FF9BFCF6F956FA87B28C" blockId="26.[835,1473,1729,1785]" pageId="26" pageNumber="49">Uropods strongly asymmetrical, left larger than right; endopods and exopods with well developed rasps.</paragraph>
<paragraph id="FA03D4272973FF9BFCF6F8B6FBACB3AC" blockId="26.[835,1475,1825,2009]" pageId="26" pageNumber="49">
Telson (
<figureCitation id="6287C8A22973FF9BFC2EF8B6FBE8B34C" box="[923,1033,1825,1849]" captionStart="Fig" captionStartId="22.[135,170,2022,2044]" captionTargetBox="[134,1471,189,2016]" captionTargetId="figure-8@22.[122,1489,183,2025]" captionTargetPageId="22" captionText="Fig. 6. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Shield and cephalic appendages; B, Interocular plate and right ocular acicle; C, Right antennal peduncle; D, Anterior lobe of sternite of third pereopods; E, Telson, dorsal view; F, Posterior lobes of telson, ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503049" httpUri="https://zenodo.org/record/4503049/files/figure.png" pageId="26" pageNumber="49">Fig. 6E, F</figureCitation>
) with lateral constrictions; marginal area partially calcified; posterior lobes separated by median cleft, left much larger than right, each with 7 (left) or 6 (right) ventral corneous spines near terminal margin, terminal margins fringed with long setae; anterior lobes with long setae on lateral margins.
</paragraph>
<paragraph id="FA03D4272973FF9AFCF6F796FDD1B2F5" blockId="26.[835,1474,2049,2105]" lastBlockId="27.[112,753,1384,1664]" lastPageId="27" lastPageNumber="50" pageId="26" pageNumber="49">
<emphasis id="C8C808352973FF9BFCF6F796FC7DBC6C" bold="true" box="[835,924,2049,2073]" pageId="26" pageNumber="49">Colour.</emphasis>
In life unknown, but the specimen was initially mistakenly identified as
<taxonomicName id="3DBCAFA42973FF9BFBEFF7B5FB0EBC4C" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[1114,1263,2081,2105]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="49" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352973FF9BFBEFF7B5FB0EBC4C" box="[1114,1263,2081,2105]" italics="true" pageId="26" pageNumber="49">D. lagopodes</emphasis>
</taxonomicName>
and therefore live colouration must resemble
<taxonomicName id="3DBCAFA42972FF9AFE3DFAFEFDF2B1F5" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[392,531,1384,1408]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFE3DFAFEFDF2B1F5" box="[392,531,1384,1408]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
or the closely related
<taxonomicName id="3DBCAFA42972FF9AFFC5FA1EFED3B1D5" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[112,306,1416,1440]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFFC5FA1EFED3B1D5" box="[112,306,1416,1440]" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
</taxonomicName>
. Colour in preservative (
<figureCitation id="6287C8A22972FF9AFDF4FA1FFD77B1D5" box="[577,662,1416,1440]" captionStart="Fig" captionStartId="17.[113,148,1651,1673]" captionTargetBox="[113,1451,189,1636]" captionTargetId="figure-90@17.[113,1451,189,1636]" captionTargetPageId="17" captionText="Fig. 4. A, Dardanus sanguinolentus (Quoy &amp; Gaimard, 1824). Female ovig. SL 6.3 mm, stn R6; B, Dardanus balhibuon, n. sp. Holotype female SL 14.3 mm, stn R38, NMCR 40109, preserved colouration; C, Dardanus megistos (Herbst, 1804). Male SL 11.0 mm, stn M8; D, Dardanus pedunculatus (Herbst, 1804). Male SL 13.4 mm, stn N3; E, Dardanus scutellatus. Male SL 9.8 mm, stn D12; F, Dardanus woodmasoni (Alcock, 1905). Male SL 5.8 mm, stn R76." figureDoi="http://doi.org/10.5281/zenodo.4503040" httpUri="https://zenodo.org/record/4503040/files/figure.png" pageId="27" pageNumber="50">Fig. 4B</figureCitation>
): shield, ocular acicles, and antennae orange with white spots; ocular peduncles pale brown; antennules and antennal flagellae pale yellow; chelipeds and ambulatory legs orange with white spots and mottlings overall; ambulatory legs with a wide deep orange-red ring on the subdistal dorsal face of the propodus; chelipeds and ambulatory legs with bright orange patches on outer faces of carpi and meri.
</paragraph>
<caption id="AEC384AF2972FF9AFFC4FF2AFB83B4A6" ID-Table-UUID="AEC384AF2972FF9AFFC4FF2AFB83B4A6" box="[113,1122,189,211]" httpUri="http://table.plazi.org/id/AEC384AF2972FF9AFFC4FF2AFB83B4A6" pageId="27" pageNumber="50" startId="27.[113,168,189,211]" targetBox="[128,1436,264,1305]" targetIsTable="true" targetPageId="27">
<paragraph id="FA03D4272972FF9AFFC4FF2AFB83B4A6" blockId="27.[113,1122,189,211]" box="[113,1122,189,211]" pageId="27" pageNumber="50">
Table 2. Morphological characters separating members of the “
<taxonomicName id="3DBCAFA42972FF9AFD6BFF2AFC48B4A7" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[734,937,189,210]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFD6BFF2AFC48B4A7" box="[734,937,189,210]" italics="true" pageId="27" pageNumber="50">Dardanus lagopodes</emphasis>
</taxonomicName>
” species complex.
</paragraph>
</caption>
<paragraph id="FA03D4272972FF9AFF1BFE81FAA6B16C" pageId="27" pageNumber="50">
<table id="88BC26872972007EFF35FE9FFA7DB16C" box="[128,1436,264,1305]" gridcols="4" gridrows="9" pageId="27" pageNumber="50">
<tr id="448CD6652972007EFF35FE9FFA7DB54E" box="[128,1436,264,315]" gridrow="0" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FE9FFEB2B54E" box="[128,339,264,315]" gridcol="0" gridrow="0" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFF1BFE81FECCB55E" bold="true" box="[174,301,278,299]" pageId="27" pageNumber="50">Morphology</emphasis>
</th>
<th id="075DBF192972007EFECEFE9FFD26B54E" box="[379,711,264,315]" gridcol="1" gridrow="0" pageId="27" pageNumber="50">
<taxonomicName id="3DBCAFA42972FF9AFE0CFE9FFD8FB54F" authority="(Forskal, 1775)" baseAuthorityName="Forskal" baseAuthorityYear="1775" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFE0CFE9FFD8FB54F" bold="true" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFE0CFE9FFD67B56B" bold="true" box="[441,646,264,286]" italics="true" pageId="27" pageNumber="50">Dardanus lagopodes</emphasis>
(
<bibRefCitation id="9E2DA9D62972FF9AFE6DFEB3FD86B54F" author="Forskal P" box="[472,615,292,314]" pageId="27" pageNumber="50" pagination="1 - 164" refId="ref42874" refString="Forskal P (1775) Descriptiones Animalium, Avium, Amphibiorum, Piscium, Insectorum, vermium; quae in itinere oriental observ avit. Post mortem auctoris edidit Carsten Niebuhr. Hauniae, pp. 1 - 19, 1 - 164." type="journal article" year="1775">Forskål, 1775</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="075DBF192972007EFD53FE9FFBD3B54E" box="[742,1074,264,315]" gridcol="2" gridrow="0" pageId="27" pageNumber="50">
<taxonomicName id="3DBCAFA42972FF9AFCBCFE9FFBF1B54F" authority="(Quoy &amp; Gaimard, 1824)" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFCBCFE9FFBF1B54F" bold="true" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFCBCFE9FFBEDB56B" bold="true" box="[777,1036,264,286]" italics="true" pageId="27" pageNumber="50">Dardanus sanguinolentus</emphasis>
(
<bibRefCitation id="9E2DA9D62972FF9AFCBAFEB3FBE8B54F" author="Quoy JRC &amp; Gaimard JP" box="[783,1033,292,315]" pageId="27" pageNumber="50" refId="ref47029" refString="Quoy JRC &amp; Gaimard JP (1824) Zoologie. In: Freycinet LCD de (ed.) Voyage autour du monde sur les corvettes de S. M. l'Uranie et la Physicienne, pendant les annees 1817, 1818, 1819 et 1820. Paris, 1. Pp. 517 - 536." type="journal volume" year="1824">Quoy &amp; Gaimard, 1824</bibRefCitation>
)
</emphasis>
</taxonomicName>
</th>
<th id="075DBF192972007EFBFAFE9FFA7DB54E" box="[1103,1436,264,315]" gridcol="3" gridrow="0" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFB3EFE9FFAD3B54E" bold="true" pageId="27" pageNumber="50">
<taxonomicName id="3DBCAFA42972FF9AFB3EFE9FFABBB56B" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[1163,1370,264,286]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C808352972FF9AFB3EFE9FFABBB56B" bold="true" box="[1163,1370,264,286]" italics="true" pageId="27" pageNumber="50">Dardanus balhibuon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D3FBB54E2972FF9AFB0FFEB2FAD3B54E" box="[1210,1330,293,315]" pageId="27" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="448CD6652972007EFF35FEC9FA7DB5D8" box="[128,1436,350,429]" gridrow="1" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FEC9FEB2B5D8" box="[128,339,350,429]" gridcol="0" gridrow="1" pageId="27" pageNumber="50">Shield</th>
<td id="075DBF192972007EFECEFEC9FD26B5D8" box="[379,711,350,429]" gridcol="1" gridrow="1" pageId="27" pageNumber="50">0.91 times as long as broad, lateral margin with tufts of setae</td>
<td id="075DBF192972007EFD53FEC9FBD3B5D8" box="[742,1074,350,429]" gridcol="2" gridrow="1" pageId="27" pageNumber="50">0.91 times as long as broad, lateral margin with tufts of setae</td>
<td id="075DBF192972007EFBFAFEC9FA7DB5D8" box="[1103,1436,350,429]" gridcol="3" gridrow="1" pageId="27" pageNumber="50">0.9 times as long as broad, lateral margin with dense tufts of long setae</td>
</tr>
<tr id="448CD6652972007EFF35FE50FA7DB663" box="[128,1436,455,534]" gridrow="2" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FE50FEB2B663" box="[128,339,455,534]" gridcol="0" gridrow="2" pageId="27" pageNumber="50">Ocular peduncles</th>
<td id="075DBF192972007EFECEFE50FD26B663" box="[379,711,455,534]" gridcol="1" gridrow="2" pageId="27" pageNumber="50">0.9 times length of shield, corneas 0.2 length of ocular peduncles</td>
<td id="075DBF192972007EFD53FE50FBD3B663" box="[742,1074,455,534]" gridcol="2" gridrow="2" pageId="27" pageNumber="50">0.9 times length of shield, corneas 0.2 length of ocular peduncles</td>
<td id="075DBF192972007EFBFAFE50FA7DB663" box="[1103,1436,455,534]" gridcol="3" gridrow="2" pageId="27" pageNumber="50">0.8 times length of shield, corneas 0.3 length of ocular peduncles</td>
</tr>
<tr id="448CD6652972007EFF35FDB8FA7DB630" box="[128,1436,559,581]" gridrow="3" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FDB8FEB2B630" box="[128,339,559,581]" gridcol="0" gridrow="3" pageId="27" pageNumber="50">Antennular peduncles</th>
<td id="075DBF192972007EFECEFDB8FD26B630" box="[379,711,559,581]" gridcol="1" gridrow="3" pageId="27" pageNumber="50">Not reaching base of corneas</td>
<td id="075DBF192972007EFD53FDB8FBD3B630" box="[742,1074,559,581]" gridcol="2" gridrow="3" pageId="27" pageNumber="50">Reaching base of corneas</td>
<td id="075DBF192972007EFBFAFDB8FA7DB630" box="[1103,1436,559,581]" gridcol="3" gridrow="3" pageId="27" pageNumber="50">Reaching middle of corneas</td>
</tr>
<tr id="448CD6652972007EFF35FDC9FA7DB6E4" box="[128,1436,606,657]" gridrow="4" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FDC9FEB2B6E4" box="[128,339,606,657]" gridcol="0" gridrow="4" pageId="27" pageNumber="50">Antennal peduncles</th>
<td id="075DBF192972007EFECEFDC9FD26B6E4" box="[379,711,606,657]" gridcol="1" gridrow="4" pageId="27" pageNumber="50">0.7 times length of ocular peduncles</td>
<td id="075DBF192972007EFD53FDC9FBD3B6E4" box="[742,1074,606,657]" gridcol="2" gridrow="4" pageId="27" pageNumber="50">0.6 times length of ocular peduncles</td>
<td id="075DBF192972007EFBFAFDC9FA7DB6E4" box="[1103,1436,606,657]" gridcol="3" gridrow="4" pageId="27" pageNumber="50">0.6 times length of ocular peduncles</td>
</tr>
<tr id="448CD6652972007EFF35FD3DFA7DB68C" box="[128,1436,682,761]" gridrow="5" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FD3DFEB2B68C" box="[128,339,682,761]" gridcol="0" gridrow="5" pageId="27" pageNumber="50">Left second pereopod</th>
<td id="075DBF192972007EFECEFD3DFD26B68C" box="[379,711,682,761]" gridcol="1" gridrow="5" pageId="27" pageNumber="50">Dactyl 1.31.4 length of propodus, ventral margin with 57 spines distally</td>
<td id="075DBF192972007EFD53FD3DFBD3B68C" box="[742,1074,682,761]" gridcol="2" gridrow="5" pageId="27" pageNumber="50">Dactyl 1.61.7 length of propodus, ventral margin with 7 spines distally</td>
<td id="075DBF192972007EFBFAFD3DFA7DB68C" box="[1103,1436,682,761]" gridcol="3" gridrow="5" pageId="27" pageNumber="50">Dactyl 1.31.4 length of propodus, ventral margin with 6 spines distally</td>
</tr>
<tr id="448CD6652972007EFF35FC85FA7DB7CD" box="[128,1436,786,952]" gridrow="6" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FC85FEB2B7CD" box="[128,339,786,952]" gridcol="0" gridrow="6" pageId="27" pageNumber="50">Left third pereopod</th>
<td id="075DBF192972007EFECEFC85FD26B7CD" box="[379,711,786,952]" gridcol="1" gridrow="6" pageId="27" pageNumber="50">Dactyl 1.31.4 length of propodus, ventral margin with 3 spines; propodus 2.32.4 as long as broad, no longitudinal sulcus on dactylus and propodus</td>
<td id="075DBF192972007EFD53FC85FBD3B7CD" box="[742,1074,786,952]" gridcol="2" gridrow="6" pageId="27" pageNumber="50">Dactyl 1.2 length of propodus, ventral margin with 3 spines; propodus 2.32.5 as long as broad, longitudinal sulcus on lateral face of dactylus and propodus</td>
<td id="075DBF192972007EFBFAFC85FA7DB7CD" box="[1103,1436,786,952]" gridcol="3" gridrow="6" pageId="27" pageNumber="50">Dactyl 1.2 length of propodus, ventral margin with 5 spines; propodus 2.1 as long as broad, indistinct midline longitudinal lobe on lateral face of propodus</td>
</tr>
<tr id="448CD6652972007EFF35FC46FA7DB02F" box="[128,1436,977,1114]" gridrow="7" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FC46FEB2B02F" box="[128,339,977,1114]" gridcol="0" gridrow="7" pageId="27" pageNumber="50">Telson</th>
<td id="075DBF192972007EFECEFC46FD26B02F" box="[379,711,977,1114]" gridcol="1" gridrow="7" pageId="27" pageNumber="50">Asymmetrical, posterior lobe separated by median cleft, each with 7 corneous spines near terminal margin</td>
<td id="075DBF192972007EFD53FC46FBD3B02F" box="[742,1074,977,1114]" gridcol="2" gridrow="7" pageId="27" pageNumber="50">Asymmetrical, posterior lobe separated by median cleft, left lobe with 7, right lobe with 4 corneous spines near terminal margin</td>
<td id="075DBF192972007EFBFAFC46FA7DB02F" box="[1103,1436,977,1114]" gridcol="3" gridrow="7" pageId="27" pageNumber="50">Asymmetrical, posterior lobe separated by median cleft, left lobe with 7, right lobe with 6 corneous spines near terminal margin</td>
</tr>
<tr id="448CD6652972007EFF35FBE4FA7DB16C" box="[128,1436,1139,1305]" gridrow="8" pageId="27" pageNumber="50">
<th id="075DBF192972007EFF35FBE4FEB2B16C" box="[128,339,1139,1305]" gridcol="0" gridrow="8" pageId="27" pageNumber="50">Setosity</th>
<td id="075DBF192972007EFECEFBE4FD26B16C" box="[379,711,1139,1305]" gridcol="1" gridrow="8" pageId="27" pageNumber="50">Long setae covering surface of pereopods and chelae, shield with sparse tufts of long setae; setae arise singly</td>
<td id="075DBF192972007EFD53FBE4FBD3B16C" box="[742,1074,1139,1305]" gridcol="2" gridrow="8" pageId="27" pageNumber="50">Long setae covering surface of pereopods and chelae, shield with sparse tufts of long setae; setae arise singly</td>
<td id="075DBF192972007EFBFAFBE4FA7DB16C" box="[1103,1436,1139,1305]" gridcol="3" gridrow="8" pageId="27" pageNumber="50">Very long, extremely dense setae nearly obscuring surface of pereopods and chelae, shield bearing dense tufts of very long setae; multiple (3+) setae arise from single spot</td>
</tr>
</table>
</paragraph>
<paragraph id="FA03D4272972FF9AFFC4F93FFD92B2B5" blockId="27.[113,627,1704,1728]" box="[113,627,1704,1728]" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFFC4F93FFF31B2B5" bold="true" box="[113,208,1704,1728]" pageId="27" pageNumber="50">Habitat.</emphasis>
Subtidal,
<quantity id="3D4479C22972FF9AFEF5F93FFE72B2CA" box="[320,403,1704,1728]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.7" metricValueMin="0.6" pageId="27" pageNumber="50" unit="m" value="21.5" valueMax="37.0" valueMin="6.0">637 m</quantity>
deep, on reef slope.
</paragraph>
</subSubSection>
<subSubSection id="B2A687AC2972FF9AFFC4F97FFEECB335" pageId="27" pageNumber="50" type="etymology">
<paragraph id="FA03D4272972FF9AFFC4F97FFEECB335" blockId="27.[112,752,1768,1856]" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFFC4F97FFF10B375" bold="true" box="[113,241,1768,1792]" pageId="27" pageNumber="50">Etymology.</emphasis>
From the Boholano/Cebuano word, “balhibuon”, meaning hairy, referring to the exceptionally setose condition of the species.
</paragraph>
</subSubSection>
<subSubSection id="B2A687AC2972FF9AFFC4F8FFFA4CB395" pageId="27" pageNumber="50" type="discussion">
<paragraph id="FA03D4272972FF9AFFC4F8FFFA4CB395" blockId="27.[112,753,1896,2048]" lastBlockId="27.[811,1453,1384,2016]" pageId="27" pageNumber="50">
<emphasis id="C8C808352972FF9AFFC4F8FFFF09B3F5" bold="true" box="[113,232,1896,1920]" pageId="27" pageNumber="50">Remarks.</emphasis>
A single specimen was found that closely resembles
<taxonomicName id="3DBCAFA42972FF9AFF6BF81EFE89B3D5" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[222,360,1928,1952]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFF6BF81EFE89B3D5" box="[222,360,1928,1952]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
and
<taxonomicName id="3DBCAFA42972FF9AFE20F81EFDB0B3D5" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[405,593,1928,1952]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFE20F81EFDB0B3D5" box="[405,593,1928,1952]" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
</taxonomicName>
in morphology and overall colouration after preservation in alcohol, and yet differs in some important respects: this new species is larger (
<quantity id="3D4479C22972FF9AFF6BF87FFEA3B38A" box="[222,322,2024,2048]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="27" pageNumber="50" unit="mm" value="14.3">14.3 mm</quantity>
SL) than typical
<taxonomicName id="3DBCAFA42972FF9AFDBCF87EFD7ABC75" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[521,667,2024,2048]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFDBCF87EFD7ABC75" box="[521,667,2024,2048]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
and
<taxonomicName id="3DBCAFA42972FF9AFD63F87EFC2FB1F5" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[726,974,1384,2048]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFD63F87EFC2FB1F5" box="[726,974,1384,2048]" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
</taxonomicName>
, and is exceedingly setose, with numerous (3 or more) long setae arising in tufts from the same spot. The excessively setose condition of the entire body is one of the most distinct features of the new species, a condition which is not mentioned in the original descriptions nor observed in the
<typeStatus id="25076A852972FF9AFCE0F99FFC62B255" box="[853,899,1544,1568]" pageId="27" pageNumber="50">type</typeStatus>
specimens of
<taxonomicName id="3DBCAFA42972FF9AFB90F99EFB52B255" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[1061,1203,1544,1568]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFB90F99EFB52B255" box="[1061,1203,1544,1568]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
,
<taxonomicName id="3DBCAFA42972FF9AFB77F99EFA64B255" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[1218,1413,1544,1568]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFB77F99EFA64B255" box="[1218,1413,1544,1568]" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
</taxonomicName>
, or any of their junior synonyms. Another easy way to see the differences between these three species is from the shape of left third pereopod. In
<taxonomicName id="3DBCAFA42972FF9AFBFEF9FEFB3CB2F5" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[1099,1245,1640,1664]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFBFEF9FEFB3CB2F5" box="[1099,1245,1640,1664]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
the lateral face of dactyl and propodus of the left third pereopod is convex or slightly flattened, in
<taxonomicName id="3DBCAFA42972FF9AFBB3F93EFB26B2B5" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[1030,1223,1704,1728]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFBB3F93EFB26B2B5" box="[1030,1223,1704,1728]" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
</taxonomicName>
those lateral surfaces are flattened with a longitudinal concavity medially, while in
<taxonomicName id="3DBCAFA42972FF9AFCF8F97EFC03B375" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[845,994,1768,1792]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C808352972FF9AFCF8F97EFC03B375" box="[845,994,1768,1792]" italics="true" pageId="27" pageNumber="50">D. balhibuon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3FBB54E2972FF9AFC58F97FFBCDB28A" box="[1005,1068,1768,1791]" pageId="27" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
they are flattened with indistinct longitudinal lobe along the midline (
<tableCitation id="B73EE19C2972FF9AFB77F89FFAFBB355" box="[1218,1306,1800,1824]" captionStart="Table 2" captionStartId="27.[113,168,189,211]" captionTargetPageId="27" captionText="Table 2. Morphological characters separating members of the “Dardanus lagopodes” species complex." httpUri="http://table.plazi.org/id/AEC384AF2972FF9AFFC4FF2AFB83B4A6" pageId="27" pageNumber="50" tableUuid="AEC384AF2972FF9AFFC4FF2AFB83B4A6">Table 2</tableCitation>
,
<figureCitation id="6287C8A22972FF9AFA9CF89FFA97B355" box="[1321,1398,1800,1824]" captionStart="Fig" captionStartId="18.[136,171,1335,1357]" captionTargetBox="[139,777,188,1321]" captionTargetId="figure-125@18.[122,790,183,1324]" captionTargetPageId="18" captionText="Fig. 5. Lateral view of left third pereopod. A, Dardanus lagopodes (Forskål, 1775). Male SL 9.8 mm, stn N2, ZRC 2015.0489; B, Dardanus lagopodes (Forskål, 1775). Female SL 7.6 mm, stn R1, ZRC 2015.0488; C, Dardanus sanguinolentus (Quoy &amp; Gaimard, 1824). Male SL 7.8 mm, stn R38, ZRC 2015.0491; D, Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503046" httpUri="https://zenodo.org/record/4503046/files/figure.png" pageId="27" pageNumber="50">Figs. 5</figureCitation>
,
<figureCitation id="6287C8A22972FF9AFA30F89FFA46B355" box="[1413,1447,1800,1824]" captionStart="Fig" captionStartId="25.[113,148,1993,2015]" captionTargetBox="[104,1451,262,1977]" captionTargetId="figure-14@25.[101,1458,248,1987]" captionTargetPageId="25" captionText="Fig. 8. Dardanus balhibuon, new species, holotype. Female SL 14.3 mm, stn R38, NMCR 40109. A, Left second pereopod, lateral view; B, Propodus of left third pereopod, lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4503053" httpUri="https://zenodo.org/record/4503053/files/figure.png" pageId="27" pageNumber="50">8B</figureCitation>
; indistinct longitudinal lobe in
<taxonomicName id="3DBCAFA42972FF9AFBCBF8BEFAF1B335" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[1150,1296,1832,1856]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="balhibuon">
<emphasis id="C8C808352972FF9AFBCBF8BEFAF1B335" box="[1150,1296,1832,1856]" italics="true" pageId="27" pageNumber="50">D. balhibuon</emphasis>
</taxonomicName>
not shown in line figures). While it is allied to
<taxonomicName id="3DBCAFA42972FF9AFB09F8DEFAB2B315" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[1212,1363,1864,1888]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFB09F8DEFAB2B315" box="[1212,1363,1864,1888]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
and
<taxonomicName id="3DBCAFA42972FF9AFA24F8DEFBEBB3F5" authority=", COI" authorityName="COI" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="sanguinolentus">
<emphasis id="C8C808352972FF9AFA24F8DEFC2FB3F5" italics="true" pageId="27" pageNumber="50">D. sanguinolentus</emphasis>
, COI
</taxonomicName>
sequence data show that
<taxonomicName id="3DBCAFA42972FF9AFAABF8FEFA4AB3F5" authority="Celia &amp; Machel &amp; Malay &amp; Chan, 2018" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[1310,1451,1896,1920]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="balhibuon" status="sp. nov.">
<emphasis id="C8C808352972FF9AFAABF8FEFA4AB3F5" box="[1310,1451,1896,1920]" italics="true" pageId="27" pageNumber="50">D. balhibuon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3FBB54E2972FF9AFC98F81FFC8BB3EA" box="[813,874,1928,1951]" pageId="27" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
is genetically very distinct from other members of the
<taxonomicName id="3DBCAFA42972FF9AFCE0F83EFC07B3B5" baseAuthorityName="Forskal" baseAuthorityYear="1775" box="[853,998,1960,1984]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="lagopodes">
<emphasis id="C8C808352972FF9AFCE0F83EFC07B3B5" box="[853,998,1960,1984]" italics="true" pageId="27" pageNumber="50">D. lagopodes</emphasis>
</taxonomicName>
complex (
<figureCitation id="6287C8A22972FF9AFBEFF83FFB7CB3B5" box="[1114,1181,1960,1984]" captionStart="Fig" captionStartId="26.[136,171,1079,1101]" captionTargetBox="[134,1473,191,1070]" captionTargetId="figure-453@26.[124,1491,167,1083]" captionTargetPageId="26" captionText="Fig. 9. Maximum likelihood phylogram based on COI mitochondrial DNA sequences. Branch support for the major clades are indicated above (= likelihood bootstrap values) and below (= Bayesian posterior probabilities) the branches. Bootstrap values &lt;70% were not reported." figureDoi="http://doi.org/10.5281/zenodo.4503057" httpUri="https://zenodo.org/record/4503057/files/figure.png" pageId="27" pageNumber="50">Fig. 9</figureCitation>
).
<taxonomicName id="3DBCAFA42972FF9AFB01F83FFA72B3B5" authorityName="Celia &amp; Machel &amp; Malay &amp; Chan" authorityYear="2018" box="[1204,1427,1960,1984]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="50" phylum="Arthropoda" rank="species" species="balhibuon">
<emphasis id="C8C808352972FF9AFB01F83FFA72B3B5" box="[1204,1427,1960,1984]" italics="true" pageId="27" pageNumber="50">Dardanus balhibuon</emphasis>
</taxonomicName>
is presently only known from Pamilacan Island, the
<collectingCountry id="82AB94B72972FF9AFA85F85FFA4BB395" box="[1328,1450,1992,2016]" name="Philippines" pageId="27" pageNumber="50">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>