460 lines
63 KiB
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460 lines
63 KiB
XML
<document id="0BD6C5FC5891E15B83B931A1F68C1440" ID-DOI="10.5281/zenodo.4665088" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665088" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617711800044" checkinUser="felipe" docAuthor="Valiukevičius, Juozas" docDate="2003" docId="586B87E1FF88FF89FEC2F0E46C6DC1FB" docLanguage="en" docName="Geodiversitas.25.1.131-204.pdf" docOrigin="Geodiversitas 25 (1)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Nostolepis watsoni Valiukevičius 2003, n. sp." docType="treatment" docVersion="5" lastPageNumber="162" masterDocId="A452FF99FF94FF96FFDBF1036E5CC578" masterDocTitle="Devonian acanthodians from Severnaya Zemlya Archipelago (Russia)" masterLastPageNumber="204" masterPageNumber="131" pageNumber="159" updateTime="1698900742125" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="ACBEF5B840F78AAE50D8B884C3B353CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C4722C1B29CF1CD1B13251B1166B15C4">
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<mods:title id="251D64D77538C8F57FE05E0A981CC818">Devonian acanthodians from Severnaya Zemlya Archipelago (Russia)</mods:title>
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</mods:titleInfo>
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<mods:name id="2139CBDDA7C0A3A118F7E4546E164A73" type="personal">
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<mods:role id="19C1A2759636FB00D797A94D38A1E6ED">
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<mods:roleTerm id="006983DB180FA0148BD7DA54CE3B2CF9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1677E81362ED145EEFCA138D4749E71E">Valiukevičius, Juozas</mods:namePart>
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<mods:affiliation id="F7EA19FC6D346DD01DFE04957BAA0887">Institute of Geology and Geography, T. Ševčenkos 13, LT 2600 Vilnius (Lithuania) valiuk @ geologin. lt Acanthodians from the stratigraphic interval of late Pridoli (Late Silurian) to Givetian (Middle Devonian) are studied. Based on 48 acanthodian taxa, biostratigraphic zones are established within the late Pridoli (Poracanthodes punctatus Zone, Krasnaya Bukhta Formation) and Lochkovian (Nostolepis minima Zone), the latter divided into two subzones: Poracanthodes menneri (Severnaya Zemlya Formation) and Diplacanthus poltnigi n. sp. (Pod ” emnaya Formation). Two more biostratigraphic units are characterised by index species, but they have no acanthodian-based boundaries (separated by intervals without effective acanthodian characteristics). These are beds with Watsonacanthus costatus n. sp. typical for the Emsian Rusanov and lower Al’banov formations, and Eifelian? or Eifelian / Givetian? beds with Diplacanthus solidus n. sp. (Vstrechnaya Formation). Five new genera and 14 new species are described. They are based on articulated specimens from the topmost Severnaya Zemlya Formation (Acritolepis ushakovi n. gen., n. sp., A. urvantsevi n. gen., n. sp., Acanthacanthus ornatus n. gen., n. sp., Nostolepis decora n. sp., N. fragilis n. sp., Acanthopora transitans n. gen., n. sp. and Acanthospina irregulare n. gen., n. sp.) or on isolated scales identified from different formations (Nostolepis timanica n. sp., N. watsoni n. sp., Watsonacanthus costatus n. sp., Diplacanthus poltnigi n. sp., D. solidus n. sp., Gomphonchus nordicus n. sp. and Arcticacanthus bicostatus n. gen., n. sp.).</mods:affiliation>
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<mods:nameIdentifier id="7216B3E5D6B26B1C2F1F793808F35B85" type="email">valiuk@geologin.lt</mods:nameIdentifier>
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<mods:title id="6BF99ECF249C9736110A438459BAC487">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="EA58AE1C4A921D75CEC3F34EBB99D690">
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<mods:date id="7FAD87A13BC3624F0D7A74ACB4B0D040">2003</mods:date>
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<mods:detail id="E56577C853497E48D211F3CF6C1D0DCE" type="volume">
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<mods:number id="0AF12E94E7DE7CC966313A25C2FDF425">25</mods:number>
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</mods:detail>
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<mods:detail id="F412908F4963C20F0561584C51514352" type="issue">
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<mods:number id="9FAD6E99C04A6BE31CAE4594DF7DA7A0">1</mods:number>
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<mods:classification id="A133138E611CACE9824F19B01E1D4F1A">journal article</mods:classification>
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<mods:identifier id="12953F0F892DC979C8DB60A11E4164CF" type="DOI">10.5281/zenodo.4665088</mods:identifier>
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<mods:identifier id="BF8A116555B1139B2061B06D1EE9D209" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="1760E1F2DE223F79F7BFF26FEBA23D24" type="Zenodo-Dep">4665088</mods:identifier>
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<treatment id="586B87E1FF88FF89FEC2F0E46C6DC1FB" LSID="urn:lsid:plazi:treatment:586B87E1FF88FF89FEC2F0E46C6DC1FB" httpUri="http://treatment.plazi.org/id/586B87E1FF88FF89FEC2F0E46C6DC1FB" lastPageId="31" lastPageNumber="162" pageId="28" pageNumber="159">
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<subSubSection id="98D8657CFF88FF8AFEC2F0E46FA0C766" pageId="28" pageNumber="159" type="nomenclature">
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<paragraph id="D07D36F7FF88FF8AFEC2F0E46FA0C766" blockId="28.[281,554,487,542]" pageId="28" pageNumber="159">
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<taxonomicName id="17C24D74FF88FF8AFEC2F0E46FB5C487" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[281,489,487,511]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
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<emphasis id="E2B6EAE5FF88FF8AFEC2F0E46FB5C487" bold="true" box="[281,489,487,511]" italics="true" pageId="28" pageNumber="159">Nostolepis watsoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F985579EFF88FF8AFE2BF0EB6C76C487" box="[496,554,488,511]" pageId="28" pageNumber="159" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="48F92A72FF88FF8AFE95F3056FC3C766" box="[334,415,518,542]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Figs 23</figureCitation>
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A-F; 24)
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</paragraph>
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</subSubSection>
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<subSubSection id="98D8657CFF88FF8AFF4CF3436F40C79B" pageId="28" pageNumber="159" type="materials_examined">
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<paragraph id="D07D36F7FF88FF8AFF4CF3436F3FC7F0" blockId="28.[151,685,575,1281]" pageId="28" pageNumber="159">
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<materialsCitation id="60AA3CAAFF88FF8AFF4CF3436F3FC7F0" county="Member" location="Scale" municipality="Lower Devonian" pageId="28" pageNumber="159" specimenCount="1" typeStatus="holotype">
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<typeStatus id="0F798855FF88FF8AFF4CF3436F59C72C" box="[151,261,576,597]" pageId="28" pageNumber="159" type="holotype">HOLOTYPE</typeStatus>
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. — LIG 35-2020.
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<location id="D51D602CFF88FF8AFE62F33C6FB6C72D" LSID="urn:lsid:plazi:treatment:586B87E1FF88FF89FEC2F0E46C6DC1FB:D51D602CFF88FF8AFE62F33C6FB6C72D" box="[441,490,575,597]" county="Member" municipality="Lower Devonian" name="Scale" pageId="28" pageNumber="159">Scale</location>
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(
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<figureCitation id="48F92A72FF88FF8AFE2CF33C6C13C72D" box="[503,591,575,597]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Fig. 23A</figureCitation>
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).
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<location id="D51D602CFF88FF8AFD84F33C6E84C717" LSID="urn:lsid:plazi:treatment:586B87E1FF88FF89FEC2F0E46C6DC1FB:D51D602CFF88FF8AFD84F33C6E84C717" county="Member" municipality="Lower Devonian" name="Pioneer Island" pageId="28" pageNumber="159">Pioneer Island</location>
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, sample P-10-1.
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<collectingMunicipality id="3019AC8DFF88FF8AFE5EF35A6C6BC717" box="[389,567,601,623]" pageId="28" pageNumber="159">Lower Devonian</collectingMunicipality>
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,
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<collectingCounty id="391C4E7BFF88FF8AFD99F35A6CC4C717" box="[578,664,601,623]" pageId="28" pageNumber="159">Member</collectingCounty>
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8 (
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||
<bibRefCitation id="B4534B06FF88FF8AFF45F3716F0BC7F0" author="KLUBOV B. & KACHANOV E. & TALIMAA V." box="[158,343,626,648]" pageId="28" pageNumber="159" pagination="50 - 56" refId="ref36299" refString="KLUBOV B., KACHANOV E. & KARATAJUTE- TALIMAA V. 1980. - Stratigrafia silura i devona o. Pioner (Severnaya Zemlya) [= Stratigraphy of Silurian and Devonian of the Pioneer Island (Severnaya Zemlya)]. Izvestiya Akademii Nauk SSSR, seriya geologicheskaya 11: 50 - 56 (in Russian)." type="journal article" year="1980">
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Klubov
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||
<emphasis id="E2B6EAE5FF88FF8AFF2BF3706F7DC7FF" box="[240,289,626,648]" italics="true" pageId="28" pageNumber="159">et al.</emphasis>
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||
1980
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||
</bibRefCitation>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="D07D36F7FF88FF8AFF4CF3986F40C79B" blockId="28.[151,685,575,1281]" pageId="28" pageNumber="159">
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<typeStatus id="0F798855FF88FF8AFF4CF3986F56C7D7" box="[151,266,667,688]" pageId="28" pageNumber="159" type="paratype">PARATYPES</typeStatus>
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. — LIG 35-1937, 1938, 1940, 1968 and 1969: scales (
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<figureCitation id="48F92A72FF88FF8AFECDF3B76F00C7B2" box="[278,348,692,714]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Fig. 23</figureCitation>
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B-F). All from the same sample as the
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<typeStatus id="0F798855FF88FF8AFF67F3CE6F44C79B" box="[188,280,717,739]" pageId="28" pageNumber="159" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="98D8657CFF88FF8AFF4CF3F56F09C65D" pageId="28" pageNumber="159" type="etymology">
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<paragraph id="D07D36F7FF88FF8AFF4CF3F56F09C65D" blockId="28.[151,685,575,1281]" pageId="28" pageNumber="159">ETYMOLOGY. — In honour of D. M. S. Watson, acanthodian researcher.</paragraph>
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</subSubSection>
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<subSubSection id="98D8657CFF88FF8AFF4CF23B6FB9C138" pageId="28" pageNumber="159" type="materials_examined">
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<paragraph id="D07D36F7FF88FF8AFF4CF23B6FADC635" blockId="28.[151,685,575,1281]" box="[151,497,823,845]" pageId="28" pageNumber="159">
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||
<collectionCode id="B6D3AE32FF88FF8AFF4CF23B6F30C634" box="[151,364,824,845]" pageId="28" pageNumber="159">MATERIAL EXAMINED</collectionCode>
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. — 83 scales.
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</paragraph>
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<paragraph id="D07D36F7FF88FF8AFF4CF2636FB9C138" blockId="28.[151,685,575,1281]" pageId="28" pageNumber="159">
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LOCALITY
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<collectionCode id="B6D3AE32FF88FF8AFEDDF2676F6EC60D" box="[262,306,868,885]" pageId="28" pageNumber="159">AND</collectionCode>
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AGE. — October Revolution Island: Matusevich River, outcrop 5, bed 55; Spokojnaya River, outcrop 48, beds 5, 7 and outcrop 49, bed 11; Pod”emnaya River, observation point 26-3. Pioneer Island: samples P-12-7, 12-12, 10-1, 9-1 and observation point 2478. Lower Devonian, Emsian, Rusanov and Al’banov formations of the October Revolution Island and supposed analogues, in age, of the first one on Pioneer Island (Members 6-8).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="98D8657CFF88FF8AFF4CF5506FB4C079" pageId="28" pageNumber="159" type="diagnosis">
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||
<paragraph id="D07D36F7FF88FF8AFF4CF5506FB4C079" blockId="28.[151,685,575,1281]" pageId="28" pageNumber="159">
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||
DIAGNOSIS. —
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<taxonomicName id="17C24D74FF88FF8AFEE1F5506FFAC110" authorityName="Pander" authorityYear="1856" box="[314,422,1107,1128]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="genus">
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<emphasis id="E2B6EAE5FF88FF8AFEE1F5506FFAC110" box="[314,422,1107,1128]" italics="true" pageId="28" pageNumber="159">Nostolepis</emphasis>
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</taxonomicName>
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with scales of moderate size, rhombic crowns ornamented by six short, rounded, sub-parallel or sub-radial ridges, low neck and strongly anteriorly advanced base. Extremely thinlamellose cellular to acellular base bone, with a simple network of mesodentine and Stranggewebe in crown which can be replaced by dentine.
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</paragraph>
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</subSubSection>
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<subSubSection id="98D8657CFF88FF8AFF4CF4296D88C61B" pageId="28" pageNumber="159" type="description">
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||
<paragraph id="D07D36F7FF88FF8AFF4CF4296F74C03A" blockId="28.[151,684,1322,1667]" box="[151,296,1322,1348]" pageId="28" pageNumber="159">DESCRIPTION</paragraph>
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||
<paragraph id="D07D36F7FF88FF8AFF4CF4486A39C45C" blockId="28.[151,684,1322,1667]" lastBlockId="28.[714,1247,171,867]" pageId="28" pageNumber="159">
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||
Species described from disarticulated scales. Their size varies in length from
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<quantity id="173A9B12FF88FF8AFE26F4686CF4C0FB" box="[509,680,1387,1411]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="8.5" metricValueMin="4.0" pageId="28" pageNumber="159" unit="mm" value="0.625" valueMax="0.85" valueMin="0.4">0.4 to 0.85 mm</quantity>
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||
. Crown is rhombic, with a slightly elongated posterior area. Six short, rounded, sub-parallel or sub-radial converging ridges (
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<figureCitation id="48F92A72FF88FF8AFDC1F4C86CFFC09B" box="[538,675,1483,1507]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Fig. 23A, B</figureCitation>
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) occur along the proximal margin. Another scale variety (
|
||
<figureCitation id="48F92A72FF88FF8AFF25F7086FD3C35B" box="[254,399,1547,1571]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Fig. 23D, F</figureCitation>
|
||
) shows more regularly parallel, lower and longer ridges reaching the middle crown part. The scale base is massive, strongly vaulted anteriorly and protruding the crown (
|
||
<figureCitation id="48F92A72FF88FF8AFCFEF1A86DEFC5BB" box="[805,947,171,195]" captionStart="FIG" captionStartId="29.[151,162,1460,1477]" captionTargetBox="[160,1235,267,1396]" captionTargetId="figure-259@29.[162,1236,267,1396]" captionTargetPageId="29" captionText="FIG. 23. — A-F, Nostolepis watsoni n. sp., scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after Klubov et al. 1980); A, holotype, LIG 35-2020, crown; B, LIG 35-1968, crown; C, LIG 35-1938, base; D, LIG 35-1937, crown; E, LIG 35-1969, side; F, LIG 35-1940, crown; A, B, D, F, anterior upwards; C, to left; E, to right; G-K, Watsonacanthus costatus n. sp., scales; G, LIG 35-1967, crown; H, LIG 35-2140, crown; I, LIG 35-1422, side; J, LIG 35-1993, crown; K, holotype, LIG 35-2074, crown; anterior upwards (G, K, J), to the left (H) and to right (I); G-I, Pioneer Island, sample P-10-1, Lower Devonian, Member 8; J, K, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation; L, M, Acanthodii indet., scales, crowns, anterior upwards; L, LIG 35-1997; M, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation; N-P, Gomphonchus nordicus n. sp., scales; N, LIG 35-2006, crown, anterior upwards; O, LIG 35-2008, side, anterior to left; P, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="28" pageNumber="159">Fig. 23A, C</figureCitation>
|
||
). A distinct sub-rhombic rim, separating it from a very low neck that thickens posteriorly, outlines it. Postero-lateral neck walls may bear vertical grooves.
|
||
</paragraph>
|
||
<paragraph id="D07D36F7FF88FF8AFD11F0286D88C61B" blockId="28.[714,1247,171,867]" pageId="28" pageNumber="159">
|
||
The two noted scale varieties have a slightly different histological structure. The short-ridged crowns (like the
|
||
<typeStatus id="0F798855FF88FF8AFC43F0686A59C4FB" box="[920,1029,363,387]" pageId="28" pageNumber="159" type="holotype">holotype</typeStatus>
|
||
) are composed of mesodentine with a typical network of tubules interconnected with osteocyte spaces (
|
||
<figureCitation id="48F92A72FF88FF8AFBACF0A86CBFC49B" captionStart="FIG" captionStartId="30.[151,162,1011,1028]" captionTargetBox="[177,1214,172,973]" captionTargetId="figure-364@30.[163,1234,220,992]" captionTargetPageId="30" captionText="FIG. 24. — Nostolepis watsoni n. sp., scale histology, Pioneer Island, sample P-9-1, Lower Devonian, Member 8; A, B, vertical transverse sections through the scales with elongated flattened ridges (see Fig. 23 D), thin sections 3286 (A) and 3266 (B); C, D, vertical longitudinal sections through the scales with short, rounded, higher ridges (see Fig. 23A), thin sections 3267 (C) and 3268 (D). Abbreviations: abo, acellular bone; avc, ascending vascular canal; cbo, cellular bone; gl, growth lamella; oc, osteocyte cavity; prgl, primordial growth lamella; smd, simple mesodentine; stg, Sranggewebe (oriented mesodentine). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665157" httpUri="https://zenodo.org/record/4665157/files/figure.png" pageId="28" pageNumber="159">Fig. 24C, D</figureCitation>
|
||
). The small posterior area demonstrates a Stranggewebe only superficially developed. It never occurs in the primordial lamella. The extremely thin-layered base is composed of cellular bone. The second scale
|
||
<taxonomicName id="17C24D74FF88FF8AFBC7F3486AC6C71B" box="[1052,1178,587,611]" pageId="28" pageNumber="159" rank="variety" variety="has">variety has</taxonomicName>
|
||
bases consisting of almost acellular bone and crowns made of dentine (
|
||
<figureCitation id="48F92A72FF88FF8AFC4FF3886A40C7DB" box="[916,1052,651,675]" captionStart="FIG" captionStartId="30.[151,162,1011,1028]" captionTargetBox="[177,1214,172,973]" captionTargetId="figure-364@30.[163,1234,220,992]" captionTargetPageId="30" captionText="FIG. 24. — Nostolepis watsoni n. sp., scale histology, Pioneer Island, sample P-9-1, Lower Devonian, Member 8; A, B, vertical transverse sections through the scales with elongated flattened ridges (see Fig. 23 D), thin sections 3286 (A) and 3266 (B); C, D, vertical longitudinal sections through the scales with short, rounded, higher ridges (see Fig. 23A), thin sections 3267 (C) and 3268 (D). Abbreviations: abo, acellular bone; avc, ascending vascular canal; cbo, cellular bone; gl, growth lamella; oc, osteocyte cavity; prgl, primordial growth lamella; smd, simple mesodentine; stg, Sranggewebe (oriented mesodentine). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665157" httpUri="https://zenodo.org/record/4665157/files/figure.png" pageId="28" pageNumber="159">Fig. 24A, B</figureCitation>
|
||
). The number of growth lamellae is the same: eight. Ascending vascular canals are smooth, with long widened principal and shorter side branches. No lacunae or osteocytes are met. A complex central knot (
|
||
<figureCitation id="48F92A72FF88FF8AFD09F2286D65C63B" box="[722,825,811,835]" captionStart="FIG" captionStartId="30.[151,162,1011,1028]" captionTargetBox="[177,1214,172,973]" captionTargetId="figure-364@30.[163,1234,220,992]" captionTargetPageId="30" captionText="FIG. 24. — Nostolepis watsoni n. sp., scale histology, Pioneer Island, sample P-9-1, Lower Devonian, Member 8; A, B, vertical transverse sections through the scales with elongated flattened ridges (see Fig. 23 D), thin sections 3286 (A) and 3266 (B); C, D, vertical longitudinal sections through the scales with short, rounded, higher ridges (see Fig. 23A), thin sections 3267 (C) and 3268 (D). Abbreviations: abo, acellular bone; avc, ascending vascular canal; cbo, cellular bone; gl, growth lamella; oc, osteocyte cavity; prgl, primordial growth lamella; smd, simple mesodentine; stg, Sranggewebe (oriented mesodentine). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4665157" httpUri="https://zenodo.org/record/4665157/files/figure.png" pageId="28" pageNumber="159">Fig. 24A</figureCitation>
|
||
) of dentine canals is seen in the primordial growth lamella.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="98D8657CFF88FF89FD11F2896C6DC1FB" lastPageId="31" lastPageNumber="162" pageId="28" pageNumber="159" type="discussion">
|
||
<paragraph id="D07D36F7FF88FF8AFD11F2896D10C6DA" blockId="28.[714,1248,906,1667]" box="[714,844,906,932]" pageId="28" pageNumber="159">DISCUSSION</paragraph>
|
||
<paragraph id="D07D36F7FF88FF8AFD11F2AF6A83C3FB" blockId="28.[714,1248,906,1667]" pageId="28" pageNumber="159">
|
||
By its crown ornamentation and its specific, largely advanced, massive and anteriorly vaulted base,
|
||
<taxonomicName id="17C24D74FF88FF8AFCD2F2E86D8BC17B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[777,983,1003,1027]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF88FF8AFCD2F2E86D8BC17B" box="[777,983,1003,1027]" italics="true" pageId="28" pageNumber="159">Nostolepis watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF88FF8AFC3BF2EF6A41C17B" box="[992,1053,1004,1027]" pageId="28" pageNumber="159" rank="species">n. sp.</taxonomicNameLabel>
|
||
is comparable to several representatives belonging to different genera. The closest one is
|
||
<taxonomicName id="17C24D74FF88FF8AFBCCF5286ACBC13B" authorityName="Pander" authorityYear="1856" box="[1047,1175,1067,1091]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF88FF8AFBCCF5286ACBC13B" box="[1047,1175,1067,1091]" italics="true" pageId="28" pageNumber="159">Nostolepis</emphasis>
|
||
</taxonomicName>
|
||
sp. B defined by Vyushkova from the Telengitian upper Salairka beds and Belovo Horizon of Salair (
|
||
<bibRefCitation id="B4534B06FF88FF8AFCC7F5886DB4C1DB" author="VYUSHKOVA L." box="[796,1000,1163,1187]" pageId="28" pageNumber="159" pagination="281 - 287" refId="ref37602" refString="VYUSHKOVA L. 1992. - Fish assemblages and facies in the Telengitian Suprahorizon of Salair, in MARK- KURIK E. (ed.), Fossil fishes as living animals. Academia 1: 281 - 287." type="journal article" year="1992">Vyushkova 1992</bibRefCitation>
|
||
: pl. 1, figs 7-9, pl. 3, fig. 1). The histological structure of scales has not been investigated. Morphologic differences concern the more numerous and regular subparallel ridges occurring in the Salair specimens. This is also supposedly said about the weakly ridged scales of
|
||
<taxonomicName id="17C24D74FF88FF8AFCA1F4486AFCC01B" box="[890,1184,1355,1379]" genus="Trundlelepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="cervicostulata">
|
||
<emphasis id="E2B6EAE5FF88FF8AFCA1F4486AFCC01B" box="[890,1184,1355,1379]" italics="true" pageId="28" pageNumber="159">Trundlelepis cervicostulata</emphasis>
|
||
</taxonomicName>
|
||
morphotype 2 (
|
||
<bibRefCitation id="B4534B06FF88FF8AFC8BF4686DB1C0FB" author="BURROW C. J." box="[848,1005,1387,1411]" pageId="28" pageNumber="159" pagination="1 - 5" refId="ref35734" refString="BURROW C. J. 1997. - Microvertebrate assemblages from the Lower Devonian (pesavis / sulcatus zones) of central New South Wales, Australia. Modern Geology 21: 43 - 77; Addendum, October 1997: 1 - 5, 7 figs." type="book chapter" year="1997">Burrow 1997</bibRefCitation>
|
||
), identified from the Lochkovian Trundle beds of
|
||
<collectingRegion id="1206F815FF88FF8AFBD4F48F6A87C0DB" box="[1039,1243,1419,1443]" country="Australia" name="New South Wales" pageId="28" pageNumber="159">New South Wales</collectingRegion>
|
||
,
|
||
<collectingCountry id="A8D57667FF88FF8AFD11F4A86D6CC0BB" box="[714,816,1451,1475]" name="Australia" pageId="28" pageNumber="159">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B4534B06FF88FF8AFC9AF4A86D86C0BB" author="BURROW C. J." box="[833,986,1451,1475]" pageId="28" pageNumber="159" pagination="1 - 5" refId="ref35734" refString="BURROW C. J. 1997. - Microvertebrate assemblages from the Lower Devonian (pesavis / sulcatus zones) of central New South Wales, Australia. Modern Geology 21: 43 - 77; Addendum, October 1997: 1 - 5, 7 figs." type="book chapter" year="1997">Burrow 1997</bibRefCitation>
|
||
: pl. 1, fig. 17a, b, pl. 3, fig. 3). As opposed to
|
||
<taxonomicName id="17C24D74FF88FF8AFC0FF4CF6A0FC09B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[980,1107,1484,1507]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF88FF8AFC0FF4CF6DB6C09B" box="[980,1002,1484,1507]" italics="true" pageId="28" pageNumber="159">N</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF88FF8AFC21F4CF6A0FC09B" box="[1018,1107,1484,1507]" italics="true" pageId="28" pageNumber="159">watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF88FF8AFB87F4CF6AC1C09B" box="[1116,1181,1484,1507]" pageId="28" pageNumber="159" rank="species">n. sp.</taxonomicNameLabel>
|
||
, they bear six to 12 anterior ridges, and about 10% of scales contain crown pore openings, which are not observed in
|
||
<taxonomicName id="17C24D74FF88FF8AFC5DF72F6A5FC33B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[902,1027,1580,1603]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF88FF8AFC5DF72F6DC0C33B" box="[902,924,1580,1603]" italics="true" pageId="28" pageNumber="159">N</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF88FF8AFC77F72F6A5FC33B" box="[940,1027,1580,1603]" italics="true" pageId="28" pageNumber="159">watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF88FF8AFBD7F72F6A16C33B" box="[1036,1098,1580,1603]" pageId="28" pageNumber="159" rank="species">n. sp.</taxonomicNameLabel>
|
||
<taxonomicName id="17C24D74FF88FF8AFB89F7286D3FC31B" genus="Trundlelepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="28" pageNumber="159" phylum="Chordata" rank="species" species="cervicostulata">
|
||
<emphasis id="E2B6EAE5FF88FF8AFB89F7286D3FC31B" italics="true" pageId="28" pageNumber="159">Trundlelepis cervicostulata</emphasis>
|
||
</taxonomicName>
|
||
scale bases are also composed of cellular (
|
||
<bibRefCitation id="B4534B06FF88FF8AFCF7F7686D9AC3FB" author="BURROW C. J." box="[812,966,1643,1667]" pageId="28" pageNumber="159" pagination="1 - 5" refId="ref35734" refString="BURROW C. J. 1997. - Microvertebrate assemblages from the Lower Devonian (pesavis / sulcatus zones) of central New South Wales, Australia. Modern Geology 21: 43 - 77; Addendum, October 1997: 1 - 5, 7 figs." type="book chapter" year="1997">Burrow 1997</bibRefCitation>
|
||
: fig. 5C and explanatory
|
||
</paragraph>
|
||
<paragraph id="D07D36F7FF89FF8BFF7FF1ED6D99C47F" blockId="29.[164,695,238,263]" box="[164,965,238,263]" lastBlockId="29.[946,965,238,263]" pageId="29" pageNumber="160">A B C D</paragraph>
|
||
<caption id="84BD667FFF89FF8BFF4CF4B76C19C3FD" ID-DOI="http://doi.org/10.5281/zenodo.4665155" ID-Zenodo-Dep="4665155" httpUri="https://zenodo.org/record/4665155/files/figure.png" pageId="29" pageNumber="160" startId="29.[151,162,1460,1477]" targetBox="[160,1235,267,1396]" targetPageId="29">
|
||
<paragraph id="D07D36F7FF89FF8BFF4CF4B76C19C3FD" blockId="29.[151,1249,1460,1669]" pageId="29" pageNumber="160">
|
||
FIG. 23. —
|
||
<emphasis id="E2B6EAE5FF89FF8BFF2DF4B76F4AC0BD" bold="true" box="[246,278,1460,1477]" pageId="29" pageNumber="160">A -F</emphasis>
|
||
,
|
||
<taxonomicName id="17C24D74FF89FF8BFEF9F4B76F9DC0BD" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[290,449,1460,1477]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="29" pageNumber="160" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF89FF8BFEF9F4B76F9DC0BD" box="[290,449,1460,1477]" italics="true" pageId="29" pageNumber="160">Nostolepis watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF89FF8BFE13F4B76FA1C0BD" box="[456,509,1460,1477]" pageId="29" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
, scales, Pioneer Island, sample P-10-1, Lower Devonian, Member 8. (Here and further members of the Pioneer Island geological sequence taken after
|
||
<bibRefCitation id="B4534B06FF89FF8BFD78F4CA6D66C0A2" author="KLUBOV B. & KACHANOV E. & TALIMAA V." box="[675,826,1481,1498]" pageId="29" pageNumber="160" pagination="50 - 56" refId="ref36299" refString="KLUBOV B., KACHANOV E. & KARATAJUTE- TALIMAA V. 1980. - Stratigrafia silura i devona o. Pioner (Severnaya Zemlya) [= Stratigraphy of Silurian and Devonian of the Pioneer Island (Severnaya Zemlya)]. Izvestiya Akademii Nauk SSSR, seriya geologicheskaya 11: 50 - 56 (in Russian)." type="journal article" year="1980">
|
||
Klubov
|
||
<emphasis id="E2B6EAE5FF89FF8BFD3AF4CA6D56C0A2" box="[737,778,1481,1498]" italics="true" pageId="29" pageNumber="160">et al.</emphasis>
|
||
1980
|
||
</bibRefCitation>
|
||
);
|
||
<emphasis id="E2B6EAE5FF89FF8BFC9EF4CA6D0EC0A2" bold="true" box="[837,850,1481,1498]" pageId="29" pageNumber="160">A</emphasis>
|
||
, holotype, LIG 35-2020, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFB87F4CA6A35C0A2" bold="true" box="[1116,1129,1481,1498]" pageId="29" pageNumber="160">B</emphasis>
|
||
, LIG 35-1968, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFF0DF4DC6EB8C088" bold="true" box="[214,228,1503,1520]" pageId="29" pageNumber="160">C</emphasis>
|
||
, LIG 35-1938, base;
|
||
<emphasis id="E2B6EAE5FF89FF8BFE43F4DC6FFAC088" bold="true" box="[408,422,1503,1520]" pageId="29" pageNumber="160">D</emphasis>
|
||
, LIG 35-1937, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFDBEF4DC6C2DC088" bold="true" box="[613,625,1503,1520]" pageId="29" pageNumber="160">E</emphasis>
|
||
, LIG 35-1969, side;
|
||
<emphasis id="E2B6EAE5FF89FF8BFCFBF4DC6D77C088" bold="true" box="[800,811,1503,1520]" pageId="29" pageNumber="160">F</emphasis>
|
||
, LIG 35-1940, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFC31F4DC6DABC088" bold="true" box="[1002,1015,1503,1520]" pageId="29" pageNumber="160">A</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFBD8F4DC6A4CC088" bold="true" box="[1027,1040,1503,1520]" pageId="29" pageNumber="160">B</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFBC0F4DC6A75C088" bold="true" box="[1051,1065,1503,1520]" pageId="29" pageNumber="160">D</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFBEEF4DC6A1CC088" bold="true" box="[1077,1088,1503,1520]" pageId="29" pageNumber="160">F</emphasis>
|
||
, anterior upwards;
|
||
<emphasis id="E2B6EAE5FF89FF8BFF4CF4F76EF9C37D" bold="true" box="[151,165,1524,1541]" pageId="29" pageNumber="160">C</emphasis>
|
||
, to left;
|
||
<emphasis id="E2B6EAE5FF89FF8BFF31F4F76EAAC37D" bold="true" box="[234,246,1524,1541]" pageId="29" pageNumber="160">E</emphasis>
|
||
, to right;
|
||
<emphasis id="E2B6EAE5FF89FF8BFE93F4F76F36C37D" bold="true" box="[328,362,1524,1541]" pageId="29" pageNumber="160">G -K</emphasis>
|
||
,
|
||
<taxonomicName id="17C24D74FF89FF8BFEAEF4F76C17C37D" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[373,587,1524,1541]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="29" pageNumber="160" phylum="Chordata" rank="species" species="costatus" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF89FF8BFEAEF4F76C17C37D" box="[373,587,1524,1541]" italics="true" pageId="29" pageNumber="160">Watsonacanthus costatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF89FF8BFD89F4F76CD8C37D" box="[594,644,1524,1541]" pageId="29" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
, scales;
|
||
<emphasis id="E2B6EAE5FF89FF8BFD10F4F76C85C37D" bold="true" box="[715,729,1524,1541]" pageId="29" pageNumber="160">G</emphasis>
|
||
, LIG 35-1967, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFC4DF4F76DF8C37D" bold="true" box="[918,932,1524,1541]" pageId="29" pageNumber="160">H</emphasis>
|
||
, LIG 35-2140, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFBBAF4F76A3BC37D" bold="true" box="[1121,1127,1524,1541]" pageId="29" pageNumber="160">I</emphasis>
|
||
, LIG 35-1422, side;
|
||
<emphasis id="E2B6EAE5FF89FF8BFF1FF70A6E92C362" bold="true" box="[196,206,1545,1562]" pageId="29" pageNumber="160">J</emphasis>
|
||
, LIG 35-1993, crown;
|
||
<emphasis id="E2B6EAE5FF89FF8BFE51F70A6FCBC362" bold="true" box="[394,407,1545,1562]" pageId="29" pageNumber="160">K</emphasis>
|
||
, holotype, LIG 35-2074, crown; anterior upwards (
|
||
<emphasis id="E2B6EAE5FF89FF8BFCE5F70A6D10C362" bold="true" box="[830,844,1545,1562]" pageId="29" pageNumber="160">G</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFC8DF70A6D3FC362" bold="true" box="[854,867,1545,1562]" pageId="29" pageNumber="160">K</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFCB4F70A6D25C362" bold="true" box="[879,889,1545,1562]" pageId="29" pageNumber="160">J</emphasis>
|
||
), to the left (
|
||
<emphasis id="E2B6EAE5FF89FF8BFC39F70A6DACC362" bold="true" box="[994,1008,1545,1562]" pageId="29" pageNumber="160">H</emphasis>
|
||
) and to right (
|
||
<emphasis id="E2B6EAE5FF89FF8BFBBEF70A6A37C362" bold="true" box="[1125,1131,1545,1562]" pageId="29" pageNumber="160">I</emphasis>
|
||
);
|
||
<emphasis id="E2B6EAE5FF89FF8BFBA1F70A6AC9C362" bold="true" box="[1146,1173,1545,1562]" pageId="29" pageNumber="160">G -I</emphasis>
|
||
, Pioneer Island, sample P-10-1, Lower Devonian, Member 8;
|
||
<emphasis id="E2B6EAE5FF89FF8BFD8EF71C6C03C348" bold="true" box="[597,607,1567,1584]" pageId="29" pageNumber="160">J</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFDB0F71C6C24C348" bold="true" box="[619,632,1567,1584]" pageId="29" pageNumber="160">K</emphasis>
|
||
, October Revolution Island, Spokojnaya River, outcrop 48, bed 5, lower Al’banov Subformation;
|
||
<emphasis id="E2B6EAE5FF89FF8BFEB9F7376F31C33D" bold="true" box="[354,365,1588,1605]" pageId="29" pageNumber="160">L</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF89FF8BFEA2F7376FD6C33D" bold="true" box="[377,394,1588,1605]" pageId="29" pageNumber="160">M</emphasis>
|
||
,
|
||
<taxonomicName id="17C24D74FF89FF8BFE4EF7376FB2C33D" box="[405,494,1588,1605]" class="Acanthodii" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="29" pageNumber="160" phylum="Chordata" rank="class">Acanthodii</taxonomicName>
|
||
indet., scales, crowns, anterior upwards;
|
||
<emphasis id="E2B6EAE5FF89FF8BFC8BF7376D07C33D" bold="true" box="[848,859,1588,1605]" pageId="29" pageNumber="160">L</emphasis>
|
||
, LIG 35-1997;
|
||
<emphasis id="E2B6EAE5FF89FF8BFC00F7376DB0C33D" bold="true" box="[987,1004,1588,1605]" pageId="29" pageNumber="160">M</emphasis>
|
||
, LIG 35-1998, Komsomolets Island, outcrop 18077, bed 9, Spokojnaya Formation;
|
||
<emphasis id="E2B6EAE5FF89FF8BFD86F74A6C23C322" bold="true" box="[605,639,1609,1626]" pageId="29" pageNumber="160">N -P</emphasis>
|
||
,
|
||
<taxonomicName id="17C24D74FF89FF8BFD51F74A6D0FC322" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[650,851,1609,1626]" class="Acanthodii" family="Ischnacanthidae" genus="Gomphonchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="29" pageNumber="160" phylum="Chordata" rank="species" species="nordicus" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF89FF8BFD51F74A6D0FC322" box="[650,851,1609,1626]" italics="true" pageId="29" pageNumber="160">Gomphonchus nordicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF89FF8BFC82F74A6DD0C322" box="[857,908,1609,1626]" pageId="29" pageNumber="160" rank="species">n. sp.</taxonomicNameLabel>
|
||
, scales;
|
||
<emphasis id="E2B6EAE5FF89FF8BFC08F74A6DBDC322" bold="true" box="[979,993,1609,1626]" pageId="29" pageNumber="160">N</emphasis>
|
||
, LIG 35-2006, crown, anterior upwards;
|
||
<emphasis id="E2B6EAE5FF89FF8BFF31F75C6EA5C308" bold="true" box="[234,249,1631,1648]" pageId="29" pageNumber="160">O</emphasis>
|
||
, LIG 35-2008, side, anterior to left;
|
||
<emphasis id="E2B6EAE5FF89FF8BFDF2F75C6C69C308" bold="true" box="[553,565,1631,1648]" pageId="29" pageNumber="160">P</emphasis>
|
||
, holotype, LIG 35-2007, crown, anterior upwards, Pioneer Island, sample 12-76, Upper Silurian, Pridoli, Member 3. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="84BD667FFF8AFF88FF4CF2F06AD0C121" ID-DOI="http://doi.org/10.5281/zenodo.4665157" ID-Zenodo-Dep="4665157" httpUri="https://zenodo.org/record/4665157/files/figure.png" pageId="30" pageNumber="161" startId="30.[151,162,1011,1028]" targetBox="[177,1214,172,973]" targetPageId="30">
|
||
<paragraph id="D07D36F7FF8AFF88FF4CF2F06AD0C121" blockId="30.[151,1249,1011,1113]" pageId="30" pageNumber="161">
|
||
FIG. 24. —
|
||
<taxonomicName id="17C24D74FF8AFF88FF29F2F06FD3C17C" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[242,399,1011,1028]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8AFF88FF29F2F06FD3C17C" box="[242,399,1011,1028]" italics="true" pageId="30" pageNumber="161">Nostolepis watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8AFF88FE4FF2F06F9AC17C" box="[404,454,1011,1028]" pageId="30" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
, scale histology, Pioneer Island, sample P-9-1, Lower Devonian, Member 8;
|
||
<emphasis id="E2B6EAE5FF8AFF88FBE4F2F06A10C17C" bold="true" box="[1087,1100,1011,1028]" pageId="30" pageNumber="161">A</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF8AFF88FB8DF2F06A3FC17C" bold="true" box="[1110,1123,1011,1028]" pageId="30" pageNumber="161">B</emphasis>
|
||
, vertical transverse sections through the scales with elongated flattened ridges (see Fig. 23 D), thin sections 3286 (
|
||
<emphasis id="E2B6EAE5FF8AFF88FC3EF50B6DAEC161" bold="true" box="[997,1010,1032,1049]" pageId="30" pageNumber="161">A</emphasis>
|
||
) and 3266 (
|
||
<emphasis id="E2B6EAE5FF8AFF88FB8DF50B6A3FC161" bold="true" box="[1110,1123,1032,1049]" pageId="30" pageNumber="161">B</emphasis>
|
||
);
|
||
<emphasis id="E2B6EAE5FF8AFF88FBA9F50B6ADCC161" bold="true" box="[1138,1152,1032,1049]" pageId="30" pageNumber="161">C</emphasis>
|
||
,
|
||
<emphasis id="E2B6EAE5FF8AFF88FB50F50B6AC5C161" bold="true" box="[1163,1177,1032,1049]" pageId="30" pageNumber="161">D</emphasis>
|
||
, vertical longitudinal sections through the scales with short, rounded, higher ridges (see Fig. 23A), thin sections 3267 (
|
||
<emphasis id="E2B6EAE5FF8AFF88FB8BF51E6A02C156" bold="true" box="[1104,1118,1053,1070]" pageId="30" pageNumber="161">C</emphasis>
|
||
) and 3268 (
|
||
<emphasis id="E2B6EAE5FF8AFF88FB1CF51E6A89C156" bold="true" box="[1223,1237,1053,1070]" pageId="30" pageNumber="161">D</emphasis>
|
||
). Abbreviations:
|
||
<emphasis id="E2B6EAE5FF8AFF88FEC3F5306F66C13C" bold="true" box="[280,314,1075,1092]" pageId="30" pageNumber="161">abo</emphasis>
|
||
, acellular bone;
|
||
<emphasis id="E2B6EAE5FF8AFF88FE10F5306FB6C13C" bold="true" box="[459,490,1075,1092]" pageId="30" pageNumber="161">avc</emphasis>
|
||
, ascending vascular canal;
|
||
<emphasis id="E2B6EAE5FF8AFF88FD06F5306CA3C13C" bold="true" box="[733,767,1075,1092]" pageId="30" pageNumber="161">cbo</emphasis>
|
||
, cellular bone;
|
||
<emphasis id="E2B6EAE5FF8AFF88FC5EF5306DC9C13C" bold="true" box="[901,917,1075,1092]" pageId="30" pageNumber="161">gl</emphasis>
|
||
, growth lamella;
|
||
<emphasis id="E2B6EAE5FF8AFF88FBF7F5306A1EC13C" bold="true" box="[1068,1090,1075,1092]" pageId="30" pageNumber="161">oc</emphasis>
|
||
, osteocyte cavity;
|
||
<emphasis id="E2B6EAE5FF8AFF88FF4CF54B6EE6C121" bold="true" box="[151,186,1096,1113]" pageId="30" pageNumber="161">prgl</emphasis>
|
||
, primordial growth lamella;
|
||
<emphasis id="E2B6EAE5FF8AFF88FE79F54B6F94C121" bold="true" box="[418,456,1096,1113]" pageId="30" pageNumber="161">smd</emphasis>
|
||
, simple mesodentine;
|
||
<emphasis id="E2B6EAE5FF8AFF88FD5FF54B6CFCC121" bold="true" box="[644,672,1096,1113]" pageId="30" pageNumber="161">stg</emphasis>
|
||
, Sranggewebe (oriented mesodentine). Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D07D36F7FF8AFF88FF4CF5A86EA3C0BB" blockId="30.[151,684,1195,1667]" pageId="30" pageNumber="161">
|
||
fig. 5-
|
||
<quantity id="173A9B12FF8AFF88FF05F5A86F51C1BB" box="[222,269,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="30" pageNumber="161" unit="in" value="3.0">3 in</quantity>
|
||
addendum 1997) to acellular bone (fig. 5D and fig. 5-4). Their simple mesodentine (figs 5-2 and 5-
|
||
<quantity id="173A9B12FF8AFF88FE9CF5E86F28C07B" box="[327,372,1259,1283]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="30" pageNumber="161" unit="in" value="5.0">5 in</quantity>
|
||
addendum 1997) in crown contains short winding dentine tubules with lacunae expansions comparable only to the first variety of scales in
|
||
<taxonomicName id="17C24D74FF8AFF88FE09F44F6C00C01B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[466,604,1356,1379]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8AFF88FE09F44F6FB4C01B" box="[466,488,1356,1379]" italics="true" pageId="30" pageNumber="161">N</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FE27F44F6C00C01B" box="[508,604,1356,1379]" italics="true" pageId="30" pageNumber="161">watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8AFF88FDBCF44F6CF0C01B" box="[615,684,1356,1379]" pageId="30" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
Stranggewebe or dentine tissue with long ascending vascular canals is not seen in
|
||
<taxonomicName id="17C24D74FF8AFF88FD96F48F6EA4C0BB" genus="Trundlelepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="cervicostulata">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD96F48F6C02C0DB" box="[589,606,1420,1443]" italics="true" pageId="30" pageNumber="161">T</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FDB7F48F6EA4C0BB" italics="true" pageId="30" pageNumber="161">cervicostulata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D07D36F7FF8AFF88FF4CF4C86A3AC19B" blockId="30.[151,684,1195,1667]" lastBlockId="30.[714,1248,1195,1667]" pageId="30" pageNumber="161">
|
||
Several morphologic similarities unite
|
||
<taxonomicName id="17C24D74FF8AFF88FD89F4CF6E98C37B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD89F4CF6C34C09B" box="[594,616,1484,1507]" italics="true" pageId="30" pageNumber="161">N</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FDACF4CF6E98C37B" italics="true" pageId="30" pageNumber="161">watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8AFF88FF17F4EF6F57C37B" box="[204,267,1516,1539]" pageId="30" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
with the scales of
|
||
<taxonomicName id="17C24D74FF8AFF88FE3DF4E86CF0C37B" authorityName="Wells" authorityYear="1944" box="[486,684,1515,1539]" class="Acanthodii" family="Climatiidae" genus="Cheiracanthoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8AFF88FE3DF4E86CF0C37B" box="[486,684,1515,1539]" italics="true" pageId="30" pageNumber="161">Cheiracanthoides</emphasis>
|
||
</taxonomicName>
|
||
sp. indet. from the Emsian La Grange Limestone of Armorican Massif,
|
||
<collectingCountry id="A8D57667FF8AFF88FE0BF72F6C41C33B" box="[464,541,1580,1603]" name="France" pageId="30" pageNumber="161">France</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B4534B06FF8AFF88FDF7F7286E8CC31B" author="VIDAL M. & GOUJET D. & JANVIER P. & LARDEUX H." pageId="30" pageNumber="161" pagination="321 - 341" refId="ref37509" refString="VIDAL M., GOUJET D., JANVIER P. & LARDEUX H. 1994. - Micro-restes de Vertebres du Calcaire de La Grange, Devonien inferieur (Emsien) du Sud- Est du Massif Armoricain. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 194 (2 / 3): 321 - 341." type="journal article" year="1994">
|
||
Vidal
|
||
<emphasis id="E2B6EAE5FF8AFF88FDA8F72F6CF0C33B" box="[627,684,1579,1603]" italics="true" pageId="30" pageNumber="161">et al.</emphasis>
|
||
1994
|
||
</bibRefCitation>
|
||
: fig. 4.11). However the crown ridges of figured specimen seem sharper when compared to
|
||
<taxonomicName id="17C24D74FF8AFF88FD33F5AF6D03C1BB" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[744,863,1196,1219]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD33F5AF6CA2C1BB" box="[744,766,1196,1219]" italics="true" pageId="30" pageNumber="161">N</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FCD1F5AF6D03C1BB" box="[778,863,1196,1219]" italics="true" pageId="30" pageNumber="161">watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8AFF88FCBDF5AF6DFCC1BB" box="[870,928,1196,1219]" pageId="30" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
The histological structure of these scales is not taken into account.
|
||
</paragraph>
|
||
<paragraph id="D07D36F7FF8AFF88FD11F5EF6A25C3FB" blockId="30.[714,1248,1195,1667]" pageId="30" pageNumber="161">
|
||
By most morphologic and histological features
|
||
<taxonomicName id="17C24D74FF8AFF88FD11F4086DF6C05B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[714,938,1291,1315]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="watsoni" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD11F4086DF6C05B" box="[714,938,1291,1315]" italics="true" pageId="30" pageNumber="161">Nostolepis watsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8AFF88FC68F40F6DAFC05B" box="[947,1011,1292,1315]" pageId="30" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
|
||
resembles scales of
|
||
<taxonomicName id="17C24D74FF8AFF88FD11F4286A33C03B" authority="Valiukevicius, 1979" authorityName="Valiukevicius" authorityYear="1979" box="[714,1135,1323,1347]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="161" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD11F4286DD8C03B" box="[714,900,1323,1347]" italics="true" pageId="30" pageNumber="161">Watsonacanthus</emphasis>
|
||
<bibRefCitation id="B4534B06FF8AFF88FC57F4286A33C03B" author="VALIUKEVICIUS J." box="[908,1135,1323,1347]" pageId="30" pageNumber="161" pagination="101 - 111" refId="ref36831" refString="VALIUKEVICIUS J. 1979. - Acanthodian scales from the Eifelian of Spitsbergen. Paleontological Journal 4: 101 - 111." type="journal article" year="1979">Valiukevicius, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
. It can be marked as an intermediate form between certain
|
||
<taxonomicName id="17C24D74FF8AFF88FD11F4686D68C0FB" authorityName="Pander" authorityYear="1856" box="[714,820,1387,1411]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="30" pageNumber="161" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD11F4686D68C0FB" box="[714,820,1387,1411]" italics="true" pageId="30" pageNumber="161">Nostolepis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="17C24D74FF8AFF88FCBFF4686A51C0FB" authorityName="Valiukevicius" authorityYear="1979" box="[868,1037,1387,1411]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="161" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8AFF88FCBFF4686A51C0FB" box="[868,1037,1387,1411]" italics="true" pageId="30" pageNumber="161">Watsonacanthus</emphasis>
|
||
</taxonomicName>
|
||
lineages. By its converging rounded and flattened ridges it is closer to
|
||
<taxonomicName id="17C24D74FF8AFF88FD11F4AF6A78C0BB" authority="Valiukevicius, 1979" authorityName="Valiukevicius" authorityYear="1979" box="[714,1060,1451,1475]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="oervigi">
|
||
<emphasis id="E2B6EAE5FF8AFF88FD11F4AF6CBFC0BB" box="[714,739,1452,1475]" italics="true" pageId="30" pageNumber="161">W</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FD2AF4AF6D1CC0BB" box="[753,832,1452,1475]" italics="true" pageId="30" pageNumber="161">oervigi</emphasis>
|
||
<bibRefCitation id="B4534B06FF8AFF88FC93F4A86A78C0BB" author="VALIUKEVICIUS J." box="[840,1060,1451,1475]" pageId="30" pageNumber="161" pagination="101 - 111" refId="ref36831" refString="VALIUKEVICIUS J. 1979. - Acanthodian scales from the Eifelian of Spitsbergen. Paleontological Journal 4: 101 - 111." type="journal article" year="1979">Valiukevicius, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
(pl. 11, figs 1-8, pl. 12, figs 1-4).
|
||
<taxonomicName id="17C24D74FF8AFF88FCA2F4CF6DB5C09B" authorityName="Valiukevicius" authorityYear="1979" box="[889,1001,1484,1507]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="30" pageNumber="161" phylum="Chordata" rank="species" species="oervigi">
|
||
<emphasis id="E2B6EAE5FF8AFF88FCA2F4CF6DCDC09B" box="[889,913,1484,1507]" italics="true" pageId="30" pageNumber="161">W</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8AFF88FC44F4CF6DB5C09B" box="[927,1001,1484,1507]" italics="true" pageId="30" pageNumber="161">oervigi</emphasis>
|
||
</taxonomicName>
|
||
scale bases are characterised by a highly cellular bone (
|
||
<bibRefCitation id="B4534B06FF8AFF88FB9DF4E86D5DC35B" author="VALIUKEVICIUS J." pageId="30" pageNumber="161" pagination="101 - 111" refId="ref36831" refString="VALIUKEVICIUS J. 1979. - Acanthodian scales from the Eifelian of Spitsbergen. Paleontological Journal 4: 101 - 111." type="journal article" year="1979">Valiukevičius 1979</bibRefCitation>
|
||
: fig. 1a-d; 1994: fig. 75.5), containing dense multiangular osteocytes, and a simple mesodentine network in crowns without forming main branches. Stranggewebe has also not been observed.
|
||
</paragraph>
|
||
<paragraph id="D07D36F7FF8BFF89FF72F1AD6F83C5BF" blockId="31.[169,479,174,199]" box="[169,479,174,199]" pageId="31" pageNumber="162">A B</paragraph>
|
||
<caption id="84BD667FFF8BFF89FF4CF0F76E80C73D" ID-DOI="http://doi.org/10.5281/zenodo.4665159" ID-Zenodo-Dep="4665159" httpUri="https://zenodo.org/record/4665159/files/figure.png" pageId="31" pageNumber="162" startId="31.[151,162,500,517]" targetBox="[169,667,201,462]" targetPageId="31">
|
||
<paragraph id="D07D36F7FF8BFF89FF4CF0F76E80C73D" blockId="31.[151,685,500,581]" pageId="31" pageNumber="162">
|
||
FIG. 25. —
|
||
<taxonomicName id="17C24D74FF8BFF89FF2CF0F76FC1C77D" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" box="[247,413,500,517]" class="Acanthodii" family="Climatiidae" genus="Nostolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="31" pageNumber="162" phylum="Chordata" rank="species" species="timanica" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8BFF89FF2CF0F76FC1C77D" box="[247,413,500,517]" italics="true" pageId="31" pageNumber="162">Nostolepis timanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8BFF89FE7FF0F76F84C77D" box="[420,472,500,517]" pageId="31" pageNumber="162" rank="species">n. sp.</taxonomicNameLabel>
|
||
, scales, crowns, anterior upwards, Pioneer Island, sample P-10-1, Lower Devonian, Member 8;
|
||
<emphasis id="E2B6EAE5FF8BFF89FF2EF31D6F5EC757" bold="true" box="[245,258,542,559]" pageId="31" pageNumber="162">A</emphasis>
|
||
, LIG 35-2022;
|
||
<emphasis id="E2B6EAE5FF8BFF89FEA6F31D6FD6C757" bold="true" box="[381,394,542,559]" pageId="31" pageNumber="162">B</emphasis>
|
||
, holotype, LIG 35-2023. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D07D36F7FF8BFF89FF4CF3896FA1C7DA" blockId="31.[151,684,650,1155]" box="[151,509,650,676]" pageId="31" pageNumber="162">BIOSTRATIGRAPHIC SIGNIFICANCE</paragraph>
|
||
<paragraph id="D07D36F7FF8BFF89FF4CF3AF6C6DC1FB" blockId="31.[151,684,650,1155]" pageId="31" pageNumber="162">
|
||
Key species of the beds with
|
||
<taxonomicName id="17C24D74FF8BFF89FE2AF3A86EB0C79B" authority="Valiukevičius, 2003" authorityName="Valiukevičius" authorityYear="2003" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="162" phylum="Chordata" rank="species" species="costatus" status="sp. nov.">
|
||
<emphasis id="E2B6EAE5FF8BFF89FE2AF3A86EB0C79B" italics="true" pageId="31" pageNumber="162">Watsonacanthus costatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F985579EFF8BFF89FF28F3CF6F72C79B" box="[243,302,716,739]" pageId="31" pageNumber="162" rank="species">n. sp.</taxonomicNameLabel>
|
||
based on acanthodian, a dispersed scale assemblage, an age analogue of the nominate acanthodian zone, marked earlier as
|
||
<taxonomicName id="17C24D74FF8BFF89FF4CF2286FBFC63B" authority="Zone" authorityName="Zone" box="[151,483,811,835]" class="Actinopterygii" genus="Watsonacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="162" phylum="Chordata" rank="species" species="oervigi">
|
||
<emphasis id="E2B6EAE5FF8BFF89FF4CF2286FFCC63B" box="[151,416,811,835]" italics="true" pageId="31" pageNumber="162">Watsonacanthus oervigi</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
can be proposed. On Severnaya Zemlya Archipelago it is attached to the Rusanov Formation and lower Al’banov Subformation. The Conodont-grounded age, based on ranges of the dominating
|
||
<taxonomicName id="17C24D74FF8BFF89FD99F2A86CFBC69B" authority="(Uyeno & Mason, 1975)" baseAuthorityName="Uyeno & Mason" baseAuthorityYear="1975" class="Conodonta" family="Polygnathidae" genus="Pandorinellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="31" pageNumber="162" phylum="Chordata" rank="species" species="expansa">
|
||
<emphasis id="E2B6EAE5FF8BFF89FD99F2A86F02C69B" italics="true" pageId="31" pageNumber="162">Pandorinellina expansa</emphasis>
|
||
(Uyeno & Mason, 1975)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="17C24D74FF8BFF89FF4CF2EF6FA2C17B" authority="exigua (Philip, 1966)" authorityName="exigua (Philip" authorityYear="1966" baseAuthorityName="Philip" baseAuthorityYear="1966" box="[151,510,1003,1027]" class="Conodonta" family="Polygnathidae" genus="Pandorinellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="31" pageNumber="162" phylum="Chordata" rank="subSpecies" species="exigua" subSpecies="exigua">
|
||
<emphasis id="E2B6EAE5FF8BFF89FF4CF2EF6EFBC17B" box="[151,167,1004,1027]" italics="true" pageId="31" pageNumber="162">P</emphasis>
|
||
.
|
||
<emphasis id="E2B6EAE5FF8BFF89FF6CF2EF6F0AC17B" box="[183,342,1004,1027]" italics="true" pageId="31" pageNumber="162">exigua exigua</emphasis>
|
||
(Philip, 1966)
|
||
</taxonomicName>
|
||
with a smaller numbers of
|
||
<taxonomicName id="17C24D74FF8BFF89FEC6F5086E91C13B" authority="Klapper & Johnson, 1980" authorityName="Klapper & Johnson" authorityYear="1980" class="Conodonta" family="Icriodontidae" genus="Pelekysgnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="31" pageNumber="162" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8BFF89FEC6F5086F98C15B" box="[285,452,1035,1059]" italics="true" pageId="31" pageNumber="162">Pelekysgnathus</emphasis>
|
||
Klapper & Johnson, 1980
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="17C24D74FF8BFF89FED3F52F6CFBC13B" authority="Uyeno & Klapper, 1980" authorityName="Uyeno & Klapper" authorityYear="1980" box="[264,679,1067,1092]" class="Conodonta" genus="Steptotaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="31" pageNumber="162" phylum="Chordata" rank="genus">
|
||
<emphasis id="E2B6EAE5FF8BFF89FED3F52F6FD8C13B" box="[264,388,1068,1091]" italics="true" pageId="31" pageNumber="162">Steptotaxis</emphasis>
|
||
<bibRefCitation id="B4534B06FF8BFF89FE57F52F6CFBC13B" author="UYENO T. T. & KLAPPER J." box="[396,679,1067,1092]" pageId="31" pageNumber="162" pagination="81 - 93" refId="ref36769" refString="UYENO T. T. & KLAPPER J. 1980. - Summary of conodont biostratigraphy of the Blue Fiord and Bird Fiord formations (Lower-Middle Devonian) at the type and adjacent areas, southwerstern Ellesmere Island, Canadian Arctic Archipelago, in Current Research. Part C. Geological Survey of Canada, paper 80: 81 - 93." type="journal article" year="1980">Uyeno & Klapper, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
, gives an Emsian age, within the
|
||
<emphasis id="E2B6EAE5FF8BFF89FE31F5486CF0C11B" box="[490,684,1099,1123]" italics="true" pageId="31" pageNumber="162">dehiscens-inversus</emphasis>
|
||
zones of the standard conodont scale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |