407 lines
36 KiB
XML
407 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.60072" ID-GBIF-Dataset="a8375122-971c-4804-92a6-fe16e10e3224" ID-PMC="PMC8222295" ID-Pensoft-Pub="1313-2970-1044-427" ID-Pensoft-UUID="8320077CEFA5571D971EFEA66B0C6268" ID-PubMed="34183882" ID-ZooBank="9AA72654B6CC4081B557CD0BAE3939A3" ModsDocID="1313-2970-1044-427" checkinTime="1623875398087" checkinUser="pensoft" docAuthor="Allegro, Gianni & Giachino, Pier Mauro" docDate="2021" docId="582C63BBE91F5B488E6973704D0409DA" docLanguage="en" docName="ZooKeys 1044: 427-448" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.60072" docTitle="Diplocheila (Diplocheila) walterrossii Allegro & Giachino 2021, sp. nov." docType="treatment" docUuid="714A6638-4FF9-4169-AC5F-E3F937247446" docUuidSource="ZooBank" docVersion="4" id="8320077CEFA5571D971EFEA66B0C6268" lastPageNumber="427" masterDocId="8320077CEFA5571D971EFEA66B0C6268" masterDocTitle="The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="448" masterPageNumber="427" pageNumber="427" updateTime="1668150474377" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Allegro, Gianni</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5079-6671</mods:nameIdentifier>
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<mods:affiliation>Strada Patro 11, I- 14036 Moncalvo (AT), Italy</mods:affiliation>
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<mods:nameIdentifier type="email">gianni.allegro54@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Giachino, Pier Mauro</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1167-5447</mods:nameIdentifier>
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<mods:affiliation>Via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>427</mods:start>
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<mods:end>448</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.60072</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-427</mods:identifier>
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||
<mods:identifier type="ZooBank">9AA72654B6CC4081B557CD0BAE3939A3</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">8320077CEFA5571D971EFEA66B0C6268</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-GBIF-Taxon="182754011" LSID="urn:lsid:zoobank.org:act:714A6638-4FF9-4169-AC5F-E3F937247446" httpUri="http://treatment.plazi.org/id/582C63BBE91F5B488E6973704D0409DA" lastPageNumber="427" pageId="0" pageNumber="427">
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<subSubSection pageId="0" pageNumber="427" type="nomenclature">
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<paragraph pageId="0" pageNumber="427">
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<taxonomicName LSID="http://zoobank.org/714A6638-4FF9-4169-AC5F-E3F937247446" authority="Allegro & Giachino, 2021" authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila) walterrossii" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="walterrossii" status="sp. nov." subGenus="Diplocheila">Diplocheila (Diplocheila) walterrossii</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="427">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diplocheila walterrossii sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure4" httpUri="https://binary.pensoft.net/fig/554095" pageId="0" pageNumber="427">Figures 4</figureCitation>
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<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">, 10</figureCitation>
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<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">, 16</figureCitation>
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<figureCitation captionStart="Figures 18–19" captionStartId="F8" captionText="Figures 18 - 19. Male protarsi of Diplocheila species 18 D. laevigata from Cambodia 19 D. walterrossii sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures18-19" httpUri="https://binary.pensoft.net/fig/554099" pageId="0" pageNumber="427">, 19</figureCitation>
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<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">, 22</figureCitation>
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<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 29</figureCitation>
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<figureCitation captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">, 33</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="type locality">
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<paragraph pageId="0" pageNumber="427">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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Cambodia, Siem Reap Province, N Siem Reap City,
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<geoCoordinate degrees="13" direction="north" minutes="26" orientation="latitude" precision="15" seconds="29" value="13.441389">13°26'29"N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="52" orientation="longitude" precision="15" seconds="25" value="103.87361">103°52'25"E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="materials_examined">
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<paragraph pageId="0" pageNumber="427">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<materialsCitation collectingDate="2018-11-13" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" latitude="13.441389" location="City" longLatPrecision="21" longitude="103.87361" specimenCount="♂" stateProvince="Siem Reap Province" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount>♂</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Siem Reap">Siem Reap Province</collectingRegion>
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, N
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<collectingRegion country="Cambodia" name="Siem Reap">Siem Reap</collectingRegion>
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:82EE76030A32525F473E44BD90BF8D02" country="Cambodia" latitude="13.441389" longLatPrecision="21" longitude="103.87361" name="City" stateProvince="Siem Reap Province">City</location>
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,
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<geoCoordinate degrees="13" direction="north" minutes="26" orientation="latitude" precision="15" seconds="29" value="13.441389">13°26'29"N</geoCoordinate>
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,
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<geoCoordinate degrees="103" direction="east" minutes="52" orientation="longitude" precision="15" seconds="25" value="103.87361">103°52'25"E</geoCoordinate>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2018-11-13">13.XI.2018</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
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and
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<collectorName>V. Kong</collectorName>
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leg. (CGi).
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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: (
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<specimenCount type="male">7 ♂♂</specimenCount>
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and
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<specimenCount type="female">3 ♀♀</specimenCount>
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);
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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<specimenCount type="male">2 ♂♂</specimenCount>
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, same data as HT
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</materialsCitation>
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;
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<materialsCitation collectingDate="2018-05-21" collectingMethod="light trap" collectorName="Rossi, Bernardi, Kong" country="Cambodia" latitude="12.237223" location="Tonle Sap Lake" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" specimenCount="2" specimenCount-male="2" stateProvince="Kampong Chhnang Province" typeStatus="paratype">
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<specimenCount type="male">2 ♂♂</specimenCount>
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang Province</collectingRegion>
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,
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<collectingMunicipality>Khom Domnatpopol</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:4BEECA77A45DDC3391DB7EF380573DDF" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" name="Tonle Sap Lake" stateProvince="Kampong Chhnang Province">Tonle Sap Lake</location>
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,
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<geoCoordinate degrees="12" direction="north" minutes="14" orientation="latitude" precision="15" seconds="14" value="12.237223">12°14'14"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="41" orientation="longitude" precision="15" seconds="15" value="104.6875">104°41'15"E</geoCoordinate>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2018-05-21">21.V.2018</collectingDate>
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,
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<collectorName>Rossi</collectorName>
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,
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<collectorName>Bernardi</collectorName>
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and
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<collectorName>Kong</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation collectingDate="2019-05-17" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" location="Tonle Sap Lake" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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<specimenCount type="female">1 ♀</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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, banks of
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:D7FF07485072ABAE72EF55DA565D3886" country="Cambodia" name="Tonle Sap Lake" stateProvince="Kampong Chhnang">Tonle Sap Lake</location>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2019-05-17">17.V.2019</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
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and
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<collectorName>V. Kong</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation collectingDate="2018-11-07" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" county="Rolea" latitude="12.225277" location="Toulkrolanh Village" longLatPrecision="21" longitude="104.66389" municipality="District" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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,
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<collectingCounty>Rolea</collectingCounty>
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<normalizedToken originalValue="B’ier">B'ier</normalizedToken>
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<collectingMunicipality>District</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:24B9720454447FD5C4D186D413844E99" country="Cambodia" county="Rolea" latitude="12.225277" longLatPrecision="21" longitude="104.66389" municipality="District" name="Toulkrolanh Village" stateProvince="Kampong Chhnang">Toulkrolanh Village</location>
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,
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<geoCoordinate degrees="12" direction="north" minutes="13" orientation="latitude" precision="15" seconds="31" value="12.225277">12°13'31"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="39" orientation="longitude" precision="15" seconds="50" value="104.66389">104°39'50"E</geoCoordinate>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2018-11-07">7.XI.2018</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
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and
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<collectorName>V. Kong</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation collectingDate="2019-10-22" collectingDateMax="2019-11-23" collectingDateMin="2019-10-22" collectingMethod="light trap" collectorName="P. Bun, W. Rossi" country="Cambodia" county="Sisophon" location="Mean Chey University" municipality="Campus" specimenCount="2" specimenCount-female="2" stateProvince="Banteay Meanchey Province" typeStatus="paratype">
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<specimenCount type="female">2 ♀♀</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Banteay Meanchey">Banteay Meanchey Province</collectingRegion>
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, near
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<collectingCounty>Sisophon</collectingCounty>
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,
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<collectingMunicipality>Campus</collectingMunicipality>
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of the
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:B6126BD009F088E62247F2BB0C5A6963" country="Cambodia" county="Sisophon" municipality="Campus" name="Mean Chey University" stateProvince="Banteay Meanchey Province">Mean Chey University</location>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2019-10-22" valueMax="2019-11-23" valueMin="2019-10-22">22.X-23.XI.2019</collectingDate>
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,
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<collectorName>P. Bun</collectorName>
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and
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<collectorName>W. Rossi</collectorName>
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leg.
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||
</materialsCitation>
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||
;
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<materialsCitation collectingDate="2019-11-29" collectingDateMax="2019-12-03" collectingDateMin="2019-11-29" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" latitude="12.279722" location="Khsam" longLatPrecision="15" longitude="104.65794" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
|
||
,
|
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<location LSID="urn:lsid:plazi:treatment:582C63BBE91F5B488E6973704D0409DA:D3F4B58394FB0CA772F845F750ED9515" country="Cambodia" latitude="12.279722" longLatPrecision="15" longitude="104.65794" name="Khsam" stateProvince="Kampong Chhnang">Khsam</location>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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,
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<geoCoordinate degrees="12" direction="north" minutes="16" orientation="latitude" precision="15" seconds="47" value="12.279722">12°16'47"N</geoCoordinate>
|
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,
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<geoCoordinate degrees="104" direction="east" minutes="39" orientation="longitude" precision="1" seconds="28.6" value="104.65794">104°39'28.6"E</geoCoordinate>
|
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,
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<collectingMethod>light trap</collectingMethod>
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||
,
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<collectingDate value="2019-11-29" valueMax="2019-12-03" valueMin="2019-11-29">29.XI-3.XII.2019</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
|
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and
|
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<collectorName>V. Kong</collectorName>
|
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leg.
|
||
</materialsCitation>
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(CAl, CCa, CGi, BMNH, MCSNG).
|
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="diagnosis">
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<paragraph pageId="0" pageNumber="427">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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A medium-sized (ABL: 15-18 mm)
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<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
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</taxonomicName>
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of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1792" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila polita" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="polita">
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<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila polita</emphasis>
|
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</taxonomicName>
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group in the subgenus
|
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<taxonomicName authority="Diplocheila" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila)" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="subGenus" subGenus="Diplocheila">Diplocheila Diplocheila</taxonomicName>
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(sensu
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
). Among the species of this group having a sexsetose labrum, it is easily distinguished from
|
||
<taxonomicName lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by the larger body size (15-18 mm vs 12-14 mm), from
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
by the more transverse pronotum (PW/PL = 1.38 vs 1.28-1.32), from
|
||
<taxonomicName lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
|
||
</taxonomicName>
|
||
by the hind angles of pronotum not protruding (externally protruding in
|
||
<taxonomicName lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
|
||
</taxonomicName>
|
||
) and from all these species by the morphology of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="description">
|
||
<paragraph pageId="0" pageNumber="427">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Habitus</emphasis>
|
||
: ABL: 15-18 mm (HT ♂ 15.6 mm). Body parallel-sided, moderately shiny, black with antennae and palpi piceous-brown (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diplocheila walterrossii sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure4" httpUri="https://binary.pensoft.net/fig/554095" pageId="0" pageNumber="427">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1044.60072.figure4" httpUri="https://binary.pensoft.net/fig/554095" pageId="0" pageNumber="427" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila walterrossii" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.: habitus of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Head</emphasis>
|
||
: almost quadrangular, robust, glabrous except for the supraorbital setae. Eyes markedly convex; a single supraorbital seta on each side. Dorsum with microsculpture not evident, only with scattered punctures visible at>100
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
magnification; frontal impressions short and superficial. Labrum symmetrically and deeply (LR = 0.72) emarginate, with six setigerous punctures on anterior margin (4 medial equidistant + 2 lateral on lobes). Clypeus trapezoid, distinctly concave anteriorly, with 1 seta at each anterolateral corner. Antennae moderately long, densely pubescent from segment 4, with terminal two articles surpassing base of pronotum; segments elongate, the second one short, as long as a half of first. Mandibles elongate, broad, approximately similar to one another (the left with apical cutting edge more concave), with scrobe well-defined and glabrous and apex blunt; terebral tooth triangular and prominent. Labial and maxillary palps fusiform, with apices narrowly truncate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Thorax</emphasis>
|
||
: pronotum smooth, with very faint isodiametric microsculpture evident at>200
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
magnification and with scattered punctures, transverse (PW/PL = 1.38), widest just above middle (Fig.
|
||
<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">10</figureCitation>
|
||
). Disk moderately convex. Sides moderately rounded in anterior half, delicately sinuate backwards. Hind angles rounded obtuse, with a postero-lateral seta. Posterior margin rectilinear between basal impressions, which are linear and markedly impressed; anterior margin with front angles nearly obsolete. A single lateral seta on each side at anterior third. Lateral bead continuous, separated from the discal area by a narrow groove, only scarcely dilated before hind angles. Medial longitudinal impression fine, nearly reaching anterior and posterior margins; anterior transversal impression absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Elytra</emphasis>
|
||
: moderately long (EL/EW = 1.65), parallel-sided, slightly convex and flattened on disk, widest at middle, with rounded shoulders and sides delicately sinuate before apex. Surface moderately shiny; microsculpture evident only at high magnification (>100
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), consisting of fine, slightly transverse meshes. Epipleura without any distinct external plicae ("uncrossed epipleura"). Intervals moderately convex, smooth; striae deeply impressed on the whole length, delicately punctate. Parascutellar stria present; scutellar setigerous pore present at base of stria 1, just before conjunction with stria 2. Basal margin complete. Discal setigerous punctures absent; umbilicate series of setigerous punctures continuous, but punctures more widely spaced at middle. Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Ventral surface</emphasis>
|
||
(thorax and abdomen)
|
||
</emphasis>
|
||
: prosternum and proepisterna glabrous and impunctate (only with very fine punctures). Metepisterna twice as long as their width at anterior side; metepimera narrow, nearly rectangular. Prosternal intercoxal process parallel-sided with blunt apex, delicately bordered. Abdominal ventrites IV-VI shiny but shagreened at sides, glabrous except one pair of subapical central setae; males with 2, females with 4 setae at apex of ventrite VII.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Legs</emphasis>
|
||
: moderately slender. Posterior face of femora with 1 seta in profemora, 2 setae in mesofemora and metafemora. Metatrochanters glabrous and as long as one-third of metafemora. Protibial antennal cleaning organ well developed, with 2 clip setae. Protibiae robust, with 6 or 7 outer apical spines; mesotibiae with a group of setae at middle of inner face; metatibiae longitudinally furrowed at inner face. Dorsal face of tarsomeres smooth. Protarsomeres 1-3 of males moderately dilated, slightly asymmetrical; meso- and metatarsomeres not dilated in both sexes; tarsomere 5 ventrally glabrous, dorsally with 2 apical setae; claws smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Male genitalia</emphasis>
|
||
: median lobe of aedeagus short and moderately swollen before apex in lateral view (Fig.
|
||
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">22</figureCitation>
|
||
); apical lamella very short and apically rounded in dorsal view (Fig.
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">29</figureCitation>
|
||
), apex thick and very slightly bent downwards in lateral view. Ostium long, in dorsal position. Right paramere elongate and subtruncate at apex; left paramere conchoid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="etymology">
|
||
<paragraph pageId="0" pageNumber="427">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">This species is named after its collector, Walter Rossi, a world-renowned specialist in entomoparasitic fungi, as a token of our esteem and as a sign of gratitude for the gift to the authors of the specimens of the new species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="distribution">
|
||
<paragraph pageId="0" pageNumber="427">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<taxonomicName genus="Geographical" lsidName="Geographical distribution" pageId="0" pageNumber="427" rank="species" species="distribution">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Geographical distribution</emphasis>
|
||
</taxonomicName>
|
||
: this species is recorded from Central and North-Western Cambodia (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">33</figureCitation>
|
||
).
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Life habits</emphasis>
|
||
: the specimens of the type series were collected by light trapping. No other data are available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="remarks">
|
||
<paragraph pageId="0" pageNumber="427">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<taxonomicName authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila walterrossii" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the
|
||
<taxonomicName lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
group (sensu
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
), as it shares the characters distinguishing this group and, at first sight, is very similar in external morphology to
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, at a deeper examination its aedeagus (Figs
|
||
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">22</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">29</figureCitation>
|
||
) reveals evident morphological differences and is easy distinguished from that of
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">20</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">26</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">27</figureCitation>
|
||
), as well as from that of
|
||
<taxonomicName lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Figs
|
||
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">23</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">30</figureCitation>
|
||
), both species sympatric and syntopic with
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in Cambodia. Moreover, the new species shows a character almost unique in this group, as far as we know, that is the less enlarged male fore tarsi 1-3. For these reasons,
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. could represent a rather isolated species in the
|
||
<taxonomicName lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
group, and its closest relatives remain uncertain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |