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<mods:title id="797E7FC9F4DB6F37A71EA83A99EE1CF7">The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with descriptions of two new species</mods:title>
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<mods:namePart id="09B182B170FE495F3539513FA5302803">Tongkerd, Piyoros</mods:namePart>
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<paragraph id="A15D947210758618D6E6FAFDB012FAA7" blockId="3.[950,1359,1299,1355]" box="[950,1359,1299,1323]" pageId="3" pageNumber="333">
<heading id="FA15231E10758618D6E6FAFDB012FAA7" bold="true" box="[950,1359,1299,1323]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="333" reason="2">
<taxonomicName id="66E2EFF110758618D6E6FAFDB012FAA7" ID-CoL="884FM" authority="(Tomlin, 1932)" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[950,1359,1299,1323]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="333" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="9396486010758618D6E6FAFDB012FAA7" bold="true" box="[950,1359,1299,1323]" pageId="3" pageNumber="333">
<emphasis id="9396486010758618D6E6FAFDB1C2FAA7" bold="true" box="[950,1183,1299,1323]" italics="true" pageId="3" pageNumber="333">Pterocyclos spaleotes</emphasis>
(
<bibRefCitation id="C573E98310758618D1FFFAFAB015FAA7" author="le Tomlin JRB" box="[1199,1352,1300,1323]" pageId="3" pageNumber="333" pagination="227 - 228" refId="ref8293" refString="le Tomlin JRB (1932) Two new species of Rhiostoma. Proceedings of the Malacological Society of London, 19: 227 - 228." type="journal article" year="1932">Tomlin, 1932</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="E9F8C7F910758618D6B8FADAB040FAC7" box="[1000,1309,1332,1355]" pageId="3" pageNumber="333" type="description">
<paragraph id="A15D947210758618D6B8FADAB040FAC7" blockId="3.[950,1359,1299,1355]" box="[1000,1309,1332,1355]" pageId="3" pageNumber="333">
(
<figureCitation id="39D988F710758618D6A0FADAB161FAC7" box="[1008,1084,1332,1355]" captionStart="Fig" captionStartId="1.[136,171,2006,2027]" captionTargetBox="[136,774,1032,1975]" captionTargetId="figure-176@1.[136,774,1032,1975]" captionTargetPageId="1" captionText="Fig. 1. Distribution map of Pterocyclos species examined in this study. The numbered locality names are detailed in Table 1. The approximate type locality of Pterocyclos subalatus, and Pterocyclos umbraticus, are indicated by locality numbers 3 and 5, respectively." figureDoi="http://doi.org/10.5281/zenodo.5458833" httpUri="https://zenodo.org/record/5458833/files/figure.png" pageId="3" pageNumber="333">Figs. 1</figureCitation>
, 
<figureCitation id="39D988F710758618D11BFADAB133FAC7" box="[1099,1134,1332,1355]" captionStart="Fig" captionStartId="2.[113,148,2006,2027]" captionTargetBox="[113,1450,778,1971]" captionTargetId="figure-151@2.[113,1451,776,1972]" captionTargetPageId="2" captionText="Fig. 2. A–C, General anatomy of Pterocyclos frednaggsi Sutcharit &amp; Panha, new species, from Gua Musang, Kelantan, Malaysia CUMZ 4944, showing: A, right side of male with testis and external penis; B, right side of female with ovary and vaginal groove; C, left side of female with lung cavity and heart. D, E, Radula morphology of: D, Pterocyclos spaleotes, topotype specimen CUMZ 4585; E, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species, paratype CUMZ 4581." figureDoi="http://doi.org/10.5281/zenodo.5458835" httpUri="https://zenodo.org/record/5458835/files/figure.png" pageId="3" pageNumber="333">2D</figureCitation>
, 
<figureCitation id="39D988F710758618D12BFADAB1EBFAC7" box="[1147,1206,1332,1355]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">
3H 
<emphasis id="9396486010758618D1CCFADAB1F4FAC7" bold="true" box="[1180,1193,1332,1355]" pageId="3" pageNumber="333">–</emphasis>
J
</figureCitation>
; 
<tableCitation id="EC60A1C910758618D193FADAB04EFAC7" box="[1219,1299,1332,1355]" captionStart="Table 1" captionStartId="5.[136,189,189,210]" captionText="Table 1. Shell size variation in Pterocyclos spp. Specimen collections and catalogue numbers are indicated in parentheses." pageId="3" pageNumber="333">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9F8C7F910758618D613FA9DB09EFA4E" pageId="3" pageNumber="333" type="reference_group">
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<treatmentCitationGroup id="81F2B35C10758618D613FA9DB09EFA4E" pageId="3" pageNumber="333">
<treatmentCitation id="2043B26310758618D613FA9DB192FA04" author="le Tomlin JRB" box="[835,1231,1395,1416]" page="227" pageId="3" pageNumber="333" year="1932">
<taxonomicName id="66E2EFF110758618D613FA9DB192FA04" ID-CoL="884FM" authority="Tomlin, 1932: 227" authorityName="Tomlin" authorityPageNumber="227" authorityYear="1932" box="[835,1231,1395,1416]" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="333" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="9396486010758618D613FA9DB151FA04" box="[835,1036,1395,1416]" italics="true" pageId="3" pageNumber="333">Rhiostoma spaleotes</emphasis>
<bibRefCitation id="C573E98310758618D145FA9DB192FA04" author="le Tomlin JRB" box="[1045,1231,1395,1416]" pageId="3" pageNumber="333" pagination="227 - 228" refId="ref8293" refString="le Tomlin JRB (1932) Two new species of Rhiostoma. Proceedings of the Malacological Society of London, 19: 227 - 228." type="journal article" year="1932">Tomlin, 1932: 227</bibRefCitation>
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, pl. 26 text figure. 
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locality: In a hole in the rock just outside the Batu Caves, near Kula Lumpur, 
<collectingRegion id="63265A9010758618D6A4FA43B116FA4E" box="[1012,1099,1453,1474]" country="Malaysia" name="Selangor" pageId="3" pageNumber="333">Selangor</collectingRegion>
. 
<treatmentCitation id="2043B26310758618D103FA43B05FFA4E" author="Laidlaw FF" box="[1107,1282,1453,1474]" page="36" pageId="3" pageNumber="333" year="1932">
<bibRefCitation id="C573E98310758618D103FA43B05FFA4E" author="Laidlaw FF" box="[1107,1282,1453,1474]" pageId="3" pageNumber="333" pagination="35 - 41" refId="ref7504" refString="Laidlaw FF (1932) Notes on the land Mollusca of the Batu Cave, Selangor, with description of two new species. Bulletin of the Raffles Museum, 7: 35 - 41." type="journal article" year="1932">Laidlaw, 1932: 36</bibRefCitation>
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. 
<treatmentCitation id="2043B26310758618D059FA43B0E2FA4E" author="Maassen WJM" box="[1289,1471,1453,1474]" page="28" pageId="3" pageNumber="333" year="2001">
<bibRefCitation id="C573E98310758618D059FA43B0E2FA4E" author="Maassen WJM" box="[1289,1471,1453,1474]" pageId="3" pageNumber="333" pagination="1 - 155" refId="ref7540" refString="Maassen WJM (2001) A preliminary checklist of the non-marine Molluscs of west Malaysia, A hand list. De Kreukel Supplement: 1 - 155." type="book chapter" year="2001">Maassen, 2001: 28</bibRefCitation>
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.
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<subSubSection id="E9F8C7F910758618D613FA08B019F972" pageId="3" pageNumber="333" type="materials_examined">
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<emphasis id="9396486010758618D613FA08B6B7FA77" bold="true" box="[835,1002,1510,1531]" pageId="3" pageNumber="333">Type specimens.</emphasis>
<materialsCitation id="118A9E2F10758618D6A0FA08B042F907" collectionCode="NMW" pageId="3" pageNumber="333" specimenCode="NMW 1955.158, NMW 1981.118" specimenCount="3" typeStatus="lectotype">
The specimen herein designated as the 
<typeStatus id="7E592AD010758618D03AFA08B09FFA77" box="[1386,1474,1510,1531]" pageId="3" pageNumber="333" type="lectotype">lectotype</typeStatus>
<specimenCode id="F1443C0910758618D613F9EDB680F994" box="[835,989,1539,1560]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="333">NMW 1955.158</specimenCode>
.01107 (height 11.4 × diameter 
<quantity id="661A399710758618D1ACF9EDB00FF994" box="[1276,1362,1539,1560]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="3" pageNumber="333" unit="mm" value="22.8">22.8 mm</quantity>
; 
<figureCitation id="39D988F710758618D00AF9EDB0FFF994" box="[1370,1442,1539,1560]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">Fig. 3H</figureCitation>
, no operculum), to stabilise the name, is the shell figured in the original description (
<bibRefCitation id="C573E98310758618D6EBF9D3B11CF9DE" author="le Tomlin JRB" box="[955,1089,1597,1618]" pageId="3" pageNumber="333" pagination="227 - 228" refId="ref8293" refString="le Tomlin JRB (1932) Two new species of Rhiostoma. Proceedings of the Malacological Society of London, 19: 227 - 228." type="journal article" year="1932">Tomlin, 1932</bibRefCitation>
, pl. 26 text figure). The other specimen from the same lot of the 
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becomes the 
<typeStatus id="7E592AD010758618D1AFF9B7B0DDF9E2" box="[1279,1408,1625,1646]" pageId="3" pageNumber="333" type="paralectotype">paralectotype</typeStatus>
<specimenCode id="F1443C0910758618D0D5F9B7B6C3F907" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="333">NMW 1981.118</specimenCode>
.02705 (1 shell, 
<figureCitation id="39D988F710758618D16DF998B1DDF907" box="[1085,1152,1654,1675]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">Fig. 3I</figureCitation>
, no operculum)
</materialsCitation>
.
</paragraph>
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<emphasis id="9396486010758618D613F95EB104F949" bold="true" box="[835,1113,1712,1733]" pageId="3" pageNumber="333">Other material examined.</emphasis>
<materialsCitation id="118A9E2F10758618D133F95EB01DF972" collectionCode="CUMZ" country="Malaysia" latitude="3.2379334" location="Topotypes" longLatPrecision="1" longitude="101.68465" municipality="Batu Cave" pageId="3" pageNumber="333" specimenCode="CUMZ 4585" specimenCount="35" stateProvince="Kuala Lumpur">
<location id="A43DC2A910758618D133F95EB197F949" LSID="urn:lsid:plazi:treatment:294B25641075861ED6E6FAFDB72CF8E3:A43DC2A910758618D133F95EB197F949" box="[1123,1226,1712,1733]" country="Malaysia" latitude="3.2379334" longLatPrecision="1" longitude="101.68465" municipality="Batu Cave" name="Topotypes" pageId="3" pageNumber="333" stateProvince="Kuala Lumpur">Topotypes</location>
from 
<collectingMunicipality id="41390E0810758618D05DF95EB027F949" box="[1293,1402,1712,1733]" pageId="3" pageNumber="333">Batu Cave</collectingMunicipality>
, 
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, 
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(
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, 
<geoCoordinate id="C4D6F2B510758618D1CDF923B073F96E" box="[1181,1326,1741,1762]" degrees="101" direction="east" minutes="41.079" orientation="longitude" pageId="3" pageNumber="333" precision="1" value="101.68465">101°41.079' E</geoCoordinate>
): 
<specimenCode id="F1443C0910758618D013F923B09FF96E" box="[1347,1474,1741,1762]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="3" pageNumber="333">CUMZ 4585</specimenCode>
(
<specimenCount id="B7E45FFB10758618D61AF907B692F972" box="[842,975,1769,1790]" pageId="3" pageNumber="333" type="generic">35 specimens</specimenCount>
in ethanol, 
<figureCitation id="39D988F710758618D116F907B1D6F972" box="[1094,1163,1769,1790]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">Fig. 3J</figureCitation>
), 4747 (52 shells)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E9F8C7F91075861ED613F8C6B72CF8E3" lastPageId="5" lastPageNumber="335" pageId="3" pageNumber="333" type="discussion">
<paragraph id="A15D94721075861ED613F8C6B655F983" blockId="3.[835,1477,1832,2111]" lastBlockId="5.[136,776,1368,1551]" lastPageId="5" lastPageNumber="335" pageId="3" pageNumber="333">
<emphasis id="9396486010758618D613F8C6B6F5F8B3" bold="true" box="[835,936,1832,1855]" pageId="3" pageNumber="333">Remark.</emphasis>
<taxonomicName id="66E2EFF110758618D6FEF8C6B1C8F8B3" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[942,1173,1832,1855]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="333" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="9396486010758618D6FEF8C6B1C8F8B3" box="[942,1173,1832,1855]" italics="true" pageId="3" pageNumber="333">Pterocyclos spaleotes</emphasis>
</taxonomicName>
is only known from its 
<typeStatus id="7E592AD010758618D0C4F8C6B09FF8B3" box="[1428,1474,1832,1855]" pageId="3" pageNumber="333">type</typeStatus>
locality, despite several surveys in nearby areas. Originally, it was described from two shells without an operculum, which leads 
<bibRefCitation id="C573E98310758618D6D5F866B17BF813" author="le Tomlin JRB" box="[901,1062,1928,1951]" pageId="3" pageNumber="333" pagination="227 - 228" refId="ref8293" refString="le Tomlin JRB (1932) Two new species of Rhiostoma. Proceedings of the Malacological Society of London, 19: 227 - 228." type="journal article" year="1932">Tomlin (1932)</bibRefCitation>
to assign it to the genus 
<taxonomicName id="66E2EFF110758618D01EF866B6BCF833" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="333" phylum="Mollusca" rank="genus">
<emphasis id="9396486010758618D01EF866B09FF813" box="[1358,1474,1928,1951]" italics="true" pageId="3" pageNumber="333">Rhiostoma</emphasis>
Benson, 1860
</taxonomicName>
. However, after examining the 
<typeStatus id="7E592AD010758618D019F846B0DEF833" box="[1353,1411,1960,1983]" pageId="3" pageNumber="333">types</typeStatus>
(
<figureCitation id="39D988F710758618D0C6F846B6D1F853" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">Fig. 3G, H</figureCitation>
) and topotypic specimens (
<figureCitation id="39D988F710758618D180F826B07FF853" box="[1232,1314,1992,2015]" captionStart="Fig" captionStartId="4.[113,148,1890,1911]" captionTargetBox="[113,1451,259,1818]" captionTargetId="figure-177@4.[113,1451,259,1819]" captionTargetPageId="4" captionText="Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit &amp; Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit &amp; Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597." figureDoi="http://doi.org/10.5281/zenodo.5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="3" pageNumber="333">Fig. 3I</figureCitation>
), this generic placement seems untenable. The unique characters of 
<taxonomicName id="66E2EFF110758618D0FEF806B6F5F793" baseAuthorityName="Tomlin" baseAuthorityYear="1932" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="333" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="9396486010758618D0FEF806B6F5F793" italics="true" pageId="3" pageNumber="333">P. spaleotes</emphasis>
</taxonomicName>
are: the small and thin shell, the corneous and transparent periostracum, and the surface with a variegated brown zigzag colour pattern and a dark brown peripheral band. The last whorl is usually partially disconnected from the penultimate whorl.Aperture rounded, with a white lip and a little expansion of the upper part. The accessory respiratory structure has a short, tubular shape. The operculum is calcareous, slightly concave inside and multilamellae outside.
</paragraph>
<caption id="F59DC4FA1072861FD521F88CB0F1F7CD" ID-DOI="http://doi.org/10.5281/zenodo.5458837" ID-Zenodo-Dep="5458837" httpUri="https://zenodo.org/record/5458837/files/figure.png" pageId="4" pageNumber="334" startId="4.[113,148,1890,1911]" targetBox="[113,1451,259,1818]" targetPageId="4">
<paragraph id="A15D94721072861FD521F88CB0F1F7CD" blockId="4.[113,1455,1890,2113]" pageId="4" pageNumber="334">
Fig. 3. A–D, Shell and operculum of 
<taxonomicName id="66E2EFF11072861FD4B5F88CB7EEF8FB" authorityName="Benson" authorityYear="1832" box="[485,691,1890,1911]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="rupestris">
<emphasis id="939648601072861FD4B5F88CB7EEF8FB" box="[485,691,1890,1911]" italics="true" pageId="4" pageNumber="334">Pterocyclos rupestris</emphasis>
</taxonomicName>
: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of 
<taxonomicName id="66E2EFF11072861FD0CAF891B5F3F83D" authorityName="Benson" authorityYear="1851" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="blandi">
<emphasis id="939648601072861FD0CAF891B5F3F83D" italics="true" pageId="4" pageNumber="334">P. blandi</emphasis>
</taxonomicName>
from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of 
<taxonomicName id="66E2EFF11072861FD6A8F872B133F83D" authorityName="Sykes" authorityYear="1903" box="[1016,1134,1948,1969]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="subalatus">
<emphasis id="939648601072861FD6A8F872B133F83D" box="[1016,1134,1948,1969]" italics="true" pageId="4" pageNumber="334">P. subalatus</emphasis>
</taxonomicName>
. H–J, 
<taxonomicName id="66E2EFF11072861FD1E2F872B0D9F83D" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[1202,1412,1948,1969]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="939648601072861FD1E2F872B0D9F83D" box="[1202,1412,1948,1969]" italics="true" pageId="4" pageNumber="334">Pterocyclos spaleotes</emphasis>
</taxonomicName>
: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of 
<taxonomicName id="66E2EFF11072861FD58BF83BB402F866" baseAuthorityName="Benthem Jutting" baseAuthorityYear="1949" box="[219,351,2005,2026]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="umbraticus">
<emphasis id="939648601072861FD58BF83BB402F866" box="[219,351,2005,2026]" italics="true" pageId="4" pageNumber="334">P. umbraticus</emphasis>
</taxonomicName>
. L–P, 
<taxonomicName id="66E2EFF11072861FD4CEF83BB67EF866" authority="Sutcharit &amp; Panha" authorityName="Sutcharit &amp; Panha" authorityYear="2014" box="[414,803,2005,2026]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="diluvium" status="sp. nov.">
<emphasis id="939648601072861FD4CEF83BB734F866" box="[414,617,2005,2026]" italics="true" pageId="4" pageNumber="334">Pterocyclos diluvium</emphasis>
Sutcharit &amp; Panha
</taxonomicName>
, 
<taxonomicNameLabel id="88A5F51B1072861FD67EF83BB6F8F866" box="[814,933,2005,2026]" pageId="4" pageNumber="334" rank="species">new species</taxonomicNameLabel>
: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, 
<taxonomicName id="66E2EFF11072861FD6B3F7E1B024F7A8" authority="Sutcharit &amp; Panha" authorityName="Sutcharit &amp; Panha" authorityYear="2014" box="[995,1401,2063,2084]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="334" phylum="Mollusca" rank="species" species="frednaggsi" status="sp. nov.">
<emphasis id="939648601072861FD6B3F7E1B19DF7A8" box="[995,1216,2063,2084]" italics="true" pageId="4" pageNumber="334">Pterocyclos frednaggsi</emphasis>
Sutcharit &amp; Panha
</taxonomicName>
, 
<taxonomicNameLabel id="88A5F51B1072861FD0D3F7E1B5E7F7CD" pageId="4" pageNumber="334" rank="species">new species</taxonomicNameLabel>
: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597.
</paragraph>
</caption>
<caption id="F59DC4FA1073861ED5D8FF53B064FF5E" box="[136,1337,189,210]" pageId="5" pageNumber="335" startId="5.[136,189,189,210]">
<paragraph id="A15D94721073861ED5D8FF53B064FF5E" blockId="5.[136,1337,189,210]" box="[136,1337,189,210]" pageId="5" pageNumber="335">
Table 1. Shell size variation in 
<taxonomicName id="66E2EFF11073861ED4ECFF53B700FF5E" box="[444,605,189,210]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="939648601073861ED4ECFF53B771FF5E" box="[444,556,189,210]" italics="true" pageId="5" pageNumber="335">Pterocyclos</emphasis>
spp.
</taxonomicName>
Specimen collections and catalogue numbers are indicated in parentheses.
</paragraph>
</caption>
<paragraph id="A15D94721073861ED5D8FEE9B1A5FA99" pageId="5" pageNumber="335">
<table id="D3E266D2107379E4D5D8FF19B0EDFA99" box="[136,1456,247,1301]" gridcols="6" gridrows="33" pageId="5" pageNumber="335">
<tr id="1FD29630107379E4D5D8FF19B0EDFEBC" box="[136,1456,247,304]" gridrow="0" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FF19B74CFEBC" box="[136,529,247,304]" gridcol="0" gridrow="0" pageId="5" pageNumber="335">
<emphasis id="939648601073861ED5D8FEE9B400FE90" bold="true" box="[136,349,263,284]" pageId="5" pageNumber="335">Species and Locality</emphasis>
(CUMZ nos.)
</th>
<th id="5C03FF4C107379E4D765FF19B7FCFEBC" box="[565,673,247,304]" gridcol="1" gridrow="0" pageId="5" pageNumber="335">
<emphasis id="939648601073861ED71BFF12B7FCFEA7" bold="true" pageId="5" pageNumber="335">No. of Specimens</emphasis>
</th>
<th id="5C03FF4C107379E4D7BDFF19B1A5FEBC" box="[749,1272,247,304]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="5" pageNumber="335">
<emphasis id="939648601073861ED676FF19B1A8FEBC" bold="true" pageId="5" pageNumber="335">Ranges, Mean ± S.D. (mm) of the Shell Height (H) Diameter (D) H/D Ratio</emphasis>
</th>
<th id="5C03FF4C107379E4D010FF19B0EDFEBC" box="[1344,1456,247,304]" gridcol="5" gridrow="0" pageId="5" pageNumber="335">
<emphasis id="939648601073861ED010FF12B0C3FEA7" bold="true" pageId="5" pageNumber="335">Number of Whorls</emphasis>
</th>
</tr>
<tr id="1FD29630107379E4D5D8FEADB0EDFED4" box="[136,1456,323,344]" gridrow="1" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FEADB0EDFED4" box="[136,1456,323,344]" colspan="6" colspanRight="5" gridcol="0" gridrow="1" pageId="5" pageNumber="335">
<taxonomicName id="66E2EFF11073861ED5D8FEADB460FED4" authorityName="Benson" authorityYear="1851" box="[136,317,323,344]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="blandi">
<emphasis id="939648601073861ED5D8FEADB460FED4" box="[136,317,323,344]" italics="true" pageId="5" pageNumber="335">Pterocyclos blandi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="1FD29630107379E4D5D8FE84B0EDFEF3" box="[136,1456,362,383]" gridrow="2" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FE84B74CFEF3" box="[136,529,362,383]" gridcol="0" gridrow="2" pageId="5" pageNumber="335">1. Pulau Langkawi, Malaysia</th>
<td id="5C03FF4C107379E4D765FE84B7FCFEF3" box="[565,673,362,383]" gridcol="1" gridrow="2" pageId="5" pageNumber="335">180</td>
<td id="5C03FF4C107379E4D7BDFE84B603FEF3" box="[749,862,362,383]" gridcol="2" gridrow="2" pageId="5" pageNumber="335">8.9–13.2</td>
<td id="5C03FF4C107379E4D6EAFE84B176FEF3" box="[954,1067,362,383]" gridcol="3" gridrow="2" pageId="5" pageNumber="335">19.3–25.4</td>
<td id="5C03FF4C107379E4D1D8FE84B1A5FEF3" box="[1160,1272,362,383]" gridcol="4" gridrow="2" pageId="5" pageNumber="335">0.42–0.57</td>
<td id="5C03FF4C107379E4D010FE84B0EDFEF3" box="[1344,1456,362,383]" gridcol="5" gridrow="2" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FE69B0EDFE10" box="[136,1456,391,412]" gridrow="3" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FE69B74CFE10" box="[136,529,391,412]" gridcol="0" gridrow="3" pageId="5" pageNumber="335">(3880, 4582, 4583, 4584)</th>
<td id="5C03FF4C107379E4D7BDFE69B603FE10" box="[749,862,391,412]" gridcol="2" gridrow="3" pageId="5" pageNumber="335">11.0 ± 0.84</td>
<td id="5C03FF4C107379E4D6EAFE69B176FE10" box="[954,1067,391,412]" gridcol="3" gridrow="3" pageId="5" pageNumber="335">22.2 ± 1.28</td>
<td id="5C03FF4C107379E4D1D8FE69B1A5FE10" box="[1160,1272,391,412]" gridcol="4" gridrow="3" pageId="5" pageNumber="335">0.50 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FE4AB0EDFE35" box="[136,1456,420,441]" gridrow="4" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FE4AB74CFE35" box="[136,529,420,441]" gridcol="0" gridrow="4" pageId="5" pageNumber="335">2. Gua Cerita, Pulau Langkawi,</th>
<td id="5C03FF4C107379E4D765FE4AB7FCFE35" box="[565,673,420,441]" gridcol="1" gridrow="4" pageId="5" pageNumber="335">9</td>
<td id="5C03FF4C107379E4D7BDFE4AB603FE35" box="[749,862,420,441]" gridcol="2" gridrow="4" pageId="5" pageNumber="335">8.9–10.6</td>
<td id="5C03FF4C107379E4D6EAFE4AB176FE35" box="[954,1067,420,441]" gridcol="3" gridrow="4" pageId="5" pageNumber="335">18.9–25.6</td>
<td id="5C03FF4C107379E4D1D8FE4AB1A5FE35" box="[1160,1272,420,441]" gridcol="4" gridrow="4" pageId="5" pageNumber="335">0.40–0.47</td>
<td id="5C03FF4C107379E4D010FE4AB0EDFE35" box="[1344,1456,420,441]" gridcol="5" gridrow="4" pageId="5" pageNumber="335">4¾–5</td>
</tr>
<tr id="1FD29630107379E4D5D8FE2EB0EDFE59" box="[136,1456,448,469]" gridrow="5" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FE2EB74CFE59" box="[136,529,448,469]" gridcol="0" gridrow="5" pageId="5" pageNumber="335">Malaysia (3879)</th>
<td id="5C03FF4C107379E4D7BDFE2EB603FE59" box="[749,862,448,469]" gridcol="2" gridrow="5" pageId="5" pageNumber="335">9.7 ± 0.67</td>
<td id="5C03FF4C107379E4D6EAFE2EB176FE59" box="[954,1067,448,469]" gridcol="3" gridrow="5" pageId="5" pageNumber="335">22.4 ± 1.85</td>
<td id="5C03FF4C107379E4D1D8FE2EB1A5FE59" box="[1160,1272,448,469]" gridcol="4" gridrow="5" pageId="5" pageNumber="335">0.44 ± 0.02</td>
</tr>
<tr id="1FD29630107379E4D5D8FE06B0EDFE71" box="[136,1456,488,509]" gridrow="6" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FE06B0EDFE71" box="[136,1456,488,509]" colspan="6" colspanRight="5" gridcol="0" gridrow="6" pageId="5" pageNumber="335">
<taxonomicName id="66E2EFF11073861ED5D8FE06B404FE71" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[136,345,488,509]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="939648601073861ED5D8FE06B404FE71" box="[136,345,488,509]" italics="true" pageId="5" pageNumber="335">Pterocyclos spaleotes</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="1FD29630107379E4D5D8FDE2B0EDFDAD" box="[136,1456,524,545]" gridrow="7" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FDE2B74CFDAD" box="[136,529,524,545]" gridcol="0" gridrow="7" pageId="5" pageNumber="335">4. Batu Cave, Kuala Lumpur,</th>
<td id="5C03FF4C107379E4D765FDE2B7FCFDAD" box="[565,673,524,545]" gridcol="1" gridrow="7" pageId="5" pageNumber="335">71</td>
<td id="5C03FF4C107379E4D7BDFDE2B603FDAD" box="[749,862,524,545]" gridcol="2" gridrow="7" pageId="5" pageNumber="335">8.3–12.1</td>
<td id="5C03FF4C107379E4D6EAFDE2B176FDAD" box="[954,1067,524,545]" gridcol="3" gridrow="7" pageId="5" pageNumber="335">15.6–20.7</td>
<td id="5C03FF4C107379E4D1D8FDE2B1A5FDAD" box="[1160,1272,524,545]" gridcol="4" gridrow="7" pageId="5" pageNumber="335">0.48–0.68</td>
<td id="5C03FF4C107379E4D010FDE2B0EDFDAD" box="[1344,1456,524,545]" gridcol="5" gridrow="7" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FDC7B0EDFDB2" box="[136,1456,553,574]" gridrow="8" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FDC7B74CFDB2" box="[136,529,553,574]" gridcol="0" gridrow="8" pageId="5" pageNumber="335">Malaysia (4585, 4747)</th>
<td id="5C03FF4C107379E4D7BDFDC7B603FDB2" box="[749,862,553,574]" gridcol="2" gridrow="8" pageId="5" pageNumber="335">9.7 ± 0.92</td>
<td id="5C03FF4C107379E4D6EAFDC7B176FDB2" box="[954,1067,553,574]" gridcol="3" gridrow="8" pageId="5" pageNumber="335">17.8 ± 1.16</td>
<td id="5C03FF4C107379E4D1D8FDC7B1A5FDB2" box="[1160,1272,553,574]" gridcol="4" gridrow="8" pageId="5" pageNumber="335">0.55 ± 0.04</td>
</tr>
<tr id="1FD29630107379E4D5D8FDBEB0EDFDE9" box="[136,1456,592,613]" gridrow="9" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FDBEB0EDFDE9" box="[136,1456,592,613]" colspan="6" colspanRight="5" gridcol="0" gridrow="9" pageId="5" pageNumber="335">
<taxonomicName id="66E2EFF11073861ED5D8FDBEB74FFDE9" authority="Sutcharit &amp; Panha" authorityName="Sutcharit &amp; Panha" authorityYear="2014" box="[136,530,592,613]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="diluvium" status="sp. nov.">
<emphasis id="939648601073861ED5D8FDBEB409FDE9" box="[136,340,592,613]" italics="true" pageId="5" pageNumber="335">Pterocyclos diluvium</emphasis>
Sutcharit &amp; Panha
</taxonomicName>
, 
<taxonomicNameLabel id="88A5F51B1073861ED74DFDBEB7CFFDE9" box="[541,658,592,613]" pageId="5" pageNumber="335" rank="species">new species</taxonomicNameLabel>
</th>
</tr>
<tr id="1FD29630107379E4D5D8FD9BB0EDFD06" box="[136,1456,629,650]" gridrow="10" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FD9BB74CFD06" box="[136,529,629,650]" gridcol="0" gridrow="10" pageId="5" pageNumber="335">6. Tam Sumano, Patthalung,</th>
<td id="5C03FF4C107379E4D765FD9BB7FCFD06" box="[565,673,629,650]" gridcol="1" gridrow="10" pageId="5" pageNumber="335">13</td>
<td id="5C03FF4C107379E4D7BDFD9BB603FD06" box="[749,862,629,650]" gridcol="2" gridrow="10" pageId="5" pageNumber="335">11.8–14.4</td>
<td id="5C03FF4C107379E4D6EAFD9BB176FD06" box="[954,1067,629,650]" gridcol="3" gridrow="10" pageId="5" pageNumber="335">21.8–25.2</td>
<td id="5C03FF4C107379E4D1D8FD9BB1A5FD06" box="[1160,1272,629,650]" gridcol="4" gridrow="10" pageId="5" pageNumber="335">0.52–0.61</td>
<td id="5C03FF4C107379E4D010FD9BB0EDFD06" box="[1344,1456,629,650]" gridcol="5" gridrow="10" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FD7FB0EDFD2A" box="[136,1456,657,678]" gridrow="11" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FD7FB74CFD2A" box="[136,529,657,678]" gridcol="0" gridrow="11" pageId="5" pageNumber="335">Thailand (4588)</th>
<td id="5C03FF4C107379E4D7BDFD7FB603FD2A" box="[749,862,657,678]" gridcol="2" gridrow="11" pageId="5" pageNumber="335">13.3 ± 0.74</td>
<td id="5C03FF4C107379E4D6EAFD7FB176FD2A" box="[954,1067,657,678]" gridcol="3" gridrow="11" pageId="5" pageNumber="335">23.8 ± 1.06</td>
<td id="5C03FF4C107379E4D1D8FD7FB1A5FD2A" box="[1160,1272,657,678]" gridcol="4" gridrow="11" pageId="5" pageNumber="335">0.56 ± 0.02</td>
</tr>
<tr id="1FD29630107379E4D5D8FD40B0EDFD4F" box="[136,1456,686,707]" gridrow="12" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FD40B74CFD4F" box="[136,529,686,707]" gridcol="0" gridrow="12" pageId="5" pageNumber="335">7. Tam Puttha Kodome, Patthalung,</th>
<td id="5C03FF4C107379E4D765FD40B7FCFD4F" box="[565,673,686,707]" gridcol="1" gridrow="12" pageId="5" pageNumber="335">9</td>
<td id="5C03FF4C107379E4D7BDFD40B603FD4F" box="[749,862,686,707]" gridcol="2" gridrow="12" pageId="5" pageNumber="335">12.0–15.6</td>
<td id="5C03FF4C107379E4D6EAFD40B176FD4F" box="[954,1067,686,707]" gridcol="3" gridrow="12" pageId="5" pageNumber="335">20.1–25.0</td>
<td id="5C03FF4C107379E4D1D8FD40B1A5FD4F" box="[1160,1272,686,707]" gridcol="4" gridrow="12" pageId="5" pageNumber="335">0.57–0.62</td>
<td id="5C03FF4C107379E4D010FD40B0EDFD4F" box="[1344,1456,686,707]" gridcol="5" gridrow="12" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FD25B0EDFD6C" box="[136,1456,715,736]" gridrow="13" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FD25B74CFD6C" box="[136,529,715,736]" gridcol="0" gridrow="13" pageId="5" pageNumber="335">Thailand (3812)</th>
<td id="5C03FF4C107379E4D7BDFD25B603FD6C" box="[749,862,715,736]" gridcol="2" gridrow="13" pageId="5" pageNumber="335">13.2 ± 1.18</td>
<td id="5C03FF4C107379E4D6EAFD25B176FD6C" box="[954,1067,715,736]" gridcol="3" gridrow="13" pageId="5" pageNumber="335">22.5 ± 1.65</td>
<td id="5C03FF4C107379E4D1D8FD25B1A5FD6C" box="[1160,1272,715,736]" gridcol="4" gridrow="13" pageId="5" pageNumber="335">0.59 ± 0.02</td>
</tr>
<tr id="1FD29630107379E4D5D8FD06B0EDFD71" box="[136,1456,744,765]" gridrow="14" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FD06B74CFD71" box="[136,529,744,765]" gridcol="0" gridrow="14" pageId="5" pageNumber="335">8. Tam Phaya Hong, Kong Ra,</th>
<td id="5C03FF4C107379E4D765FD06B7FCFD71" box="[565,673,744,765]" gridcol="1" gridrow="14" pageId="5" pageNumber="335">11</td>
<td id="5C03FF4C107379E4D7BDFD06B603FD71" box="[749,862,744,765]" gridcol="2" gridrow="14" pageId="5" pageNumber="335">11.3–15.0</td>
<td id="5C03FF4C107379E4D6EAFD06B176FD71" box="[954,1067,744,765]" gridcol="3" gridrow="14" pageId="5" pageNumber="335">22.9–27.1</td>
<td id="5C03FF4C107379E4D1D8FD06B1A5FD71" box="[1160,1272,744,765]" gridcol="4" gridrow="14" pageId="5" pageNumber="335">0.47–0.57</td>
<td id="5C03FF4C107379E4D010FD06B0EDFD71" box="[1344,1456,744,765]" gridcol="5" gridrow="14" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FCEBB0EDFC96" box="[136,1456,773,794]" gridrow="15" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FCEBB74CFC96" box="[136,529,773,794]" gridcol="0" gridrow="15" pageId="5" pageNumber="335">Patthalung, Thailand (4870)</th>
<td id="5C03FF4C107379E4D7BDFCEBB603FC96" box="[749,862,773,794]" gridcol="2" gridrow="15" pageId="5" pageNumber="335">13.1 ± 1.16</td>
<td id="5C03FF4C107379E4D6EAFCEBB176FC96" box="[954,1067,773,794]" gridcol="3" gridrow="15" pageId="5" pageNumber="335">25.2 ± 1.54</td>
<td id="5C03FF4C107379E4D1D8FCEBB1A5FC96" box="[1160,1272,773,794]" gridcol="4" gridrow="15" pageId="5" pageNumber="335">0.52 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FCCFB0EDFCBA" box="[136,1456,801,822]" gridrow="16" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FCCFB74CFCBA" box="[136,529,801,822]" gridcol="0" gridrow="16" pageId="5" pageNumber="335">9. Tarutao National Park, Satun,</th>
<td id="5C03FF4C107379E4D765FCCFB7FCFCBA" box="[565,673,801,822]" gridcol="1" gridrow="16" pageId="5" pageNumber="335">54</td>
<td id="5C03FF4C107379E4D7BDFCCFB603FCBA" box="[749,862,801,822]" gridcol="2" gridrow="16" pageId="5" pageNumber="335">9.8–14.4</td>
<td id="5C03FF4C107379E4D6EAFCCFB176FCBA" box="[954,1067,801,822]" gridcol="3" gridrow="16" pageId="5" pageNumber="335">18.6–27.2</td>
<td id="5C03FF4C107379E4D1D8FCCFB1A5FCBA" box="[1160,1272,801,822]" gridcol="4" gridrow="16" pageId="5" pageNumber="335">0.49–0.61</td>
<td id="5C03FF4C107379E4D010FCCFB0EDFCBA" box="[1344,1456,801,822]" gridcol="5" gridrow="16" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FCD0B0EDFCDF" box="[136,1456,830,851]" gridrow="17" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FCD0B74CFCDF" box="[136,529,830,851]" gridcol="0" gridrow="17" pageId="5" pageNumber="335">Thailand (4589, 4593, 4749, 4750)</th>
<td id="5C03FF4C107379E4D7BDFCD0B603FCDF" box="[749,862,830,851]" gridcol="2" gridrow="17" pageId="5" pageNumber="335">11.9 ± 0.95</td>
<td id="5C03FF4C107379E4D6EAFCD0B176FCDF" box="[954,1067,830,851]" gridcol="3" gridrow="17" pageId="5" pageNumber="335">21.7 ± 1.50</td>
<td id="5C03FF4C107379E4D1D8FCD0B1A5FCDF" box="[1160,1272,830,851]" gridcol="4" gridrow="17" pageId="5" pageNumber="335">0.55 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FCB5B0EDFCFC" box="[136,1456,859,880]" gridrow="18" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FCB5B74CFCFC" box="[136,529,859,880]" gridcol="0" gridrow="18" pageId="5" pageNumber="335">10. Tam Tone-din, Kuan-Don, Satun,</th>
<td id="5C03FF4C107379E4D765FCB5B7FCFCFC" box="[565,673,859,880]" gridcol="1" gridrow="18" pageId="5" pageNumber="335">41</td>
<td id="5C03FF4C107379E4D7BDFCB5B603FCFC" box="[749,862,859,880]" gridcol="2" gridrow="18" pageId="5" pageNumber="335">13.0–16.6</td>
<td id="5C03FF4C107379E4D6EAFCB5B176FCFC" box="[954,1067,859,880]" gridcol="3" gridrow="18" pageId="5" pageNumber="335">26.1–31.5</td>
<td id="5C03FF4C107379E4D1D8FCB5B1A5FCFC" box="[1160,1272,859,880]" gridcol="4" gridrow="18" pageId="5" pageNumber="335">0.43–0.55</td>
<td id="5C03FF4C107379E4D010FCB5B0EDFCFC" box="[1344,1456,859,880]" gridcol="5" gridrow="18" pageId="5" pageNumber="335">4½–5¾</td>
</tr>
<tr id="1FD29630107379E4D5D8FC96B0EDFC01" box="[136,1456,888,909]" gridrow="19" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FC96B74CFC01" box="[136,529,888,909]" gridcol="0" gridrow="19" pageId="5" pageNumber="335">Thailand (4590, 4591, 4866)</th>
<td id="5C03FF4C107379E4D7BDFC96B603FC01" box="[749,862,888,909]" gridcol="2" gridrow="19" pageId="5" pageNumber="335">14.5 ± 0.88</td>
<td id="5C03FF4C107379E4D6EAFC96B176FC01" box="[954,1067,888,909]" gridcol="3" gridrow="19" pageId="5" pageNumber="335">28.0 ± 1.42</td>
<td id="5C03FF4C107379E4D1D8FC96B1A5FC01" box="[1160,1272,888,909]" gridcol="4" gridrow="19" pageId="5" pageNumber="335">0.50 ± 0.04</td>
</tr>
<tr id="1FD29630107379E4D5D8FC7BB0EDFC26" box="[136,1456,917,938]" gridrow="20" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FC7BB74CFC26" box="[136,529,917,938]" gridcol="0" gridrow="20" pageId="5" pageNumber="335">11. Gua Kelam, Perlis,</th>
<td id="5C03FF4C107379E4D765FC7BB7FCFC26" box="[565,673,917,938]" gridcol="1" gridrow="20" pageId="5" pageNumber="335">17</td>
<td id="5C03FF4C107379E4D7BDFC7BB603FC26" box="[749,862,917,938]" gridcol="2" gridrow="20" pageId="5" pageNumber="335">10.7–13.3</td>
<td id="5C03FF4C107379E4D6EAFC7BB176FC26" box="[954,1067,917,938]" gridcol="3" gridrow="20" pageId="5" pageNumber="335">20.0–26.7</td>
<td id="5C03FF4C107379E4D1D8FC7BB1A5FC26" box="[1160,1272,917,938]" gridcol="4" gridrow="20" pageId="5" pageNumber="335">0.45–0.59</td>
<td id="5C03FF4C107379E4D010FC7BB0EDFC26" box="[1344,1456,917,938]" gridcol="5" gridrow="20" pageId="5" pageNumber="335">4–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FC5FB0EDFC4A" box="[136,1456,945,966]" gridrow="21" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FC5FB74CFC4A" box="[136,529,945,966]" gridcol="0" gridrow="21" pageId="5" pageNumber="335">Malaysia (3877, 4587)</th>
<td id="5C03FF4C107379E4D7BDFC5FB603FC4A" box="[749,862,945,966]" gridcol="2" gridrow="21" pageId="5" pageNumber="335">11.7 ± 0.70</td>
<td id="5C03FF4C107379E4D6EAFC5FB176FC4A" box="[954,1067,945,966]" gridcol="3" gridrow="21" pageId="5" pageNumber="335">22.6 ± 1.96</td>
<td id="5C03FF4C107379E4D1D8FC5FB1A5FC4A" box="[1160,1272,945,966]" gridcol="4" gridrow="21" pageId="5" pageNumber="335">0.52 ± 0.04</td>
</tr>
<tr id="1FD29630107379E4D5D8FC20B0EDFC6F" box="[136,1456,974,995]" gridrow="22" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FC20B74CFC6F" box="[136,529,974,995]" gridcol="0" gridrow="22" pageId="5" pageNumber="335">12. Perlis State Park, Malaysia (4586)</th>
<td id="5C03FF4C107379E4D765FC20B7FCFC6F" box="[565,673,974,995]" gridcol="1" gridrow="22" pageId="5" pageNumber="335">7</td>
<td id="5C03FF4C107379E4D7BDFC20B603FC6F" box="[749,862,974,995]" gridcol="2" gridrow="22" pageId="5" pageNumber="335">12.6–15.0</td>
<td id="5C03FF4C107379E4D6EAFC20B176FC6F" box="[954,1067,974,995]" gridcol="3" gridrow="22" pageId="5" pageNumber="335">24.4–28.3</td>
<td id="5C03FF4C107379E4D1D8FC20B1A5FC6F" box="[1160,1272,974,995]" gridcol="4" gridrow="22" pageId="5" pageNumber="335">0.49–0.61</td>
<td id="5C03FF4C107379E4D010FC20B0EDFC6F" box="[1344,1456,974,995]" gridcol="5" gridrow="22" pageId="5" pageNumber="335">4½–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FC05B0EDFB8C" box="[136,1456,1003,1024]" gridrow="23" pageId="5" pageNumber="335" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="5C03FF4C107379E4D7BDFC05B603FB8C" box="[749,862,1003,1024]" gridcol="2" gridrow="23" pageId="5" pageNumber="335">13.8 ± 0.97</td>
<td id="5C03FF4C107379E4D6EAFC05B176FB8C" box="[954,1067,1003,1024]" gridcol="3" gridrow="23" pageId="5" pageNumber="335">26.0 ± 1.30</td>
<td id="5C03FF4C107379E4D1D8FC05B1A5FB8C" box="[1160,1272,1003,1024]" gridcol="4" gridrow="23" pageId="5" pageNumber="335">0.53 ± 0.04</td>
</tr>
<tr id="1FD29630107379E4D5D8FBE6B0EDFB91" box="[136,1456,1032,1053]" gridrow="24" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FBE6B74CFB91" box="[136,529,1032,1053]" gridcol="0" gridrow="24" pageId="5" pageNumber="335">13. Sungi Jenia, Perlis, Malaysia (3878)</th>
<td id="5C03FF4C107379E4D765FBE6B7FCFB91" box="[565,673,1032,1053]" gridcol="1" gridrow="24" pageId="5" pageNumber="335">26</td>
<td id="5C03FF4C107379E4D7BDFBE6B603FB91" box="[749,862,1032,1053]" gridcol="2" gridrow="24" pageId="5" pageNumber="335">9.9–14.2</td>
<td id="5C03FF4C107379E4D6EAFBE6B176FB91" box="[954,1067,1032,1053]" gridcol="3" gridrow="24" pageId="5" pageNumber="335">18.7–24.0</td>
<td id="5C03FF4C107379E4D1D8FBE6B1A5FB91" box="[1160,1272,1032,1053]" gridcol="4" gridrow="24" pageId="5" pageNumber="335">0.47–0.62</td>
<td id="5C03FF4C107379E4D010FBE6B0EDFB91" box="[1344,1456,1032,1053]" gridcol="5" gridrow="24" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FBCBB0EDFBB6" box="[136,1456,1061,1082]" gridrow="25" pageId="5" pageNumber="335" rowspan-0="1" rowspan-1="1" rowspan-5="1">
<td id="5C03FF4C107379E4D7BDFBCBB603FBB6" box="[749,862,1061,1082]" gridcol="2" gridrow="25" pageId="5" pageNumber="335">11.9 ± 0.99</td>
<td id="5C03FF4C107379E4D6EAFBCBB176FBB6" box="[954,1067,1061,1082]" gridcol="3" gridrow="25" pageId="5" pageNumber="335">21.9 ± 1.49</td>
<td id="5C03FF4C107379E4D1D8FBCBB1A5FBB6" box="[1160,1272,1061,1082]" gridcol="4" gridrow="25" pageId="5" pageNumber="335">0.54 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FBAFB0EDFBDA" box="[136,1456,1089,1110]" gridrow="26" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FBAFB74CFBDA" box="[136,529,1089,1110]" gridcol="0" gridrow="26" pageId="5" pageNumber="335">14. Gua Cenderawasih, Perlis,</th>
<td id="5C03FF4C107379E4D765FBAFB7FCFBDA" box="[565,673,1089,1110]" gridcol="1" gridrow="26" pageId="5" pageNumber="335">29</td>
<td id="5C03FF4C107379E4D7BDFBAFB603FBDA" box="[749,862,1089,1110]" gridcol="2" gridrow="26" pageId="5" pageNumber="335">11.5–15.6</td>
<td id="5C03FF4C107379E4D6EAFBAFB176FBDA" box="[954,1067,1089,1110]" gridcol="3" gridrow="26" pageId="5" pageNumber="335">21.0–27.5</td>
<td id="5C03FF4C107379E4D1D8FBAFB1A5FBDA" box="[1160,1272,1089,1110]" gridcol="4" gridrow="26" pageId="5" pageNumber="335">0.49–0.61</td>
<td id="5C03FF4C107379E4D010FBAFB0EDFBDA" box="[1344,1456,1089,1110]" gridcol="5" gridrow="26" pageId="5" pageNumber="335">4¾–5½</td>
</tr>
<tr id="1FD29630107379E4D5D8FBB0B0EDFBFF" box="[136,1456,1118,1139]" gridrow="27" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FBB0B74CFBFF" box="[136,529,1118,1139]" gridcol="0" gridrow="27" pageId="5" pageNumber="335">Malaysia (3881, 4592)</th>
<td id="5C03FF4C107379E4D7BDFBB0B603FBFF" box="[749,862,1118,1139]" gridcol="2" gridrow="27" pageId="5" pageNumber="335">13.3 ± 1.25</td>
<td id="5C03FF4C107379E4D6EAFBB0B176FBFF" box="[954,1067,1118,1139]" gridcol="3" gridrow="27" pageId="5" pageNumber="335">24.3 ± 1.51</td>
<td id="5C03FF4C107379E4D1D8FBB0B1A5FBFF" box="[1160,1272,1118,1139]" gridcol="4" gridrow="27" pageId="5" pageNumber="335">0.55 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FB68B0EDFB17" box="[136,1456,1158,1179]" gridrow="28" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FB68B0EDFB17" box="[136,1456,1158,1179]" colspan="6" colspanRight="5" gridcol="0" gridrow="28" pageId="5" pageNumber="335">
<taxonomicName id="66E2EFF11073861ED5D8FB68B779FB17" authority="Sutcharit &amp; Panha" authorityName="Sutcharit &amp; Panha" authorityYear="2014" box="[136,548,1158,1179]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="frednaggsi" status="sp. nov.">
<emphasis id="939648601073861ED5D8FB68B43BFB17" box="[136,358,1158,1179]" italics="true" pageId="5" pageNumber="335">Pterocyclos frednaggsi</emphasis>
Sutcharit &amp; Panha
</taxonomicName>
, 
<taxonomicNameLabel id="88A5F51B1073861ED77FFB68B7F9FB17" box="[559,676,1158,1179]" pageId="5" pageNumber="335" rank="species">new species</taxonomicNameLabel>
</th>
</tr>
<tr id="1FD29630107379E4D5D8FB44B0EDFB33" box="[136,1456,1194,1215]" gridrow="29" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FB44B74CFB33" box="[136,529,1194,1215]" gridcol="0" gridrow="29" pageId="5" pageNumber="335">15. Bukit Chintamanis, Pahang,</th>
<td id="5C03FF4C107379E4D765FB44B7FCFB33" box="[565,673,1194,1215]" gridcol="1" gridrow="29" pageId="5" pageNumber="335">47</td>
<td id="5C03FF4C107379E4D7BDFB44B603FB33" box="[749,862,1194,1215]" gridcol="2" gridrow="29" pageId="5" pageNumber="335">10.8–14.8</td>
<td id="5C03FF4C107379E4D6EAFB44B176FB33" box="[954,1067,1194,1215]" gridcol="3" gridrow="29" pageId="5" pageNumber="335">21.8–32.0</td>
<td id="5C03FF4C107379E4D1D8FB44B1A5FB33" box="[1160,1272,1194,1215]" gridcol="4" gridrow="29" pageId="5" pageNumber="335">0.43–0.60</td>
<td id="5C03FF4C107379E4D010FB44B0EDFB33" box="[1344,1456,1194,1215]" gridcol="5" gridrow="29" pageId="5" pageNumber="335">4½–5¼</td>
</tr>
<tr id="1FD29630107379E4D5D8FB29B0EDFB50" box="[136,1456,1223,1244]" gridrow="30" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FB29B74CFB50" box="[136,529,1223,1244]" gridcol="0" gridrow="30" pageId="5" pageNumber="335">Malaysia (4581, 4571)</th>
<td id="5C03FF4C107379E4D7BDFB29B603FB50" box="[749,862,1223,1244]" gridcol="2" gridrow="30" pageId="5" pageNumber="335">13.1 ± 0.78</td>
<td id="5C03FF4C107379E4D6EAFB29B176FB50" box="[954,1067,1223,1244]" gridcol="3" gridrow="30" pageId="5" pageNumber="335">25.6 ± 1.64</td>
<td id="5C03FF4C107379E4D1D8FB29B1A5FB50" box="[1160,1272,1223,1244]" gridcol="4" gridrow="30" pageId="5" pageNumber="335">0.51 ± 0.03</td>
</tr>
<tr id="1FD29630107379E4D5D8FB0AB0EDFB75" box="[136,1456,1252,1273]" gridrow="31" pageId="5" pageNumber="335">
<th id="5C03FF4C107379E4D5D8FB0AB74CFB75" box="[136,529,1252,1273]" gridcol="0" gridrow="31" pageId="5" pageNumber="335">16. Gua Pulai, Gua Musang, Kelanta,</th>
<td id="5C03FF4C107379E4D765FB0AB7FCFB75" box="[565,673,1252,1273]" gridcol="1" gridrow="31" pageId="5" pageNumber="335">9</td>
<td id="5C03FF4C107379E4D7BDFB0AB603FB75" box="[749,862,1252,1273]" gridcol="2" gridrow="31" pageId="5" pageNumber="335">13.4–16.8</td>
<td id="5C03FF4C107379E4D6EAFB0AB176FB75" box="[954,1067,1252,1273]" gridcol="3" gridrow="31" pageId="5" pageNumber="335">26.6–31.3</td>
<td id="5C03FF4C107379E4D1D8FB0AB1A5FB75" box="[1160,1272,1252,1273]" gridcol="4" gridrow="31" pageId="5" pageNumber="335">0.49–0.56</td>
<td id="5C03FF4C107379E4D010FB0AB0EDFB75" box="[1344,1456,1252,1273]" gridcol="5" gridrow="31" pageId="5" pageNumber="335">4½–5¼</td>
</tr>
<tr id="1FD29630107379E4D5D8FAEEB0EDFA99" box="[136,1456,1280,1301]" gridrow="32" pageId="5" pageNumber="335" rowspan-1="1" rowspan-5="1">
<th id="5C03FF4C107379E4D5D8FAEEB74CFA99" box="[136,529,1280,1301]" gridcol="0" gridrow="32" pageId="5" pageNumber="335">Malaysia (4597)</th>
<td id="5C03FF4C107379E4D7BDFAEEB603FA99" box="[749,862,1280,1301]" gridcol="2" gridrow="32" pageId="5" pageNumber="335">14.8 ± 1.17</td>
<td id="5C03FF4C107379E4D6EAFAEEB176FA99" box="[954,1067,1280,1301]" gridcol="3" gridrow="32" pageId="5" pageNumber="335">28.6 ± 1.80</td>
<td id="5C03FF4C107379E4D1D8FAEEB1A5FA99" box="[1160,1272,1280,1301]" gridcol="4" gridrow="32" pageId="5" pageNumber="335">0.52 ± 0.02</td>
</tr>
</table>
</paragraph>
<paragraph id="A15D94721073861ED5D8F9D6B72CF8E3" blockId="5.[136,775,1592,1903]" pageId="5" pageNumber="335">
<taxonomicName id="66E2EFF11073861ED5D8F9D6B42BF9C3" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[136,374,1592,1615]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="939648601073861ED5D8F9D6B42BF9C3" box="[136,374,1592,1615]" italics="true" pageId="5" pageNumber="335">Pterocyclos spaleotes</emphasis>
</taxonomicName>
clearly differs from any 
<taxonomicName id="66E2EFF11073861ED7C3F9D6B588F9E3" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="939648601073861ED7C3F9D6B65BF9C3" box="[659,774,1592,1615]" italics="true" pageId="5" pageNumber="335">Rhiostoma</emphasis>
species
</taxonomicName>
by the shape of its accessory respiratory structure located on the sutural area, and by its calcareous, low cupshaped and multilamellae operculum. Moreover, 
<taxonomicName id="66E2EFF11073861ED7DBF976B65AF923" baseAuthorityName="Tomlin" baseAuthorityYear="1932" box="[651,775,1688,1711]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="939648601073861ED7DBF976B65AF923" box="[651,775,1688,1711]" italics="true" pageId="5" pageNumber="335">P. spaleotes</emphasis>
</taxonomicName>
has been recorded (
<typeStatus id="7E592AD01073861ED431F956B4CDF943" box="[353,400,1720,1743]" pageId="5" pageNumber="335">type</typeStatus>
locality) much further south than 
<taxonomicName id="66E2EFF11073861ED5D8F936B5A1F963" authorityName="Benson" authorityYear="1860" box="[136,252,1752,1775]" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="genus">
<emphasis id="939648601073861ED5D8F936B5A1F963" box="[136,252,1752,1775]" italics="true" pageId="5" pageNumber="335">Rhiostoma</emphasis>
</taxonomicName>
, which does not occur south of the limestone karst in 
<collectingRegion id="63265A901073861ED5B6F916B47AF883" box="[230,295,1784,1807]" country="Malaysia" name="Perlis" pageId="5" pageNumber="335">Perlis</collectingRegion>
, 
<collectingCountry id="D9F5D4E21073861ED465F916B4C1F883" box="[309,412,1784,1807]" name="Malaysia" pageId="5" pageNumber="335">Malaysia</collectingCountry>
. Finally, preliminary analysis of the DNA sequence of a fragment of the cytochrome oxidase subunit I gene supports the transfer of 
<taxonomicName id="66E2EFF11073861ED77AF8D6B65BF8C3" authorityName="Tomlin" authorityYear="1932" box="[554,774,1848,1871]" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="species" species="spaleotes">
<emphasis id="939648601073861ED77AF8D6B65BF8C3" box="[554,774,1848,1871]" italics="true" pageId="5" pageNumber="335">Rhiostoma spaleotes</emphasis>
</taxonomicName>
to 
<taxonomicName id="66E2EFF11073861ED5F5F8B6B47FF8E3" authorityName="Benson" authorityYear="1832" box="[165,290,1880,1903]" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="335" phylum="Mollusca" rank="genus">
<emphasis id="939648601073861ED5F5F8B6B47FF8E3" box="[165,290,1880,1903]" italics="true" pageId="5" pageNumber="335">Pterocyclos</emphasis>
</taxonomicName>
(Tongkerd, unpublished data).
</paragraph>
</subSubSection>
</treatment>
</document>