treatments-xml/data/1A/71/87/1A7187CFFFC31777FDAEFA31E0075860.xml
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<mods:title id="B88C2CB4223DC9E4E48A4E843A4F41F5">The vertebrate Fauna of the Judith River formation, Montana</mods:title>
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Deinodon
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<emphasis id="A0ACEACBFFC31774FED2F9FDE0E85C18" box="[360,656,1603,1634]" italics="true" pageId="41" pageNumber="358">Aublysodon mirandus</emphasis>
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<bibRefCitation id="F6494B28FFC31774FD24F9FAE1185C18" author="LEIDY, J." box="[670,864,1603,1634]" journalOrPublisher="Proc. Acad. Nat. Sci. Philadelphia" pageId="41" pageNumber="358" pagination="197 - 200" part="20" refId="ref60703" refString="1868. Remarks on a jaw fragment of Megalosaurus. Proc. Acad. Nat. Sci. Philadelphia, vol. 20, pp. 197 - 200." title="Remarks on a jaw fragment of Megalosaurus" type="journal article" year="1868">LEIDY, 1868</bibRefCitation>
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<bibRefCitation id="F6494B28FFC31774FD2AF9D3E1235CF6" author="COPE, E. D." box="[656,859,1645,1676]" journalOrPublisher="Ibid." pageId="41" pageNumber="358" pagination="340 - 359" part="28" refId="ref57044" refString="1876 b. On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana. Ibid., vol. 28, pp. 340 - 359." title="On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana" type="journal article" year="1876">COPE, 1876b</bibRefCitation>
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<emphasis id="A0ACEACBFFC31774FED2F928E00A5CCC" box="[360,626,1686,1719]" italics="true" pageId="41" pageNumber="358">Laelaps incrassatus</emphasis>
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<bibRefCitation id="F6494B28FFC31774FC61F929E6405CCD" author="COPE, E. D." box="[987,1080,1687,1719]" journalOrPublisher="Ibid." pageId="41" pageNumber="358" pagination="340 - 359" part="28" refId="ref57044" refString="1876 b. On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana. Ibid., vol. 28, pp. 340 - 359." title="On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana" type="journal article" year="1876">1876b</bibRefCitation>
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<taxonomicName id="55D84D5AFFC31774FED2F97FE0015C9A" baseAuthorityName="Hay" baseAuthorityYear="1902" box="[360,633,1728,1760]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="species" species="hazenianus">
<emphasis id="A0ACEACBFFC31774FED2F97FE0015C9A" box="[360,633,1728,1760]" italics="true" pageId="41" pageNumber="358">Laelaps hazenianus</emphasis>
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<bibRefCitation id="F6494B28FFC31774FD32F97FE12C5C9B" author="COPE, E. D." box="[648,852,1728,1761]" journalOrPublisher="Ibid." pageId="41" pageNumber="358" pagination="340 - 359" part="28" refId="ref57044" refString="1876 b. On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana. Ibid., vol. 28, pp. 340 - 359." title="On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana" type="journal article" year="1876">COPE, 1876b</bibRefCitation>
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<taxonomicName id="55D84D5AFFC31774FED1F955E1595D70" authority="MARSH" authorityName="Marsh" authorityYear="1890" box="[363,801,1771,1803]" class="Reptilia" family="Ornithomimidae" genus="Ornithomimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lastPageId="41" lastPageNumber="359" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="grandis">
<emphasis id="A0ACEACBFFC31774FED1F955E0E75D70" box="[363,671,1771,1803]" italics="true" pageId="41" pageNumber="358">Ornithomimus grandis</emphasis>
<bibRefCitation id="F6494B28FFC31774FD14F955E1FA5D70" author="MARSH, O. C." box="[686,898,1771,1802]" journalOrPublisher="Ibid." pageId="41" pageNumber="358" pagination="81 - 86" part="39" refId="ref61639" refString="1890. Description of new dinosaurian reptiles. Ibid., ser. 3, vol. 39, pp. 81 - 86, pl. 1." title="Description of new dinosaurian reptiles" type="journal article" year="1890">MARSH, 1890</bibRefCitation>
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<emphasis id="A0ACEACBFFC31774FED2F8ABE15D5D49" box="[360,805,1813,1843]" italics="true" pageId="41" pageNumber="358">
<taxonomicName id="55D84D5AFFC31774FED2F8ABE0B45D49" authorityName="Hay" authorityYear="1899" box="[360,716,1813,1843]" class="Reptilia" genus="Dryptosaurus" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="kenabekides">Dryptosaurus kenabekides</taxonomicName>
<bibRefCitation id="F6494B28FFC31774FD60F8ABE1FE5D49" author="HAY, O. P." box="[730,902,1813,1843]" journalOrPublisher="Amer. geol." pageId="41" pageNumber="358" pagination="345 - 349" part="24" refId="ref59027" refString="HAY, O. P. 1899. On the nomenclature of certain American fossil vertebrates. Amer. geol., vol. 24, pp. 345 - 349." title="On the nomenclature of certain American fossil vertebrates" type="journal article" year="1899">HAY, 1899</bibRefCitation>
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</paragraph>
<paragraph id="926736D9FFC31774FED3F880E1A95D24" blockId="41.[360,1205,1560,2462]" box="[361,977,1854,1886]" pageId="41" pageNumber="358">
<treatmentCitationGroup id="B2C811F7FFC31774FED3F880E1A95D24" box="[361,977,1854,1886]" pageId="41" pageNumber="358">
<treatmentCitation id="137910C8FFC31774FED3F880E1B45D24" ID-DOI="http://doi.org/10.5281/zenodo.4912562" ID-Zenodo-Dep="4912562" author="Lambe" box="[361,972,1854,1886]" httpUri="http://treatment.plazi.org/id/03807171D73AFFCDE8F0F534FD66DBBF" pageId="41" pageNumber="358" rank="species" treatmentCitationLabel="synonym" year="1914">
<taxonomicName id="55D84D5AFFC31774FED3F880E0F15D24" authority="Lambe, 1914" box="[361,649,1854,1886]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="libratus">
<emphasis id="A0ACEACBFFC31774FED3F880E0695D24" box="[361,529,1854,1886]" italics="true" pageId="41" pageNumber="358">Gorgosaurus</emphasis>
libratus
</taxonomicName>
LAMBE, 1914, p. 13
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DAC26552FFC31777FE28F8C3E0075860" lastPageId="42" lastPageNumber="359" pageId="41" pageNumber="358" type="discussion">
<paragraph id="926736D9FFC31774FE28F8C3E5485E0E" blockId="41.[360,1205,1560,2462]" lastBlockId="41.[1276,2138,358,2796]" pageId="41" pageNumber="358">
<taxonomicName id="55D84D5AFFC31774FE28F8C3E0B95DDA" authorityName="Leidy" authorityYear="1856" box="[402,705,1917,1952]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="horridus">
<emphasis id="A0ACEACBFFC31774FE28F8C3E0B95DDA" box="[402,705,1917,1952]" italics="true" pageId="41" pageNumber="358">Deinodon horridus</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation id="F6494B28FFC31774FBE8F8C0E3AF5DB4" author="LEIDY, J." journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="41" pageNumber="358" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy (1856)</bibRefCitation>
for
<materialsCitation id="22B03C84FFC31774FD84F812E6CC5222" GeologicalContext="Upper Cretaceous" ID-GBIF-Occurrence="2413160537" collectingCountry="United States" collectorName="Hayden" formation="Judith River Formation" pageId="41" pageNumber="358" preparations="about a dozen teeth" typeStatus="type series">about a dozen teeth collected by Hayden from the Judith River Formation. This species was the first of the carnivorous dinosaurs to be recovered from the Upper Cretaceous</materialsCitation>
sediments of the American continent. The collection of teeth on which the genus was based consists of two distinct morphological types, although
<bibRefCitation id="F6494B28FFC31774FDA9F751E0905368" author="LEIDY, J." box="[531,744,2287,2322]" journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="41" pageNumber="358" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy (1856</bibRefCitation>
, p. 73;
<bibRefCitation id="F6494B28FFC31774FCC3F74EE1B1536B" author="LEIDY, J." box="[889,969,2288,2321]" journalOrPublisher="Trans. Amer. Philos. Soc." pageId="41" pageNumber="358" pagination="139 - 154" part="11" refId="ref60658" refString="1860. Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska. Trans. Amer. Philos. Soc., new ser., vol. 11, pp. 139 - 154, pls. 8 - 1 1." title="Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska" type="journal article" year="1860">1860</bibRefCitation>
, p. 144) considered them as belonging to a single species.
<bibRefCitation id="F6494B28FFC31774FED3F6F4E03A5317" author="LEIDY, J." box="[361,578,2378,2413]" journalOrPublisher="Trans. Amer. Philos. Soc." pageId="41" pageNumber="358" pagination="139 - 154" part="11" refId="ref60658" refString="1860. Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska. Trans. Amer. Philos. Soc., new ser., vol. 11, pp. 139 - 154, pls. 8 - 1 1." title="Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska" type="journal article" year="1860">Leidy (1860</bibRefCitation>
, p. 145) correctly guessed that the differences involved in the two groups of teeth were positional in nature and suggested that the teeth with U-shaped cross sections were incisors. Later,
<bibRefCitation id="F6494B28FFC31774FAC7FE79E4205B91" author="LEIDY, J." box="[1405,1624,455,491]" journalOrPublisher="Proc. Acad. Nat. Sci. Philadelphia" pageId="41" pageNumber="358" pagination="197 - 200" part="20" refId="ref60703" refString="1868. Remarks on a jaw fragment of Megalosaurus. Proc. Acad. Nat. Sci. Philadelphia, vol. 20, pp. 197 - 200." title="Remarks on a jaw fragment of Megalosaurus" type="journal article" year="1868">Leidy (1868</bibRefCitation>
, p. 198) erected a separate genus,
<taxonomicName id="55D84D5AFFC31774FAC2FE48E4535860" authorityName="Leidy" authorityYear="1868" box="[1400,1579,502,538]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774FAC2FE48E4535860" box="[1400,1579,502,538]" italics="true" pageId="41" pageNumber="358">Aublysodon</emphasis>
</taxonomicName>
, for the inclusion of those teeth he had earlier regarded as incisors. These teeth have serrated crests on the posterior side.
<bibRefCitation id="F6494B28FFC31774F84DFDEFE70058D9" author="COPE, E. D." journalOrPublisher="Proc. Acad. Nat. Sci. Philadelphia" pageId="41" pageNumber="358" pagination="248 - 261" part="28" refId="ref57008" refString="1876 a. Descriptions of some vertebrate remains from the Fort Union beds of Montana. Proc. Acad. Nat. Sci. Philadelphia, vol. 28, pp. 248 - 261." title="Descriptions of some vertebrate remains from the Fort Union beds of Montana" type="journal article" year="1876">Cope (1876a</bibRefCitation>
,
<bibRefCitation id="F6494B28FFC31774FA21FDC1E47858D9" author="COPE, E. D." box="[1435,1536,639,675]" journalOrPublisher="Ibid." pageId="41" pageNumber="358" pagination="340 - 359" part="28" refId="ref57044" refString="1876 b. On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana. Ibid., vol. 28, pp. 340 - 359." title="On some extinct reptiles and batrachians from the Judith River and Fox Hills beds of Montana" type="journal article" year="1876">1876b</bibRefCitation>
) described a number of species from the Judith River Formation under the genus
<taxonomicName id="55D84D5AFFC31774FAD4FD62E792597A" box="[1390,1514,732,768]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="A0ACEACBFFC31774FAD4FD62E792597A" box="[1390,1514,732,768]" italics="true" pageId="41" pageNumber="358">Laelaps</emphasis>
</taxonomicName>
, the type of which is
<taxonomicName id="55D84D5AFFC31774F8D8FD63EA30597A" baseAuthorityName="Cope" baseAuthorityYear="1866" box="[1890,2120,732,768]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="species" species="aquilunguis">
<emphasis id="A0ACEACBFFC31774F8D8FD63E507597A" box="[1890,1919,733,768]" italics="true" pageId="41" pageNumber="358">L</emphasis>
.
<emphasis id="A0ACEACBFFC31774F82EFD62EA30597A" box="[1940,2120,732,768]" italics="true" pageId="41" pageNumber="358">aquilunguis</emphasis>
</taxonomicName>
, obtained from New Jersey. The genus
<taxonomicName id="55D84D5AFFC31774F85DFCB4E72A5921" authorityName="Leidy" authorityYear="1868" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F85DFCB4E72A5921" italics="true" pageId="41" pageNumber="358">Aublysodon</emphasis>
</taxonomicName>
was placed in synonymy with
<taxonomicName id="55D84D5AFFC31774F83FFC86EA6F5921" box="[1925,2071,824,859]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F83FFC86EA6F5921" box="[1925,2071,824,859]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
by Matthew and Brown (1922). Of the three species of
<taxonomicName id="55D84D5AFFC31774FA75FC2BE43E59C0" box="[1487,1606,917,954]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis id="A0ACEACBFFC31774FA75FC2BE43E59C0" box="[1487,1606,917,954]" italics="true" pageId="41" pageNumber="358">Laelaps</emphasis>
</taxonomicName>
collected from the Judith River Formation, two,
<taxonomicName id="55D84D5AFFC31774F91DFC7AE5075992" baseAuthorityName="Cope" baseAuthorityYear="1876" box="[1703,1919,963,1000]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="species" species="explanatus">
<emphasis id="A0ACEACBFFC31774F91DFC7AE4BC599D" box="[1703,1732,964,999]" italics="true" pageId="41" pageNumber="358">L</emphasis>
.
<emphasis id="A0ACEACBFFC31774F967FC7DE5075992" box="[1757,1919,963,1000]" italics="true" pageId="41" pageNumber="358">explanatus</emphasis>
</taxonomicName>
and
<taxonomicName id="55D84D5AFFC31774F859FC7AE7365E6C" baseAuthorityName="Cope" baseAuthorityYear="1876" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="species" species="falculus">
<emphasis id="A0ACEACBFFC31774F859FC7AE587599D" box="[2019,2047,964,999]" italics="true" pageId="41" pageNumber="358">L</emphasis>
.
<emphasis id="A0ACEACBFFC31774F7AAFC7DE7365E6C" italics="true" pageId="41" pageNumber="358">falculus</emphasis>
</taxonomicName>
, were referred to
<taxonomicName id="55D84D5AFFC31774F900FC4DE5DE5E6C" authorityName="Matthew &amp; Brown" authorityYear="1922" box="[1722,1958,1011,1046]" class="Reptilia" family="Dromaeosauridae" genus="Dromaeosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F900FC4DE5DE5E6C" box="[1722,1958,1011,1046]" italics="true" pageId="41" pageNumber="358">Dromaeosaurus</emphasis>
</taxonomicName>
and the other,
<taxonomicName id="55D84D5AFFC31774FACDFB9CE42D5E3F" baseAuthorityName="Cope" baseAuthorityYear="1876" box="[1399,1621,1056,1093]" class="Arachnida" family="Ascidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Mesostigmata" pageId="41" pageNumber="358" phylum="Arthropoda" rank="species" species="incrassatus">
<emphasis id="A0ACEACBFFC31774FACDFB9CE7EB5E3E" box="[1399,1427,1058,1092]" italics="true" pageId="41" pageNumber="358">L</emphasis>
.
<emphasis id="A0ACEACBFFC31774FA16FB9EE42D5E3F" box="[1452,1621,1056,1093]" italics="true" pageId="41" pageNumber="358">incrassatus</emphasis>
</taxonomicName>
, was transferred to
<taxonomicName id="55D84D5AFFC31774F87BFB9FEA2B5E3E" box="[1985,2131,1057,1092]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F87BFB9FEA2B5E3E" box="[1985,2131,1057,1092]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
by Matthew and Brown (1922).
</paragraph>
<paragraph id="926736D9FFC31774FA91FBC0EA6A5DF9" blockId="41.[1276,2138,358,2796]" pageId="41" pageNumber="358">
On the basis of the dentition it is not possible to separate the closely related genera
<taxonomicName id="55D84D5AFFC31774F82DFB15EA2C5EAA" box="[1943,2132,1195,1232]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F82DFB15EA2C5EAA" box="[1943,2132,1195,1232]" italics="true" pageId="41" pageNumber="358">Gorgosaurus</emphasis>
</taxonomicName>
and
<taxonomicName id="55D84D5AFFC31774FAE7FB64E78A5E87" box="[1373,1522,1242,1277]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774FAE7FB64E78A5E87" box="[1373,1522,1242,1277]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
. As suggested by a number of authors including Matthew and Brown (1922, p. 383), and
<bibRefCitation id="F6494B28FFC31774FA45FA8BE48D5F23" author="RUSSELL, L. S." box="[1535,1781,1332,1369]" journalOrPublisher="Toronto Univ. Roy. Ont. Mus., Life Sci. Div." pageId="41" pageNumber="358" pagination="1 - 24" part="61" refId="ref63199" refString="1964. Cretaceous non-marine faunas of northwestern North America. Toronto Univ. Roy. Ont. Mus., Life Sci. Div., contrib. no. 61, pp. 1 - 24." title="Cretaceous non-marine faunas of northwestern North America" type="journal article" year="1964">Russell (1964</bibRefCitation>
, p. 13), the genus
<taxonomicName id="55D84D5AFFC31774FAB9FADDE7C65FFD" box="[1283,1470,1379,1415]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774FAB9FADDE7C65FFD" box="[1283,1470,1379,1415]" italics="true" pageId="41" pageNumber="358">Gorgosaurus</emphasis>
</taxonomicName>
is probably synonymous with the Judith River Formation
<taxonomicName id="55D84D5AFFC31774F90BFA2CE53F5FCF" box="[1713,1863,1426,1461]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F90BFA2CE53F5FCF" box="[1713,1863,1426,1461]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
. Lambe (1914) described
<taxonomicName id="55D84D5AFFC31774FA01FA01E4115F9E" baseAuthorityName="Lambe" baseAuthorityYear="1914" box="[1467,1641,1471,1508]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="libratus">
<emphasis id="A0ACEACBFFC31774FA01FA01E7A15F9E" box="[1467,1497,1471,1508]" italics="true" pageId="41" pageNumber="358">G</emphasis>
.
<emphasis id="A0ACEACBFFC31774FA4EFA01E4115F9E" box="[1524,1641,1471,1508]" italics="true" pageId="41" pageNumber="358">libratus</emphasis>
</taxonomicName>
from the Oldman Formation and then restudied it in greater detail (Lambe, 1917) on the basis of more complete material than that known for
<taxonomicName id="55D84D5AFFC31774F8EFF9F4E5935C14" box="[1877,2027,1610,1646]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F8EFF9F4E5935C14" box="[1877,2027,1610,1646]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
. The name
<taxonomicName id="55D84D5AFFC31774FAC2F9C7E4755CE7" box="[1400,1549,1657,1693]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774FAC2F9C7E4755CE7" box="[1400,1549,1657,1693]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
, however, has priority and no differences can be shown to exist between the type specimen of
<taxonomicName id="55D84D5AFFC31774F9D7F968E4865C83" box="[1645,1790,1750,1785]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F9D7F968E4865C83" box="[1645,1790,1750,1785]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
and material of
<taxonomicName id="55D84D5AFFC31774FAB9F8BDE7BA5D5D" box="[1283,1474,1795,1831]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774FAB9F8BDE7BA5D5D" box="[1283,1474,1795,1831]" italics="true" pageId="41" pageNumber="358">Gorgosaurus</emphasis>
</taxonomicName>
. The name
<taxonomicName id="55D84D5AFFC31774F90EF8BAE53D5D5D" box="[1716,1861,1796,1831]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC31774F90EF8BAE53D5D5D" box="[1716,1861,1796,1831]" italics="true" pageId="41" pageNumber="358">Deinodon</emphasis>
</taxonomicName>
should be retained for the large carnivorous dinosaur found in the Judith River and Oldman formations.
</paragraph>
<paragraph id="926736D9FFC31774FA91F830E71950C6" blockId="41.[1276,2138,358,2796]" pageId="41" pageNumber="358">
The present collection consists of a number of teeth similar to those described by
<bibRefCitation id="F6494B28FFC31774F8D4F802EA285D9A" author="LEIDY, J." box="[1902,2128,1979,2016]" journalOrPublisher="Proc. Acad. Nat. Sci. Phila- delphia" pageId="41" pageNumber="358" pagination="72 - 73" part="8" refId="ref60602" refString="LEIDY, J. 1856. Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory. Proc. Acad. Nat. Sci. Phila- delphia, vol. 8, pp. 72 - 73." title="Notices of the remains of extinct reptiles and fishes discovered by Dr. F. V. Hayden in the badlands of the Judith River, Nebraska Territory" type="journal article" year="1856">Leidy (1856)</bibRefCitation>
and later figured by him (
<figureCitation id="0AE32A5CFFC31774F94DF857E7A45246" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="41" pageNumber="358" targetBox="[57,1772,29,2210]" targetPageId="17">
<bibRefCitation id="F6494B28FFC31774F94DF857E5AC5277" author="LEIDY, J." box="[1783,2004,2025,2061]" journalOrPublisher="Trans. Amer. Philos. Soc." pageId="41" pageNumber="358" pagination="139 - 154" part="11" refId="ref60658" refString="1860. Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska. Trans. Amer. Philos. Soc., new ser., vol. 11, pp. 139 - 154, pls. 8 - 1 1." title="Extinct vertebrata from the Judith River and Great Lignite formations of Nebraska" type="journal article" year="1860">Leidy, 1860</bibRefCitation>
, pl. 9, figs. 21-48)
</figureCitation>
.
<materialsCitation id="22B03C84FFC31774F9B9F7A6E4945246" GeologicalContext="Campanian" ID-GBIF-Occurrence="3396386301" box="[1539,1772,2072,2108]" collectingCountry="United States" collectingCounty="Chouteau Co." collectingRegion="Montana" collectionCode="AMNH" collectorName="A. Sahni" formation="Judith River" httpUri="http://research.amnh.org/paleontology/search.php?action=detail&amp;specimen_id=51553 " location="Clam Bank Hollow" pageId="41" pageNumber="358" preparations="large saber-shaped linguolabially compressed tooth" specimenCode="AMNH 8513">AMNH 8513</materialsCitation>
is a large saber-shaped linguolabially compressed tooth. Both its anterior and posterior edges are serrated. It is either a maxilla or dentary tooth (
<figureCitation id="0AE32A5CFFC31774F8CCF71DEA4752BC" box="[1910,2111,2210,2247]" captionStart="FIG" captionStartId="40.[200,257,2244,2275]" captionTargetBox="[331,2100,338,2536]" captionTargetPageId="39" captionText="FIG. 8. A. Chamops segnis. AMNH 8486, labial view of left dentary. B. Lingual view. Both x 5. C. AMNH 8488, lingual view of left maxilla. D. Labial view. Both x 5. E. cf. Xenosauridae, AMNH 8498, labial view of left maxilla. F. Lingual view. Both x 5. G. Peltosaurus piger, AMNH 8495, lingual view of left maxilla, x 5. H. AMNH 8494, lingual view of right dentary, x 5. I. Paraderma bogerti, AMNH 8504, lingual view of right maxilla. x 5. J. Parasaniwa wyomingensis, AMNH 8499, lingual view of left dentary, x 5. K. Leptochamops denticulatus, AMNH 8491, lingual view of right maxilla, x 5. L. Exostinus lancensis, AMNH 8497, lingual view of right maxilla, x 5. M. Deinodon horridus, AMNH 8513, posterior view of isolated tooth. N. Lateral view. Both x 13. 0. AMNH 8515, lateral view of anterior dentary tooth. P. Posterior view. Both x 13. Q. AMNH 8514, posterior view of?premaxillary tooth. R. Lateral view. Both x 12. S. Paronychodon lacustris, AMNH 8522, lingual view of isolated tooth. T. Labial view. Both x 12. U. Kritosaurus cf. K. breviceps, AMNH 8525, lateral view of isolated tooth. V. Posterior view. Both x 1 2. W. Procheneosaurus altidens, AMNH 8527, labial view of isolated tooth. X. Anterior or posterior view. Both x 12. Y, Z.?Sebecosuchia, AMNH 8549, views of an solated tooth. Both x 10." httpUri="https://zenodo.org/record/3382465/files/figure.png" pageId="41" pageNumber="358">fig. 8M, N</figureCitation>
). The premaxillary teeth are small with a Ushaped cross section. Morphologically similar, but larger, teeth come from the anterior part of the dentary. The premaxillary teeth (
<materialsCitation id="22B03C84FFC31774F872F6E4E72C53D7" GeologicalContext="Campanian" ID-GBIF-Occurrence="3396386302" collectingCountry="United States" collectingCounty="Chouteau Co." collectingRegion="Montana" collectionCode="AMNH" collectorName="A. Sahni" formation="Judith River" httpUri="http://research.amnh.org/paleontology/search.php?action=detail&amp;specimen_id=51554 " location="Clam Bank Hollow" pageId="41" pageNumber="358" preparations="premaxillary teeth" specimenCode="AMNH 8514">AMNH 8514</materialsCitation>
,
<figureCitation id="0AE32A5CFFC31774FACBF636E45753D7" box="[1393,1583,2440,2477]" captionStart="FIG" captionStartId="40.[200,257,2244,2275]" captionTargetBox="[331,2100,338,2536]" captionTargetPageId="39" captionText="FIG. 8. A. Chamops segnis. AMNH 8486, labial view of left dentary. B. Lingual view. Both x 5. C. AMNH 8488, lingual view of left maxilla. D. Labial view. Both x 5. E. cf. Xenosauridae, AMNH 8498, labial view of left maxilla. F. Lingual view. Both x 5. G. Peltosaurus piger, AMNH 8495, lingual view of left maxilla, x 5. H. AMNH 8494, lingual view of right dentary, x 5. I. Paraderma bogerti, AMNH 8504, lingual view of right maxilla. x 5. J. Parasaniwa wyomingensis, AMNH 8499, lingual view of left dentary, x 5. K. Leptochamops denticulatus, AMNH 8491, lingual view of right maxilla, x 5. L. Exostinus lancensis, AMNH 8497, lingual view of right maxilla, x 5. M. Deinodon horridus, AMNH 8513, posterior view of isolated tooth. N. Lateral view. Both x 13. 0. AMNH 8515, lateral view of anterior dentary tooth. P. Posterior view. Both x 13. Q. AMNH 8514, posterior view of?premaxillary tooth. R. Lateral view. Both x 12. S. Paronychodon lacustris, AMNH 8522, lingual view of isolated tooth. T. Labial view. Both x 12. U. Kritosaurus cf. K. breviceps, AMNH 8525, lateral view of isolated tooth. V. Posterior view. Both x 1 2. W. Procheneosaurus altidens, AMNH 8527, labial view of isolated tooth. X. Anterior or posterior view. Both x 12. Y, Z.?Sebecosuchia, AMNH 8549, views of an solated tooth. Both x 10." httpUri="https://zenodo.org/record/3382465/files/figure.png" pageId="41" pageNumber="358">fig. 8Q, R</figureCitation>
) are usually smaller than the anterior dentary teeth (
<materialsCitation id="22B03C84FFC31774F900F60BE5D053A3" GeologicalContext="Campanian" ID-GBIF-Occurrence="3396386303" box="[1722,1960,2485,2521]" collectingCountry="United States" collectingCounty="Chouteau Co." collectingRegion="Montana" collectionCode="AMNH" collectorName="A. Sahni" formation="Judith River" httpUri="http://research.amnh.org/paleontology/search.php?action=detail&amp;specimen_id=51555 " location="Clam Bank Hollow" pageId="41" pageNumber="358" preparations="anterior dentary teeth" specimenCode="AMNH 8515">AMNH 8515</materialsCitation>
,
<figureCitation id="0AE32A5CFFC31774F87FF608E7615072" captionStart="FIG" captionStartId="40.[200,257,2244,2275]" captionTargetBox="[331,2100,338,2536]" captionTargetPageId="39" captionText="FIG. 8. A. Chamops segnis. AMNH 8486, labial view of left dentary. B. Lingual view. Both x 5. C. AMNH 8488, lingual view of left maxilla. D. Labial view. Both x 5. E. cf. Xenosauridae, AMNH 8498, labial view of left maxilla. F. Lingual view. Both x 5. G. Peltosaurus piger, AMNH 8495, lingual view of left maxilla, x 5. H. AMNH 8494, lingual view of right dentary, x 5. I. Paraderma bogerti, AMNH 8504, lingual view of right maxilla. x 5. J. Parasaniwa wyomingensis, AMNH 8499, lingual view of left dentary, x 5. K. Leptochamops denticulatus, AMNH 8491, lingual view of right maxilla, x 5. L. Exostinus lancensis, AMNH 8497, lingual view of right maxilla, x 5. M. Deinodon horridus, AMNH 8513, posterior view of isolated tooth. N. Lateral view. Both x 13. 0. AMNH 8515, lateral view of anterior dentary tooth. P. Posterior view. Both x 13. Q. AMNH 8514, posterior view of?premaxillary tooth. R. Lateral view. Both x 12. S. Paronychodon lacustris, AMNH 8522, lingual view of isolated tooth. T. Labial view. Both x 12. U. Kritosaurus cf. K. breviceps, AMNH 8525, lateral view of isolated tooth. V. Posterior view. Both x 1 2. W. Procheneosaurus altidens, AMNH 8527, labial view of isolated tooth. X. Anterior or posterior view. Both x 12. Y, Z.?Sebecosuchia, AMNH 8549, views of an solated tooth. Both x 10." httpUri="https://zenodo.org/record/3382465/files/figure.png" pageId="41" pageNumber="358">fig. 80, P</figureCitation>
) and have serrated crests lateral to the apex of the tooth that become more posterior toward the base of the tooth. A strong median keel is present that is not found in the larger dentary teeth.
</paragraph>
<footnote id="F1C32AD7FFC31774FE31F66BE026509F" pageId="41" pageNumber="358">
<paragraph id="926736D9FFC31774FE31F66BE026509F" blockId="41.[359,1205,2499,2790]" pageId="41" pageNumber="358">
1 [
<bibRefCitation id="F6494B28FFC31774FE16F679E0055399" author="RUSSELL, D. A." box="[428,637,2503,2531]" journalOrPublisher="Natl. Mus. Nat. Sci., Publs. in Palaeont." pageId="41" pageNumber="358" pagination="1 - 34" part="1" refId="ref63003" refString="1970. Tyrannosaurs from the late Cretaceous of Western Canada. Natl. Mus. Nat. Sci., Publs. in Palaeont., no. 1, pp. i-viii, 1 - 34, figs. 1 - 9, pls. 1 - 4, paged." title="Tyrannosaurs from the late Cretaceous of Western Canada." type="journal article" year="1970">Russell (1970)</bibRefCitation>
revised the taxonomy of Upper Cretaceous carnosaurs and argued that
<taxonomicName id="55D84D5AFFC31774FC3DF650E60C5070" authorityName="Leidy" authorityYear="1856" box="[903,1140,2542,2570]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="horridus">
<emphasis id="A0ACEACBFFC31774FC3DF650E60C5070" box="[903,1140,2542,2570]" italics="true" pageId="41" pageNumber="358">Deinodon horridus</emphasis>
</taxonomicName>
is a nomen
<emphasis id="A0ACEACBFFC31774FE7BF5ACE0655055" box="[449,541,2578,2607]" italics="true" pageId="41" pageNumber="358">dubium</emphasis>
and that individual teeth are not generically determinable (
<emphasis id="A0ACEACBFFC31774FDF9F588E0035028" box="[579,635,2614,2642]" italics="true" pageId="41" pageNumber="358">Ibid</emphasis>
, p. 3). According to the classification presented by
<bibRefCitation id="F6494B28FFC31774FDF8F5E2E1605002" author="RUSSELL, D. A." box="[578,792,2652,2680]" journalOrPublisher="Natl. Mus. Nat. Sci., Publs. in Palaeont." pageId="41" pageNumber="358" pagination="1 - 34" part="1" refId="ref63003" refString="1970. Tyrannosaurs from the late Cretaceous of Western Canada. Natl. Mus. Nat. Sci., Publs. in Palaeont., no. 1, pp. i-viii, 1 - 34, figs. 1 - 9, pls. 1 - 4, paged." title="Tyrannosaurs from the late Cretaceous of Western Canada." type="journal article" year="1970">Russell (1970)</bibRefCitation>
and Colbert and Russell (1969) large carnivorous dinosaur teeth could be assigned most appropriately to the family
<taxonomicName id="55D84D5AFFC31774FC1FF519E6DC50BA" authorityName="Osborn" authorityYear="1906" box="[933,1188,2727,2752]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="family">Tyrannosauridae</taxonomicName>
.] M.
<emphasis id="A0ACEACBFFC31774FE27F574E3CE509C" bold="true" box="[413,438,2762,2790]" pageId="41" pageNumber="358">C</emphasis>
. McKenna
</paragraph>
</footnote>
<paragraph id="926736D9FFC31774FDE7F577E019509F" blockId="41.[359,1205,2499,2790]" box="[605,609,2761,2789]" pageId="41" pageNumber="358">.</paragraph>
<paragraph id="926736D9FFC31777FA90F579E0075860" blockId="41.[1276,2138,358,2796]" lastBlockId="42.[152,1006,364,538]" lastPageId="42" lastPageNumber="359" pageId="41" pageNumber="358">
The type specimen of
<taxonomicName id="55D84D5AFFC31777F954F579E3FF5BF5" authority="Marsh (1890)" authorityName="Marsh" authorityYear="1890" class="Reptilia" family="Ornithomimidae" genus="Ornithomimus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lastPageId="42" lastPageNumber="359" order="Dinosauria" pageId="41" pageNumber="358" phylum="Chordata" rank="species" species="grandis">
<emphasis id="A0ACEACBFFC31774F954F579EA2C5090" box="[1774,2132,2759,2794]" italics="true" pageId="41" pageNumber="358">Ornithomimus grandis</emphasis>
<bibRefCitation id="F6494B28FFC01777FF24FED3E3FF5BF5" author="MARSH, O. C." box="[158,391,364,400]" journalOrPublisher="Ibid." pageId="42" pageNumber="359" pagination="81 - 86" part="39" refId="ref61639" refString="1890. Description of new dinosaurian reptiles. Ibid., ser. 3, vol. 39, pp. 81 - 86, pl. 1." title="Description of new dinosaurian reptiles" type="journal article" year="1890">Marsh (1890)</bibRefCitation>
</taxonomicName>
was supposedly recovered from the Eagle Sandstone, but probably came from the Judith River Formation, and was referred by
<bibRefCitation id="F6494B28FFC01777FF24FE49E3DF5860" author="GILMORE, C. W." box="[158,423,503,538]" journalOrPublisher="Bull. U. S. Natl. Mus." pageId="42" pageNumber="359" pagination="1 - 154" part="110" refId="ref58357" refString="1920. Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus. Bull. U. S. Natl. Mus., vol. 110, pp. 1 - 154, figs. 1 - 78, pls. 1 - 36." title="Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus" type="journal article" year="1920">Gilmore (1920)</bibRefCitation>
to
<taxonomicName id="55D84D5AFFC01777FE5DFE46E0035860" box="[487,635,504,538]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="42" pageNumber="359" phylum="Chordata" rank="genus">
<emphasis id="A0ACEACBFFC01777FE5DFE46E0035860" box="[487,635,504,538]" italics="true" pageId="42" pageNumber="359">Deinodon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>