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<mods:title id="A30D5038DE754CAECF4B799FB56D78DA">Integrative taxonomy study of brackish water crabs of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Polynesia, with description of two new species</mods:title>
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<mods:namePart id="FAC501FE25194F28BCF058C8A4053B03">Mazancourt, Valentin De</mods:namePart>
<mods:affiliation id="7C2E50B0322D1CEB65277B89916F317A">Muséum national dHistoire naturelle, UMR BOREA, CP 026, 57, rue Cuvier, F- 75231 Paris, Cedex 05, France.</mods:affiliation>
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<mods:namePart id="6B7CA3D815527A50B63C32C1B534EDB5">Mazel, Virgile</mods:namePart>
<mods:affiliation id="4CE3BC7F8C6DFB5B12A8D5068943630C">Ichtyo-Pacific, Virgile Mazel BP 381816 Punavai, 98718 Punaauia, Tahiti, Polynésie française.</mods:affiliation>
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<mods:affiliation id="7ECB8E62BFE6471FCB1D94940C05D166">Muséum national dHistoire naturelle, UMR BOREA, CP 026, 57, rue Cuvier, F- 75231 Paris, Cedex 05, France.</mods:affiliation>
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<treatment id="03FF87F3E314FFE91CBEFA18FA3DF829" ID-DOI="http://doi.org/10.5281/zenodo.12727239" ID-GBIF-Taxon="235040516" ID-Zenodo-Dep="12727239" LSID="urn:lsid:plazi:treatment:03FF87F3E314FFE91CBEFA18FA3DF829" httpUri="http://treatment.plazi.org/id/03FF87F3E314FFE91CBEFA18FA3DF829" lastPageNumber="293" pageId="26" pageNumber="293">
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<heading id="D0A18189E314FFE91CBEFA18FD13F9D4" bold="true" box="[151,690,1514,1541]" fontSize="11" level="1" pageId="26" pageNumber="293" reason="1">
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Key to Polynesian species of
<taxonomicName id="4C564D66E314FFE91DD4FA18FD13F9D4" ID-CoL="7PWZ6" authorityName="Easteranus Rathbun" authorityYear="1907" box="[509,690,1514,1541]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAF7E314FFE91DD4FA18FD13F9D4" bold="true" box="[509,690,1514,1541]" italics="true" pageId="26" pageNumber="293">Ptychognathus</emphasis>
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<paragraph id="8BE936E5E314FFE91CBEF9D9FA3DF98F" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF9D9FA3DF98F" pageId="26" pageNumber="293">1. Cheliped carpus with spine at inner angle (female not in male). P24 merus with spine on superior border distally (male and female). Carapace fronto-orbital width 0.890.96 times CW (male and female).................................... 2</keyLead>
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<paragraph id="8BE936E5E314FFE91CBEF994FA3DF949" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF994FA3DF949" pageId="26" pageNumber="293"> Cheliped carpus of without spine at inner angle (female and male). P24 merus without spine on superior border distally (male and female). Carapace fronto-orbital width 0.780.90 times CW (male and female)................................. 3</keyLead>
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<keyStep id="30A22E40E314FFE91CBEF953FA3DF8DF" pageId="26" pageNumber="293">
<paragraph id="8BE936E5E314FFE91CBEF953FA3DF902" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF953FA3DF902" pageId="26" pageNumber="293">
2. Third maxilliped exopod 1.41.6 (male), 1.31.4 (female) times as long as ischium. P5 dactylus 2.53.0 times as long as broad (male and female). Endemic to
<collectingRegion id="4992F807E314FFE91E39F94CFD63F905" box="[528,706,1726,1748]" country="French Polynesia" name="Iles Marquises" pageId="26" pageNumber="293">Marquesas Islands</collectingRegion>
................................................
<taxonomicName id="4C564D66E314FFE91924F94DFA3DF902" ID-CoL="6WNLP" authorityName="Finnegan" authorityYear="1931" box="[1293,1436,1726,1748]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E314FFE91924F94DFA3DF902" box="[1293,1436,1726,1748]" italics="true" pageId="26" pageNumber="293">P. crassimanus</emphasis>
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<keyLead id="30A795D0E314FFE91CBEF92EFA3DF8DF" pageId="26" pageNumber="293">
Third maxilliped exopod 1.71.9 (male), 1.41.6 (female) times as long as ischium. P5 dactylus 3.03.9 times as long as broad (male and female).
<collectingCountry id="F3417675E314FFE91DB5F90AFDBDF8DF" box="[412,540,1784,1806]" name="Cook Islands" pageId="26" pageNumber="293">Cook Islands</collectingCountry>
,
<collectingCountry id="F3417675E314FFE91E0FF90AFD6DF8DF" box="[550,716,1784,1806]" name="French Polynesia" pageId="26" pageNumber="293">French Polynesia</collectingCountry>
(Gambier, Austral and Society) and Easter Island.........
<taxonomicName id="4C564D66E314FFE91934F90BFA3DF8DF" ID-CoL="6WNM3" authorityName="Rathbun" authorityYear="1907" box="[1309,1436,1785,1806]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="species" species="easteranus">
<emphasis id="B922EAF7E314FFE91934F90BFA3DF8DF" box="[1309,1436,1785,1806]" italics="true" pageId="26" pageNumber="293">P. easteranus</emphasis>
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<keyLead id="30A795D0E314FFE91CBEF8E4FA3DF899" pageId="26" pageNumber="293">
3. Chelipeds glabrous; very characteristic small tuft of setae at tip of pollex. Third maxilliped exopod 2.12.6 (male), 1.61.9 (female) times as long as ischium. Widespread.......................................................
<taxonomicName id="4C564D66E314FFE91916F8C6FA3DF899" ID-CoL="78NM4" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1868" box="[1343,1436,1843,1865]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="species" species="riedelii">
<emphasis id="B922EAF7E314FFE91916F8C6FA3DF899" box="[1343,1436,1843,1865]" italics="true" pageId="26" pageNumber="293">P. riedelii</emphasis>
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<paragraph id="8BE936E5E314FFE91CBEF8A3FA3DF852" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF8A3FA3DF852" pageId="26" pageNumber="293"> Chelipeds setose; pollex without small tuft of setae at tip. Third maxilliped exopod 0.91.3 (male and female) times as long as ischium............................................................................................. 4</keyLead>
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</keyStep>
<keyStep id="30A22E40E314FFE91CBEF879FA3DF829" pageId="26" pageNumber="293">
<paragraph id="8BE936E5E314FFE91CBEF879FA3DF86F" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF879FA3DF86F" pageId="26" pageNumber="293">
4. Anterolateral margin of carapace with only faint lobe after orbital tooth (male and female). Third maxilliped exopod 1.21.3 (male), 1.0 (female) times as long as ischium. Endemic to
<collectingCountry id="F3417675E314FFE91F3BF85BFC37F86F" box="[786,918,1960,1982]" name="Futuna Island" pageId="26" pageNumber="293">Futuna Island</collectingCountry>
.............................
<taxonomicName id="4C564D66E314FFE918DAF85BFAEBF86C" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[1267,1354,1960,1982]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B922EAF7E314FFE918DAF85BFAEBF86C" box="[1267,1354,1960,1982]" italics="true" pageId="26" pageNumber="293">P. similis</emphasis>
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<emphasis id="B922EAF7E314FFE91979F85BFA3DF86F" bold="true" box="[1360,1436,1961,1982]" pageId="26" pageNumber="293">
<taxonomicNameLabel id="A211578CE314FFE91979F85BFA3DF86F" box="[1360,1436,1961,1982]" pageId="26" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BE936E5E314FFE91CBEF834FA3DF829" blockId="26.[151,1437,1579,2041]" pageId="26" pageNumber="293">
<keyLead id="30A795D0E314FFE91CBEF834FA3DF829" pageId="26" pageNumber="293">
Anterolateral margin of carapace with two distinct teeth after orbital tooth (male). Third maxilliped exopod 0.91.0 (male) times as long as ischium. Endemic to
<collectingCountry id="F3417675E314FFE91E64F816FD71F828" box="[589,720,2019,2041]" name="Futuna Island" pageId="26" pageNumber="293">Futuna Island</collectingCountry>
..........................................
<taxonomicName id="4C564D66E314FFE918E2F816FAEBF829" authorityName="Mazancourt &amp; Mazel &amp; Marquet &amp; Poupin &amp; Keith" authorityYear="2024" box="[1227,1354,2019,2041]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="293" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="B922EAF7E314FFE918E2F816FAEBF829" box="[1227,1354,2019,2041]" italics="true" pageId="26" pageNumber="293">P. ngankeeae</emphasis>
</taxonomicName>
<emphasis id="B922EAF7E314FFE91979F811FA3DF829" bold="true" box="[1360,1436,2019,2040]" pageId="26" pageNumber="293">
<taxonomicNameLabel id="A211578CE314FFE91979F811FA3DF829" box="[1360,1436,2019,2040]" pageId="26" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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