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<document id="433766D0B5701D8B5AC4239948273FE1" ID-DOI="10.11646/zootaxa.5560.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14595436" ID-ZooBank="0F942970-010E-4775-856E-31CA016DAD50" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735931741530" checkinUser="plazi" docAuthor="Haddad, Charles Richard, Wiśniewski, Konrad &amp; Wesołowska, Wanda" docDate="2024" docId="03FF87E70C3F007DDAE385B6B020FEF5" docLanguage="en" docName="zootaxa.5560.1.1.pdf" docOrigin="Zootaxa 5560 (1)" docSource="https://doi.org/10.11646/zootaxa.5560.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Heliophanus difficilis Wesolowska 1986" docType="treatment" docVersion="2" lastPageNumber="24" masterDocId="FFC6FF9F0C29006ADA748055B129FF90" masterDocTitle="The jumping spiders of Mozambique (Araneae: Salticidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="23" updateTime="1735934017513" updateUser="ExternalLinkService">
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<mods:title id="04F62DBD7DC7FF91BC355800795D8F97">The jumping spiders of Mozambique (Araneae: Salticidae)</mods:title>
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<mods:namePart id="76E4F16258069BE44FBD79F70BC9FA6C">Haddad, Charles Richard</mods:namePart>
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<mods:affiliation id="45F7D4BAA0EF3F11C5B3ABD4A218EA0C">haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760</mods:affiliation>
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<mods:namePart id="FD66D47EB12C74A3E5E8D637F7DCAA03">Wiśniewski, Konrad</mods:namePart>
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<mods:affiliation id="DE42DA947310E67D4881CBCE60E68E5E">Institute of Biology, Pomeranian University in Słupsk, Arciszewskiego 22 a, 76 - 200 Słupsk, Poland &amp; konrad. wisniewski. araneae @ gmail. com; https: // orcid. org / 0000 - 0002 - 6780 - 3292</mods:affiliation>
<mods:nameIdentifier id="6B5C03A5BA85548FCAF9D96F7A1AF571" type="email">konrad.wisniewski.araneae@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A98510FD924E40CED682D3B38F26DC3D">Wesołowska, Wanda</mods:namePart>
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<mods:affiliation id="8F3A69D2BD679D9F5572E8D3D4B95321">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa &amp; Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland &amp; Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa &amp; wanda. h. wesolowska @ gmail. com; https: // orcid. org / 0000 - 0002 - 4411 - 1058 * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="ECDE9E37ABCD09E615F07A6162AF60E5" type="email">wanda.h.wesolowska@gmail.com</mods:nameIdentifier>
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<mods:date id="4CF05FDC1B1F9FB8860DA89612077182">2024</mods:date>
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<subSubSection id="C34C657A0C3F007CDAE385B6B3A1FA6D" box="[151,648,1507,1534]" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BE936F10C3F007CDAE385B6B3A1FA6D" blockId="22.[151,648,1507,1569]" box="[151,648,1507,1534]" pageId="22" pageNumber="23">
<heading id="D0A1819D0C3F007CDAE385B6B3A1FA6D" bold="true" box="[151,648,1507,1534]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
<taxonomicName id="4C564D720C3F007CDAE385B6B3A1FA6D" ID-CoL="3KBM8" authority="Wesolowska, 1986" authorityName="Wesolowska" authorityYear="1986" box="[151,648,1507,1534]" class="Arachnida" family="Salticidae" genus="Heliophanus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="B922EAE30C3F007CDAE385B6B3A1FA6D" bold="true" box="[151,648,1507,1534]" pageId="22" pageNumber="23">
<emphasis id="B922EAE30C3F007CDAE385B6B0B7FA6E" bold="true" box="[151,414,1507,1534]" italics="true" pageId="22" pageNumber="23">Heliophanus difficilis</emphasis>
<bibRefCitation id="EFC74B000C3F007CDBD185B6B3A1FA6D" author="Wesolowska, W." box="[421,648,1507,1533]" pageId="22" pageNumber="23" pagination="1 - 254" refId="ref48372" refString="Wesolowska, W. (1986) A revision of the genus Heliophanus C. L. Koch, 1833 (Aranei: Salticidae). Annales Zoologici, Warszawa, 40, 1 - 254." type="journal article" year="1986">Wesołowska, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C34C657A0C3F007CDAE38652B1CEF9B1" box="[151,231,1543,1569]" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE936F10C3F007CDAE38652B1CEF9B1" blockId="22.[151,648,1507,1569]" box="[151,231,1543,1569]" pageId="22" pageNumber="23">
<figureCitation id="136D2A740C3F007CDAE38652B1CEF9B1" box="[151,231,1543,1569]" captionStart="FIGURE 14" captionStartId="23.[151,250,1694,1718]" captionTargetBox="[174,1436,450,1651]" captionTargetId="figure-77@23.[151,1436,450,1668]" captionTargetPageId="23" captionText="FIGURE 14. Heliophanus difficilis Wesołowska, 1986, female. A. General appearance, dorsal view; B. Same, lateral view; C. Epigyne; D. Internal structure of epigyne." figureDoi="http://doi.org/10.5281/zenodo.14595464" httpUri="https://zenodo.org/record/14595464/files/figure.png" pageId="22" pageNumber="23">Fig. 14</figureCitation>
</paragraph>
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<subSubSection id="C34C657A0C3F007CDAE38619B205F9F4" box="[151,812,1611,1636]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8BE936F10C3F007CDAE38619B205F9F4" blockId="22.[151,812,1611,1636]" box="[151,812,1611,1636]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4611DF0C3F007CDAE38619B205F9F4" box="[151,812,1611,1636]" pageId="22" pageNumber="23">
<treatmentCitation id="0AF710E00C3F007CDAE38619B345F9F3" author="Wesolowska, W." box="[151,620,1611,1636]" page="226" pageId="22" pageNumber="23" year="1986">
<taxonomicName id="4C564D720C3F007CDAE38619B345F9F3" ID-CoL="3KBM8" authority="Wesolowska, 1986: 226" authorityName="Wesolowska" authorityPageNumber="226" authorityYear="1986" box="[151,620,1611,1636]" class="Arachnida" family="Salticidae" genus="Heliophanus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="B922EAE30C3F007CDAE38619B05BF9F4" box="[151,370,1612,1636]" italics="true" pageId="22" pageNumber="23">Heliophanus difficilis</emphasis>
<bibRefCitation id="EFC74B000C3F007CDB0C8619B345F9F3" author="Wesolowska, W." box="[376,620,1611,1636]" pageId="22" pageNumber="23" pagination="1 - 254" refId="ref48372" refString="Wesolowska, W. (1986) A revision of the genus Heliophanus C. L. Koch, 1833 (Aranei: Salticidae). Annales Zoologici, Warszawa, 40, 1 - 254." type="journal article" year="1986">Wesołowska, 1986: 226</bibRefCitation>
</taxonomicName>
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, figs 825827 (
<specimenCount id="9D50FD780C3F007CD9788619B234F9F4" box="[780,797,1612,1636]" count="1" pageId="22" pageNumber="23" type="female"></specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C657A0C3F007CDAE386C4B09CF940" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BE936F10C3F007CDAE386C4B09CF940" blockId="22.[151,1437,1681,2032]" pageId="22" pageNumber="23">
<emphasis id="B922EAE30C3F007CDAE386C4B03BF93B" bold="true" box="[151,274,1681,1707]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
The female can be distinguished from congeners by the trajectory of the seminal ducts, which form a full loop. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34C657A0C3F007CDAB3868CB540F964" box="[199,1129,1753,1780]" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BE936F10C3F007CDAB3868CB540F964" blockId="22.[151,1437,1681,2032]" box="[199,1129,1753,1780]" pageId="22" pageNumber="23">
<emphasis id="B922EAE30C3F007CDAB3868CB098F963" bold="true" box="[199,433,1753,1779]" pageId="22" pageNumber="23">Material examined.</emphasis>
<materialsCitation id="3B3E3CAC0C3F007CDBC3868CB54CF964" box="[439,1125,1753,1780]" collectionCode="SMF" country="Mozambique" location="Tete" pageId="22" pageNumber="23" specimenCode="SMF 10368" specimenCount="1" specimenCount-female="1" stateProvince="Tete">
<emphasis id="B922EAE30C3F007CDBC3868CB3E5F964" bold="true" box="[439,716,1753,1780]" pageId="22" pageNumber="23">
<collectingCountry id="F34176610C3F007CDBC3868CB3AEF963" box="[439,647,1753,1779]" name="Mozambique" pageId="22" pageNumber="23">MOZAMBIQUE</collectingCountry>
:
<emphasis id="B922EAE30C3F007CD8E0868FB3EAF964" bold="true" box="[660,707,1754,1780]" italics="true" pageId="22" pageNumber="23">
<collectingRegion id="4992F8130C3F007CD8E0868FB3EAF964" box="[660,707,1754,1780]" country="Mozambique" name="Tete" pageId="22" pageNumber="23">Tete</collectingRegion>
</emphasis>
:
</emphasis>
no further details,
<specimenCount id="9D50FD780C3F007CD9D4868FB2EAF964" box="[928,963,1754,1780]" count="1" pageId="22" pageNumber="23" type="female">1♀</specimenCount>
(
<specimenCode id="DBF09E8A0C3F007CD9A6868FB577F964" box="[978,1118,1754,1780]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="22" pageNumber="23">SMF 10368</specimenCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C34C657A0C3F007DDAB386A8B39BFF45" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE936F10C3F007DDAB386A8B39BFF45" blockId="22.[151,1437,1681,2032]" lastBlockId="23.[151,1436,151,357]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="B922EAE30C3F007CDAB386A8B0F5F888" bold="true" box="[199,476,1789,1816]" pageId="22" pageNumber="23">Redescription. Female:</emphasis>
Measurements. Cephalothorax length 1.8, width 1.3, height 0.7. Abdomen length 1.9, width 1.6. Eye field length 0.7, anterior and posterior width 1.2. General appearance as in
<figureCitation id="136D2A740C3F007CDEEA8777B401F8AC" box="[1182,1320,1826,1852]" captionStart="FIGURE 14" captionStartId="23.[151,250,1694,1718]" captionTargetBox="[174,1436,450,1651]" captionTargetId="figure-77@23.[151,1436,450,1668]" captionTargetPageId="23" captionText="FIGURE 14. Heliophanus difficilis Wesołowska, 1986, female. A. General appearance, dorsal view; B. Same, lateral view; C. Epigyne; D. Internal structure of epigyne." figureDoi="http://doi.org/10.5281/zenodo.14595464" httpUri="https://zenodo.org/record/14595464/files/figure.png" pageId="22" pageNumber="23">Fig. 14A, B</figureCitation>
. Carapace dark brown, eye field almost black. Some long brown bristles near anterior row of eyes, sparse faint white hairs on anterior part of eye field. White hairs form thin belt on lateral slopes running along eye field and reaching carapace dorsum. Mouthparts and sternum brown. Abdomen brownish-grey, with thin yellow line on anterior edge, dorsum speckled with bright dots, with yellowish median streak and pair of large round white spots near midpoint. Venter variegated, lighter medially. Spinnerets grey. Legs yellow, only femora brown. Epigyne with shallow oval depression (
<figureCitation id="136D2A740C3E007DDB5680C2B0A5FF21" box="[290,396,151,177]" captionStart="FIGURE 14" captionStartId="23.[151,250,1694,1718]" captionTargetBox="[174,1436,450,1651]" captionTargetId="figure-77@23.[151,1436,450,1668]" captionTargetPageId="23" captionText="FIGURE 14. Heliophanus difficilis Wesołowska, 1986, female. A. General appearance, dorsal view; B. Same, lateral view; C. Epigyne; D. Internal structure of epigyne." figureDoi="http://doi.org/10.5281/zenodo.14595464" httpUri="https://zenodo.org/record/14595464/files/figure.png" pageId="23" pageNumber="24">Fig. 14C</figureCitation>
). Strongly sclerotized atria hiding copulatory openings, seminal ducts form loop laterally, spermathecae question mark-shaped (
<figureCitation id="136D2A740C3E007DD84880EEB38CFF45" box="[572,677,187,213]" captionStart="FIGURE 14" captionStartId="23.[151,250,1694,1718]" captionTargetBox="[174,1436,450,1651]" captionTargetId="figure-77@23.[151,1436,450,1668]" captionTargetPageId="23" captionText="FIGURE 14. Heliophanus difficilis Wesołowska, 1986, female. A. General appearance, dorsal view; B. Same, lateral view; C. Epigyne; D. Internal structure of epigyne." figureDoi="http://doi.org/10.5281/zenodo.14595464" httpUri="https://zenodo.org/record/14595464/files/figure.png" pageId="23" pageNumber="24">Fig. 14D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C657A0C3E007DDAB3808BB07FFE8D" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BE936F10C3E007DDAB3808BB07FFE8D" blockId="23.[151,1436,151,357]" pageId="23" pageNumber="24">
<emphasis id="B922EAE30C3E007DDAB3808BB048FF68" bold="true" box="[199,353,222,248]" pageId="23" pageNumber="24">Distribution.</emphasis>
The species was previously only known from the D.R.
<collectingCountry id="F34176610C3E007DD9BE808AB53CFF69" box="[970,1045,223,249]" name="Democratic Republic of the Congo" pageId="23" pageNumber="24">Congo</collectingCountry>
and is recorded from
<collectingCountry id="F34176610C3E007DDF7C808AB4B5FF69" box="[1288,1436,223,249]" name="Mozambique" pageId="23" pageNumber="24">Mozambique</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C34C657A0C3E007DDAB38172B020FEF5" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BE936F10C3E007DDAB38172B020FEF5" blockId="23.[151,1436,151,357]" pageId="23" pageNumber="24">
<emphasis id="B922EAE30C3E007DDAB38172B012FED1" bold="true" box="[199,315,295,321]" pageId="23" pageNumber="24">Remarks.</emphasis>
This species was only known from the female
<typeStatus id="54ED88530C3E007DD9F98172B2DBFED1" box="[909,1010,295,321]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, so this is only the second known specimen.
</paragraph>
</subSubSection>
</treatment>
</document>