treatments-xml/data/03/FF/87/03FF87BCFFAAEA19FF34F9DFFE9EFA9F.xml

<document id="A5FB401E1F0576DDB6DC156D8ECBAFF7" ID-CLB-Dataset="25537" ID-DOI="10.11646/zootaxa.4648.1.2" ID-GBIF-Dataset="2a809cd9-f17a-4eea-8eb0-ad8f2ee0ef91" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353871" ID-ZooBank="ECE2A8C4-9CAA-4580-B589-D693C2F3EEB6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564383872264" checkinUser="plazi" docAuthor="Cruz, Carlos Alberto Gonçalves, Caramaschi, Ulisses, Fusinatto, Luciana Ardenghi &amp; Brasileiro, Cinthia Aguirre" docDate="2019" docId="03FF87BCFFAAEA19FF34F9DFFE9EFA9F" docLanguage="en" docName="zootaxa.4648.1.2.pdf" docOrigin="Zootaxa 4648 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dendrophryniscus imitator Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro 2019, new status" docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFC6FFC4FFADEA12FFA3FFF4FFE5FFB4" masterDocTitle="Taxonomic review of Dendrophryniscus brevipollicatus Jiménez de la Espada, 1870, with revalidation of D. imitator (Miranda-Ribeiro, 1920) and D. lauroi Miranda-Ribeiro, 1926, and description of four new related species (Anura, Bufonidae)" masterLastPageNumber="62" masterPageNumber="27" pageNumber="34" updateTime="1698747731810" updateUser="ExternalLinkService">
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<mods:title id="CA0D8B13C4A81DED96640BE49BCBD18C">Taxonomic review of Dendrophryniscus brevipollicatus Jiménez de la Espada, 1870, with revalidation of D. imitator (Miranda-Ribeiro, 1920) and D. lauroi Miranda-Ribeiro, 1926, and description of four new related species (Anura, Bufonidae)</mods:title>
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<mods:namePart id="D60E45EA1D8066EABBFF34A25F8CD907">Cruz, Carlos Alberto Gonçalves</mods:namePart>
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<mods:namePart id="25D06F8DA910C76413A2227504B6B214">Fusinatto, Luciana Ardenghi</mods:namePart>
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<taxonomicName id="4C564D29FFAAEA15FF34F9DFFCFFF9F1" ID-CoL="J8YW" authority="(Miranda-Ribeiro, 1920)" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[151,794,1579,1606]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="imitator" status="stat. nov.">
<emphasis id="B922EAB8FFAAEA15FF34F9DFFE3BF9F2" bold="true" box="[151,478,1579,1606]" italics="true" pageId="7" pageNumber="34">Dendrophryniscus imitator</emphasis>
(
<bibRefCitation id="EFC74B5BFFAAEA15FE4EF9DFFCF4F9F1" author="Miranda-Ribeiro, A." box="[493,785,1579,1605]" pageId="7" pageNumber="34" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
)
</taxonomicName>
, 
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, revalidated
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</paragraph>
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(
<figureCitation id="136D2A2FFFAAEA15FF3CF9A4FEECF9DE" box="[159,265,1615,1642]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="8.[151,250,1459,1484]" captionStartId-1="9.[151,250,1928,1953]" captionStartId-2="10.[151,250,1967,1992]" captionTargetBox-0="[151,1437,561,1440]" captionTargetBox-1="[151,1436,1137,1908]" captionTargetBox-2="[302,1285,885,1947]" captionTargetId-0="figure@8.[151,1437,561,1440]" captionTargetId-1="figure@9.[151,1436,1137,1908]" captionTargetId-2="figure@10.[302,1285,885,1947]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 3. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), lectotype, MZUSP 847 (SVL 21.5 mm). Dorsal and ventral views." captionText-1="FIGURE 4. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and ventral views." captionText-2="FIGURE 5. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and lateral views of head, ventral views of hand and foot." httpUri-0="https://zenodo.org/record/3353877/files/figure.png" httpUri-1="https://zenodo.org/record/3353879/files/figure.png" httpUri-2="https://zenodo.org/record/3353881/files/figure.png" pageId="7" pageNumber="34">Figs. 3–5</figureCitation>
)
</paragraph>
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<subSubSection id="C34C6521FFAAEA15FF34F979FABBF957" pageId="7" pageNumber="34" type="reference_group">
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<taxonomicName id="4C564D29FFAAEA15FF34F979FDDEF911" ID-CoL="J8YW" authority="Miranda-Ribeiro, 1920" authorityName="Miranda-Ribeiro" authorityYear="1920" box="[151,571,1677,1702]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFAAEA15FF34F979FEADF911" box="[151,328,1677,1701]" italics="true" pageId="7" pageNumber="34">Atelopus imitator</emphasis>
<bibRefCitation id="EFC74B5BFFAAEA15FEEDF979FDDEF911" author="Miranda-Ribeiro, A." box="[334,571,1677,1702]" pageId="7" pageNumber="34" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE936AAFFAAEA15FF34F958FCADF970" blockId="7.[151,1374,1677,1763]" box="[151,840,1707,1732]" pageId="7" pageNumber="34">
<emphasis id="B922EAB8FFAAEA15FF34F958FDB8F970" box="[151,605,1708,1732]" italics="true" pageId="7" pageNumber="34">
<taxonomicName id="4C564D29FFAAEA15FF34F958FE0AF970" ID-CoL="6CL67" authorityName="Jimenez de la Espada" authorityYear="1870" box="[151,495,1708,1732]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="brevipollicatus">Dendrophryniscus brevipollicatus</taxonomicName>
imitator—
</emphasis>
<bibRefCitation id="EFC74B5BFFAAEA15FDFEF95FFCA1F970" author="Miranda-Ribeiro, A." box="[605,836,1707,1732]" pageId="7" pageNumber="34" pagination="1 - 227" refId="ref22228" refString="Miranda-Ribeiro, A. (1926) Notas para servirem ao estudo dos gymnobatrachios (Anura) brasileiros. Arquivos do Museu Nacional, 27, 1 - 227, 22 pls." type="book chapter" year="1926">Miranda-Ribeiro 1926</bibRefCitation>
.
</paragraph>
<paragraph id="8BE936AAFFAAEA15FF34F93FFABBF957" blockId="7.[151,1374,1677,1763]" box="[151,1374,1738,1763]" pageId="7" pageNumber="34">
<emphasis id="B922EAB8FFAAEA15FF34F93FFDE3F957" box="[151,518,1739,1763]" italics="true" pageId="7" pageNumber="34">
<taxonomicName id="4C564D29FFAAEA15FF34F93FFEB4F957" ID-CoL="4346" authorityName="Jimenez de la Espada" authorityYear="1870" box="[151,337,1739,1763]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="genus">Dendrophryniscus</taxonomicName>
brevipollicatus—
</emphasis>
<bibRefCitation id="EFC74B5BFFAAEA15FDA6F93EFD71F957" author="Cochran, D. M." box="[517,660,1738,1763]" pageId="7" pageNumber="34" pagination="1 - 423" refId="ref21045" refString="Cochran, D. M. (1955 [1954]) Frogs of Southeastern Brazil. Bulletin of the United States National Museum, 206, 1 - 423. https: // doi. org / 10.5479 / si. 03629236.206.1" type="journal article" year="1955">Cochran 1955</bibRefCitation>
(part); 
<bibRefCitation id="EFC74B5BFFAAEA15FD43F93FFC76F957" author="Heyer, W. R. &amp; Rand, A. S. &amp; Cruz, C. A. G. &amp; Peixoto, O. L. &amp; Nelson, C. E." box="[736,915,1739,1763]" pageId="7" pageNumber="34" pagination="231 - 410" refId="ref21468" refString="Heyer, W. R., Rand, A. S., Cruz, C. A. G., Peixoto, O. L. &amp; Nelson, C. E. (1990) Frogs of Boraceia. Arquivos de Zoologia, 31, 231 - 410." type="journal article" year="1990">
Heyer 
<emphasis id="B922EAB8FFAAEA15FC87F93FFCBDF957" box="[804,856,1739,1763]" italics="true" pageId="7" pageNumber="34">et al.</emphasis>
1990
</bibRefCitation>
; 
<bibRefCitation id="EFC74B5BFFAAEA15FC3EF93EFBD8F957" author="Izecksohn, E." box="[925,1085,1738,1763]" pageId="7" pageNumber="34" pagination="473 - 488" refId="ref21619" refString="Izecksohn, E. (1994 [1993]) Tres novas especies de Dendrophryniscus Jimenez de la Espada das regioes sudeste e sul do Brasil (Amphibia, Anura, Bufonidae). Revista Brasileira de Zoologia, 10, 473 - 488. https: // doi. org / 10.1590 / S 0101 - 81751993000300015" type="journal article" year="1994">Izecksohn 1994</bibRefCitation>
(part); 
<bibRefCitation id="EFC74B5BFFAAEA15FB2AF93EFABFF957" author="Malagoli, L. R. &amp; Trevine, V. &amp; Condez, T. H. &amp; Centeno, F. C. &amp; Berneck, B. M. &amp; Haddad, C. F. B." box="[1161,1370,1738,1763]" pageId="7" pageNumber="34" pagination="31 - 39" refId="ref21946" refString="Malagoli, L. R., Trevine, V., Condez, T. H., Centeno, F. C., Berneck, B. M. &amp; Haddad, C. F. B. (2017) Notes on the breeding behaviour of the Neotropical toadlet Dendrophryniscus brevipollicatus (Anura: Bufonidae), a bromeliad phytotelmata specialist. Herpetology Notes, 10, 31 - 39." type="journal article" year="2017">
Malagoli 
<emphasis id="B922EAB8FFAAEA15FB48F93FFAFBF957" box="[1259,1310,1739,1763]" italics="true" pageId="7" pageNumber="34">et al.</emphasis>
2017
</bibRefCitation>
.
</paragraph>
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<paragraph id="8BE936AAFFAAEA15FF34F8FEFB6CF8FC" blockId="7.[151,1436,1801,2045]" pageId="7" pageNumber="34">
<materialsCitation id="3B3E3CF7FFAAEA15FF34F8FEFB6CF8FC" ID-GBIF-Occurrence="2413160014" collectionCode="MZUSP" collectorName="da Serra" country="Brazil" county="Bicego" elevation="84800" latitude="-23.766666" location="Alto da Serra" longLatPrecision="1254" longitude="-46.316666" municipality="State" pageId="7" pageNumber="34" specimenCode="MZUSP 847" specimenCount="1" stateProvince="Sao Paulo" typeStatus="lectotype">
<emphasis id="B922EAB8FFAAEA15FF34F8FEFEF1F890" bold="true" box="[151,276,1802,1828]" pageId="7" pageNumber="34">
<typeStatus id="54ED8808FFAAEA15FF34F8FEFEEAF890" box="[151,271,1802,1828]" pageId="7" pageNumber="34" type="lectotype">Lectotype</typeStatus>
.
</emphasis>
<specimenCode id="DBF09ED1FFAAEA15FEBFF8FEFE49F890" box="[284,428,1802,1828]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP 847</specimenCode>
, adult female (
<figureCitation id="136D2A2FFFAAEA15FDF1F8FEFD73F890" box="[594,662,1802,1828]" captionStart="FIGURE 3" captionStartId="8.[151,250,1459,1484]" captionTargetBox="[151,1437,561,1440]" captionTargetId="figure@8.[151,1437,561,1440]" captionTargetPageId="8" captionText="FIGURE 3. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), lectotype, MZUSP 847 (SVL 21.5 mm). Dorsal and ventral views." httpUri="https://zenodo.org/record/3353877/files/figure.png" pageId="7" pageNumber="34">Fig. 3</figureCitation>
), collected at 
<location id="8E896071FFAAEA15FC90F8FEFC2DF890" LSID="urn:lsid:plazi:treatment:03FF87BCFFAAEA19FF34F9DFFE9EFA9F:8E896071FFAAEA15FC90F8FEFC2DF890" box="[819,968,1802,1828]" country="Brazil" county="Bicego" latitude="-23.766666" longLatPrecision="1254" longitude="-46.316666" municipality="State" name="Alto da Serra" pageId="7" pageNumber="34" stateProvince="Sao Paulo">
Alto 
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</location>
(
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23 
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46’S
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, 
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46 
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19’W
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, Datum WGS 
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altitude), 
<location id="8E896071FFAAEA15FEBCF8DAFD9FF8FC" LSID="urn:lsid:plazi:treatment:03FF87BCFFAAEA19FF34F9DFFE9EFA9F:8E896071FFAAEA15FEBCF8DAFD9FF8FC" box="[287,634,1838,1864]" country="Brazil" county="Bicego" latitude="-23.766666" longLatPrecision="1254" longitude="-46.316666" municipality="State" name="Municipality of Paranapiacaba" pageId="7" pageNumber="34" stateProvince="Sao Paulo">Municipality of Paranapiacaba</location>
, 
<collectingMunicipality id="6B8DACD0FFAAEA15FD26F8DAFD58F8FC" box="[645,701,1838,1864]" pageId="7" pageNumber="34">State</collectingMunicipality>
of 
<collectingRegion id="4992F848FFAAEA15FD41F8DAFCB3F8FC" box="[738,854,1838,1864]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="34">São Paulo</collectingRegion>
, 
<collectingCountry id="F341763AFFAAEA15FCC2F8DAFC4DF8FC" box="[865,936,1838,1864]" name="Brazil" pageId="7" pageNumber="34">Brazil</collectingCountry>
, by 
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in 1899.
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</paragraph>
<paragraph id="8BE936AAFFAAEA15FF64F8A6FCB0F849" blockId="7.[151,1436,1801,2045]" pageId="7" pageNumber="34">
<bibRefCitation id="EFC74B5BFFAAEA15FF64F8A6FE32F8D8" author="Miranda-Ribeiro, A." box="[199,471,1874,1900]" pageId="7" pageNumber="34" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro (1920)</bibRefCitation>
described 
<taxonomicName id="4C564D29FFAAEA15FDF7F8A6FCFDF8D8" authorityName="Miranda-Ribeiro" authorityYear="1920" box="[596,792,1874,1900]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFAAEA15FDF7F8A6FCFDF8D8" box="[596,792,1874,1900]" italics="true" pageId="7" pageNumber="34">Atelopus imitator</emphasis>
</taxonomicName>
based on five 
<typeStatus id="54ED8808FFAAEA15FC60F8A7FBC0F8D9" box="[963,1061,1875,1901]" pageId="7" pageNumber="34" type="syntype">syntypes</typeStatus>
(
<collectionCode id="ED47AE6FFFAAEA15FB96F8A6FB76F8D8" box="[1077,1171,1874,1900]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP</collectionCode>
477 and 485 from Cubatão, Santos; 
<collectionCode id="ED47AE6FFFAAEA15FE98F882FE72F824" box="[315,407,1910,1936]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP</collectionCode>
947, Alto da Serra; 
<collectionCode id="ED47AE6FFFAAEA15FDDBF882FD31F824" box="[632,724,1910,1936]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP</collectionCode>
835, Rio Grande; 
<collectionCode id="ED47AE6FFFAAEA15FC07F882FBE5F824" box="[932,1024,1910,1936]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP</collectionCode>
639, Campo Grande; all localities in the State of 
<collectingRegion id="4992F848FFAAEA15FEBFF86EFE77F801" box="[284,402,1946,1973]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="34">São Paulo</collectingRegion>
). However, he described and referred measurements for one specimen (
<emphasis id="B922EAB8FFAAEA15FB0EF86FFAABF800" box="[1197,1358,1946,1973]" italics="true" pageId="7" pageNumber="34">
Corpo 
<quantity id="4CAE9B4FFFAAEA15FB58F86EFAAFF800" box="[1275,1354,1946,1972]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="34" unit="mm" value="23.0">23 mm</quantity>
.
</emphasis>
, 
<emphasis id="B922EAB8FFAAEA15FAF8F86FFF5EF86C" italics="true" pageId="7" pageNumber="34">perna 28.</emphasis>
), which matches the specimen 
<collectionCode id="ED47AE6FFFAAEA15FD82F84AFD98F86C" box="[545,637,1982,2008]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="34">MZUSP</collectionCode>
847. This specimen is here designated 
<typeStatus id="54ED8808FFAAEA15FB9CF84AFB43F86C" box="[1087,1190,1982,2008]" pageId="7" pageNumber="34" type="lectotype">lectotype</typeStatus>
for 
<taxonomicName id="4C564D29FFAAEA15FB7BF84AFE7CF848" authority="Miranda-Ribeiro, 1920" authorityName="Miranda-Ribeiro" authorityYear="1920" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFAAEA15FB7BF84AFA79F86C" box="[1240,1436,1982,2008]" italics="true" pageId="7" pageNumber="34">Atelopus imitator</emphasis>
<bibRefCitation id="EFC74B5BFFAAEA15FF34F816FE7CF848" author="Miranda-Ribeiro, A." box="[151,409,2018,2044]" pageId="7" pageNumber="34" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
</taxonomicName>
(currently 
<taxonomicName id="4C564D29FFAAEA15FDB5F816FCA0F848" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[534,837,2018,2044]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="34" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFAAEA15FDB5F816FCA0F848" box="[534,837,2018,2044]" italics="true" pageId="7" pageNumber="34">Dendrophryniscus imitator</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE936AAFFA5EA1AFF64FF6DFD3EFF63" blockId="8.[151,1436,153,503]" pageId="8" pageNumber="35">
Currently, the color of the 
<typeStatus id="54ED8808FFA5EA1AFE51FF6DFDBCFF07" box="[498,601,153,179]" pageId="8" pageNumber="35" type="lectotype">lectotype</typeStatus>
is fade because the time of preservation, and the body surfaces are homogeneous pale brown, the dorsal blotches not visible.
</paragraph>
<paragraph id="8BE936AAFFA5EA1AFF64FF16FAC4FF4F" blockId="8.[151,1436,153,503]" box="[199,1313,224,251]" pageId="8" pageNumber="35">
<emphasis id="B922EAB8FFA5EA1AFF64FF16FE1FFF4F" box="[199,506,225,251]" italics="true" pageId="8" pageNumber="35">
Measurements of 
<typeStatus id="54ED8808FFA5EA1AFE2EFF15FE13FF4F" box="[397,502,225,251]" pageId="8" pageNumber="35" type="lectotype">lectotype</typeStatus>
.
</emphasis>
SVL 21.5; HL 7.4; HW 6.5; ED 2.4; THL 8.0; TL 8.9; TAL 4.8; FL 7.4.
</paragraph>
<paragraph id="8BE936AAFFA5EA1AFF64FEF2FE9FFE43" blockId="8.[151,1436,153,503]" pageId="8" pageNumber="35">
<emphasis id="B922EAB8FFA5EA1AFF64FEF2FEA5FEAB" box="[199,320,262,287]" italics="true" pageId="8" pageNumber="35">Diagnosis.</emphasis>
The species is characterized by: (1) medium size for the genus (
<collectionCode id="ED47AE6FFFA5EA1AFBE2FEF1FB92FEAB" box="[1089,1143,261,287]" pageId="8" pageNumber="35">SVL</collectionCode>
<quantity id="4CAE9B4FFFA5EA1AFB22FEF0FACDFEAB" box="[1153,1320,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.7399999999999998" metricValueMin="1.51" pageId="8" pageNumber="35" unit="mm" value="16.25" valueMax="17.4" valueMin="15.1">15.1–17.4 mm</quantity>
in males, 
<quantity id="4CAE9B4FFFA5EA1AFF34FEDCFED8FEF7" box="[151,317,296,323]" metricMagnitude="-2" metricUnit="m" metricValue="1.795" metricValueMax="2.15" metricValueMin="1.44" pageId="8" pageNumber="35" unit="mm" value="17.95" valueMax="21.5" valueMin="14.4">14.4–21.5 mm</quantity>
in females); (2) body slender; (3) snout mucronate in dorsal view; (4) 
<emphasis id="B922EAB8FFA5EA1AFBDCFEDDFADBFEF7" box="[1151,1342,297,323]" italics="true" pageId="8" pageNumber="35">canthus rostralis</emphasis>
slightly curved; (5) presence of two pronounced granules posterior to the corner of mouth; (6) surfaces of upper eyelid with few granules sparse and external margin prominent; (7) tip of the third and fourth fingers poorly or not expanded laterally; (8) skinfold poor developed on the articulation of the first and second phalanges of the fingers II, III, and IV (9) male with moderate nuptial pad with minuscule horny unpigmented asperities on finger I; (10) fingers not fringed nor webbed.
</paragraph>
<caption id="DF296622FFA5EA1AFF34FA47FF3CFA44" httpUri="https://zenodo.org/record/3353877/files/figure.png" pageId="8" pageNumber="35" startId="8.[151,250,1459,1484]" targetBox="[151,1437,561,1440]" targetPageId="8">
<paragraph id="8BE936AAFFA5EA1AFF34FA47FF3CFA44" blockId="8.[151,1436,1459,1520]" pageId="8" pageNumber="35">
<emphasis id="B922EAB8FFA5EA1AFF34FA47FEF1FA78" bold="true" box="[151,276,1459,1484]" pageId="8" pageNumber="35">FIGURE 3.</emphasis>
<taxonomicName id="4C564D29FFA5EA1AFEB8FA40FCD7FA78" authority="(Miranda-Ribeiro, 1920)" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[283,818,1459,1484]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFEB8FA40FDCBFA78" box="[283,558,1460,1484]" italics="true" pageId="8" pageNumber="35">Dendrophryniscus imitator</emphasis>
(
<bibRefCitation id="EFC74B5BFFA5EA1AFD9FFA47FCCEFA78" author="Miranda-Ribeiro, A." box="[572,811,1459,1484]" pageId="8" pageNumber="35" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
)
</taxonomicName>
, lectotype, MZUSP 847 (SVL 21.5 mm). Dorsal and ventral views.
</paragraph>
</caption>
<paragraph id="8BE936AAFFA5EA1BFF64F9EBFCA8FDD0" blockId="8.[151,1437,1566,2025]" lastBlockId="9.[151,1437,153,1115]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
<emphasis id="B922EAB8FFA5EA1AFF64F9EBFC65F98C" box="[199,896,1566,1593]" italics="true" pageId="8" pageNumber="35">
Comparisons with other species. 
<taxonomicName id="4C564D29FFA5EA1AFDEEF9EAFC65F98C" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[589,896,1566,1592]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">Dendrophryniscus imitator</taxonomicName>
</emphasis>
is distinguished from 
<taxonomicName id="4C564D29FFA5EA1AFB2FF9EBFAA2F98C" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[1164,1351,1566,1592]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B922EAB8FFA5EA1AFB2FF9EBFAA2F98C" box="[1164,1351,1566,1592]" italics="true" pageId="8" pageNumber="35">D. proboscideus</emphasis>
</taxonomicName>
by the smaller size (
<collectionCode id="ED47AE6FFFA5EA1AFE93F9B6FE83F9E8" box="[304,358,1602,1628]" pageId="8" pageNumber="35">SVL</collectionCode>
<quantity id="4CAE9B4FFFA5EA1AFECDF9B6FDF1F9E9" box="[366,532,1602,1629]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.7399999999999998" metricValueMin="1.51" pageId="8" pageNumber="35" unit="mm" value="16.25" valueMax="17.4" valueMin="15.1">15.1–17.4 mm</quantity>
in males of 
<taxonomicName id="4C564D29FFA5EA1AFD04F9B7FCC3F9E8" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[679,806,1603,1628]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFD04F9B7FCC3F9E8" box="[679,806,1603,1628]" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
; SVL 
<quantity id="4CAE9B4FFFA5EA1AFCD6F9B6FBFFF9E9" box="[885,1050,1602,1629]" metricMagnitude="-2" metricUnit="m" metricValue="4.279999999999999" metricValueMax="4.64" metricValueMin="3.92" pageId="8" pageNumber="35" unit="mm" value="42.8" valueMax="46.4" valueMin="39.2">39.2–46.4 mm</quantity>
in males of 
<taxonomicName id="4C564D29FFA5EA1AFB0DF9B7FA83F9E8" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[1198,1382,1602,1628]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B922EAB8FFA5EA1AFB0DF9B7FA83F9E8" box="[1198,1382,1602,1628]" italics="true" pageId="8" pageNumber="35">D. proboscideus</emphasis>
</taxonomicName>
); 
<taxonomicName id="4C564D29FFA5EA1AFA22F9B7FF14F934" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFA22F9B7FF14F934" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA5EA1AFE52F993FD88F934" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[497,621,1638,1664]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA5EA1AFE52F993FD88F934" box="[497,621,1638,1664]" italics="true" pageId="8" pageNumber="35">D. krausae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C564D29FFA5EA1AFD06F993FCACF934" authorityName="Izecksohn" authorityYear="1994" box="[677,841,1638,1664]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA5EA1AFD06F993FCACF934" box="[677,841,1638,1664]" italics="true" pageId="8" pageNumber="35">D. stawiarskyi</emphasis>
</taxonomicName>
by the slender body with uniform shape (robust and enlarged posteriorly in those species); 
<taxonomicName id="4C564D29FFA5EA1AFDEBF97FFD20F910" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[584,709,1675,1700]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFDEBF97FFD20F910" box="[584,709,1675,1700]" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA5EA1AFC62F97FFB8AF910" authorityName="Izecksohn" authorityYear="1994" box="[961,1135,1674,1700]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="berthalutzae">
<emphasis id="B922EAB8FFA5EA1AFC62F97FFB8AF910" box="[961,1135,1674,1700]" italics="true" pageId="8" pageNumber="35">D. berthalutzae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFBDDF97FFAEEF910" authorityName="Izecksohn" authorityYear="1994" box="[1150,1291,1674,1700]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA5EA1AFBDDF97FFAEEF910" box="[1150,1291,1674,1700]" italics="true" pageId="8" pageNumber="35">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFAB9F97FFA73F910" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[1306,1430,1674,1700]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA5EA1AFAB9F97FFA73F910" box="[1306,1430,1674,1700]" italics="true" pageId="8" pageNumber="35">D. krausae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFF34F95BFF1BF97C" box="[151,254,1710,1736]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA5EA1AFF34F95BFF1BF97C" box="[151,254,1710,1736]" italics="true" pageId="8" pageNumber="35">D. lauroi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFEAEF95BFE99F97C" authorityName="Recoder, Teixeira, Cassimiro, Camacho &amp; Rodrigues" authorityYear="2010" box="[269,380,1711,1736]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="oreites">
<emphasis id="B922EAB8FFA5EA1AFEAEF95BFE99F97C" box="[269,380,1711,1736]" italics="true" pageId="8" pageNumber="35">D. oreites</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE28F95BFDA6F97C" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[395,579,1710,1736]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B922EAB8FFA5EA1AFE28F95BFDA6F97C" box="[395,579,1710,1736]" italics="true" pageId="8" pageNumber="35">D. proboscideus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFDF1F95BFD55F97C" box="[594,688,1710,1736]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="skuki">
<emphasis id="B922EAB8FFA5EA1AFDF1F95BFD55F97C" box="[594,688,1710,1736]" italics="true" pageId="8" pageNumber="35">D. skuki</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFD4CF95BFC76F97C" authorityName="Izecksohn" authorityYear="1994" box="[751,915,1710,1736]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA5EA1AFD4CF95BFC76F97C" box="[751,915,1710,1736]" italics="true" pageId="8" pageNumber="35">D. stawiarskyi</emphasis>
</taxonomicName>
by the snout mucronate in dorsal view (snout rounded in 
<taxonomicName id="4C564D29FFA5EA1AFEB6F927FE27F958" authorityName="Izecksohn" authorityYear="1994" box="[277,450,1746,1772]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="berthalutzae">
<emphasis id="B922EAB8FFA5EA1AFEB6F927FE27F958" box="[277,450,1746,1772]" italics="true" pageId="8" pageNumber="35">D. berthalutzae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE73F927FDB9F958" authorityName="Izecksohn" authorityYear="1994" box="[464,604,1746,1772]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA5EA1AFE73F927FDB9F958" box="[464,604,1746,1772]" italics="true" pageId="8" pageNumber="35">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFDCAF927FD2BF958" box="[617,718,1746,1772]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA5EA1AFDCAF927FD2BF958" box="[617,718,1746,1772]" italics="true" pageId="8" pageNumber="35">D. lauroi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFCA9F927FC4EF958" authorityName="Izecksohn" authorityYear="1994" box="[778,939,1746,1772]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA5EA1AFCA9F927FC4EF958" box="[778,939,1746,1772]" italics="true" pageId="8" pageNumber="35">D. stawiarskyi</emphasis>
</taxonomicName>
; snout truncate in dorsal view in 
<taxonomicName id="4C564D29FFA5EA1AFAB8F927FA73F958" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[1307,1430,1746,1772]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA5EA1AFAB8F927FA73F958" box="[1307,1430,1746,1772]" italics="true" pageId="8" pageNumber="35">D. krausae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFF34F903FEE3F8A4" authorityName="Recoder, Teixeira, Cassimiro, Camacho &amp; Rodrigues" authorityYear="2010" box="[151,262,1783,1808]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="oreites">
<emphasis id="B922EAB8FFA5EA1AFF34F903FEE3F8A4" box="[151,262,1783,1808]" italics="true" pageId="8" pageNumber="35">D. oreites</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFEE4F903FE1BF8A4" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[327,510,1782,1808]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B922EAB8FFA5EA1AFEE4F903FE1BF8A4" box="[327,510,1782,1808]" italics="true" pageId="8" pageNumber="35">D. proboscideus</emphasis>
</taxonomicName>
; snout long, narrow, spatulate, with parallel lateral borders and rounded tip in 
<taxonomicName id="4C564D29FFA5EA1AFA22F903FF34F880" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="skuki">
<emphasis id="B922EAB8FFA5EA1AFA22F903FF34F880" italics="true" pageId="8" pageNumber="35">D. skuki</emphasis>
</taxonomicName>
); by the 
<emphasis id="B922EAB8FFA5EA1AFE94F8EEFE16F880" box="[311,499,1818,1844]" italics="true" pageId="8" pageNumber="35">canthus rostralis</emphasis>
slightly curved, 
<taxonomicName id="4C564D29FFA5EA1AFD12F8EFFCCBF880" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[689,814,1819,1844]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFD12F8EFFCCBF880" box="[689,814,1819,1844]" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA5EA1AFB88F8EFFB72F880" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[1067,1175,1818,1844]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="davori" status="sp. nov.">
<emphasis id="B922EAB8FFA5EA1AFB88F8EFFB72F880" box="[1067,1175,1818,1844]" italics="true" pageId="8" pageNumber="35">D. davori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA5EA1AFB3DF8EFFB1CF881" box="[1182,1273,1819,1845]" pageId="8" pageNumber="35" rank="species">
<emphasis id="B922EAB8FFA5EA1AFB3DF8EFFB1CF881" bold="true" box="[1182,1273,1819,1845]" pageId="8" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C564D29FFA5EA1AFAA4F8EFFA70F880" authorityName="Izecksohn" authorityYear="1994" box="[1287,1429,1818,1844]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA5EA1AFAA4F8EFFA70F880" box="[1287,1429,1818,1844]" italics="true" pageId="8" pageNumber="35">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFF34F8CBFEF6F8EC" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[151,275,1854,1880]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA5EA1AFF34F8CBFEF6F8EC" box="[151,275,1854,1880]" italics="true" pageId="8" pageNumber="35">D. krausae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE82F8CBFE62F8EC" box="[289,391,1854,1880]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA5EA1AFE82F8CBFE62F8EC" box="[289,391,1854,1880]" italics="true" pageId="8" pageNumber="35">D. lauroi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFE64F8CBFD8CF8EC" authorityName="Izecksohn" authorityYear="1994" box="[455,617,1854,1880]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA5EA1AFE64F8CBFD8CF8EC" box="[455,617,1854,1880]" italics="true" pageId="8" pageNumber="35">D. stawiarskyi</emphasis>
</taxonomicName>
(
<emphasis id="B922EAB8FFA5EA1AFDD9F8CAFCD2F8EC" box="[634,823,1854,1880]" italics="true" pageId="8" pageNumber="35">canthus rostralis</emphasis>
straight); the presence of two white pronounced granules posterior to the corner of mouth distinguishes 
<taxonomicName id="4C564D29FFA5EA1AFD6FF897FCADF8C8" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[716,840,1891,1916]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFD6FF897FCADF8C8" box="[716,840,1891,1916]" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C564D29FFA5EA1AFC2FF897FC15F8C8" box="[908,1008,1890,1916]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA5EA1AFC2FF897FC15F8C8" box="[908,1008,1890,1916]" italics="true" pageId="8" pageNumber="35">D. lauroi</emphasis>
</taxonomicName>
(elliptical set of unpigmented shallow granules posterior to the corner of mouth), 
<taxonomicName id="4C564D29FFA5EA1AFDDDF873FCADF814" authorityName="Jimenez de la Espada" authorityYear="1870" box="[638,840,1926,1952]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA5EA1AFDDDF873FCADF814" box="[638,840,1926,1952]" italics="true" pageId="8" pageNumber="35">D. brevipollicatus</emphasis>
</taxonomicName>
(elliptical set of white pronounced granules), and 
<taxonomicName id="4C564D29FFA5EA1AFA22F873FEC6F870" authorityName="Izecksohn" authorityYear="1994" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="berthalutzae">
<emphasis id="B922EAB8FFA5EA1AFA22F873FEC6F870" italics="true" pageId="8" pageNumber="35">D. berthalutzae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE91F85FFE4AF870" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[306,431,1962,1988]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA5EA1AFE91F85FFE4AF870" box="[306,431,1962,1988]" italics="true" pageId="8" pageNumber="35">D. krausae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE1EF85FFD8EF870" authorityName="Izecksohn" authorityYear="1968" box="[445,619,1962,1988]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="B922EAB8FFA5EA1AFE1EF85FFD8EF870" box="[445,619,1962,1988]" italics="true" pageId="8" pageNumber="35">D. leucomystax</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFD08F85FFCFCF870" authorityName="Recoder, Teixeira, Cassimiro, Camacho &amp; Rodrigues" authorityYear="2010" box="[683,793,1963,1988]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="oreites">
<emphasis id="B922EAB8FFA5EA1AFD08F85FFCFCF870" box="[683,793,1963,1988]" italics="true" pageId="8" pageNumber="35">D. oreites</emphasis>
</taxonomicName>
(longitudinal set); 
<taxonomicName id="4C564D29FFA5EA1AFC51F85FFB64F870" authorityName="Izecksohn" authorityYear="1994" box="[1010,1153,1962,1988]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA5EA1AFC51F85FFB64F870" box="[1010,1153,1962,1988]" italics="true" pageId="8" pageNumber="35">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFB33F85FFAEAF870" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[1168,1295,1962,1988]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="haddadi" status="sp. nov.">
<emphasis id="B922EAB8FFA5EA1AFB33F85FFAEAF870" box="[1168,1295,1962,1988]" italics="true" pageId="8" pageNumber="35">D. haddadi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA5EA1AFAB4F85FFA97F871" box="[1303,1394,1963,1989]" pageId="8" pageNumber="35" rank="species">
<emphasis id="B922EAB8FFA5EA1AFAB4F85FFA97F871" bold="true" box="[1303,1394,1963,1989]" pageId="8" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C564D29FFA5EA1AFA22F85FFEF4F85C" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="organensis">
<emphasis id="B922EAB8FFA5EA1AFA22F85FFEF4F85C" italics="true" pageId="8" pageNumber="35">D. organensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE82F83BFE3DF85C" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[289,472,1998,2024]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="proboscideus">
<emphasis id="B922EAB8FFA5EA1AFE82F83BFE3DF85C" box="[289,472,1998,2024]" italics="true" pageId="8" pageNumber="35">D. proboscideus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA5EA1AFE4BF83BFDA3F85C" box="[488,582,1998,2024]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="skuki">
<emphasis id="B922EAB8FFA5EA1AFE4BF83BFDA3F85C" box="[488,582,1998,2024]" italics="true" pageId="8" pageNumber="35">D. skuki</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA5EA1AFD25F83BFCCCF85C" authorityName="Izecksohn" authorityYear="1994" box="[646,809,1998,2024]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA5EA1AFD25F83BFCCCF85C" box="[646,809,1998,2024]" italics="true" pageId="8" pageNumber="35">D. stawiarskyi</emphasis>
</taxonomicName>
(set absent); 
<taxonomicName id="4C564D29FFA5EA1AFC60F83BFBA4F85C" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[963,1089,1999,2024]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="35" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA5EA1AFC60F83BFBA4F85C" box="[963,1089,1999,2024]" italics="true" pageId="8" pageNumber="35">D. imitator</emphasis>
</taxonomicName>
presents the surfaces of upper eyelid with few granules sparse and external margin prominent (surfaces of upper eyelid with numerous granules densely distributed and with a conspicuous margin in 
<taxonomicName id="4C564D29FFA4EA1BFD52FF4AFC5FFF63" authorityName="Jimenez de la Espada" authorityYear="1870" box="[753,954,189,215]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA4EA1BFD52FF4AFC5FFF63" box="[753,954,189,215]" italics="true" pageId="9" pageNumber="36">D. brevipollicatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C564D29FFA4EA1BFC53FF4AFB6DFF63" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[1008,1160,189,215]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="izecksohni" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFC53FF4AFB6DFF63" box="[1008,1160,189,215]" italics="true" pageId="9" pageNumber="36">D. izecksohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFB2DFF49FB02FF63" box="[1166,1255,189,215]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFB2DFF49FB02FF63" bold="true" box="[1166,1255,189,215]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
; surfaces of upper eyelid with numerous granules with the external margin slightly prominent in 
<taxonomicName id="4C564D29FFA4EA1BFB83FF16FB60FF4F" box="[1056,1157,225,251]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA4EA1BFB83FF16FB60FF4F" box="[1056,1157,225,251]" italics="true" pageId="9" pageNumber="36">D. lauroi</emphasis>
</taxonomicName>
; surfaces of upper eyelid with few granules sparse and external margin conspicuous in 
<taxonomicName id="4C564D29FFA4EA1BFCEBFEF2FC51FEAB" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[840,948,261,287]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="davori" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFCEBFEF2FC51FEAB" box="[840,948,261,287]" italics="true" pageId="9" pageNumber="36">D. davori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFC18FEF1FBF1FEAB" box="[955,1044,261,287]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFC18FEF1FBF1FEAB" bold="true" box="[955,1044,261,287]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
); 
<taxonomicName id="4C564D29FFA4EA1BFB8FFEF2FB4CFEAB" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[1068,1193,262,287]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFB8FFEF2FB4CFEAB" box="[1068,1193,262,287]" italics="true" pageId="9" pageNumber="36">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA4EA1BFF34FEDEFE87FEF7" authorityName="Jimenez de la Espada" authorityYear="1870" box="[151,354,297,323]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA4EA1BFF34FEDEFE87FEF7" box="[151,354,297,323]" italics="true" pageId="9" pageNumber="36">D. brevipollicatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA4EA1BFED1FEDEFE3CFEF7" box="[370,473,297,323]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="lauroi">
<emphasis id="B922EAB8FFA4EA1BFED1FEDEFE3CFEF7" box="[370,473,297,323]" italics="true" pageId="9" pageNumber="36">D. lauroi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA4EA1BFDB9FEDEFD5DFEF7" box="[538,696,298,323]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="organensis">
<emphasis id="B922EAB8FFA4EA1BFDB9FEDEFD5DFEF7" box="[538,696,298,323]" italics="true" pageId="9" pageNumber="36">D. organensis</emphasis>
</taxonomicName>
by the tip of the third and fourth fingers poorly or not expanded laterally (laterally expanded in those species); 
<taxonomicName id="4C564D29FFA4EA1BFD00FEBAFCC5FED3" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[675,800,334,359]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFD00FEBAFCC5FED3" box="[675,800,334,359]" italics="true" pageId="9" pageNumber="36">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA4EA1BFBBFFEBAFB03FED3" authorityName="Jimenez de la Espada" authorityYear="1870" box="[1052,1254,333,359]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA4EA1BFBBFFEBAFB03FED3" box="[1052,1254,333,359]" italics="true" pageId="9" pageNumber="36">D. brevipollicatus</emphasis>
</taxonomicName>
by the presence of a skinfold poorly developed on the articulation of the first and second phalanges of the fingers II, III, and IV (well developed in this species); 
<taxonomicName id="4C564D29FFA4EA1BFE61FE62FDD8FE1B" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[450,573,406,431]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFE61FE62FDD8FE1B" box="[450,573,406,431]" italics="true" pageId="9" pageNumber="36">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA4EA1BFC90FE62FC78FE1B" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[819,925,405,431]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="davori" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFC90FE62FC78FE1B" box="[819,925,405,431]" italics="true" pageId="9" pageNumber="36">D. davori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFC00FE61FC1EFE1B" box="[931,1019,405,431]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFC00FE61FC1EFE1B" bold="true" box="[931,1019,405,431]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C564D29FFA4EA1BFBABFE62FB71FE1B" authorityName="Izecksohn" authorityYear="1994" box="[1032,1172,405,431]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA4EA1BFBABFE62FB71FE1B" box="[1032,1172,405,431]" italics="true" pageId="9" pageNumber="36">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA4EA1BFB02FE62FAD2FE1B" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[1185,1335,405,431]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="izecksohni" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFB02FE62FAD2FE1B" box="[1185,1335,405,431]" italics="true" pageId="9" pageNumber="36">D. izecksohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFA9EFE61FA70FE1B" box="[1341,1429,405,431]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFA9EFE61FA70FE1B" bold="true" box="[1341,1429,405,431]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C564D29FFA4EA1BFF34FE4EFEA7FE67" authorityName="Izecksohn" authorityYear="1968" box="[151,322,441,467]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="B922EAB8FFA4EA1BFF34FE4EFEA7FE67" box="[151,322,441,467]" italics="true" pageId="9" pageNumber="36">D. leucomystax</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C564D29FFA4EA1BFEDDFE4EFDC5FE67" authorityName="Izecksohn" authorityYear="1994" box="[382,544,441,467]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA4EA1BFEDDFE4EFDC5FE67" box="[382,544,441,467]" italics="true" pageId="9" pageNumber="36">D. stawiarskyi</emphasis>
</taxonomicName>
by the presence of moderate nuptial pad in male (absent in 
<taxonomicName id="4C564D29FFA4EA1BFB12FE4EFAB9FE67" authorityName="Izecksohn" authorityYear="1968" box="[1201,1372,441,467]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="B922EAB8FFA4EA1BFB12FE4EFAB9FE67" box="[1201,1372,441,467]" italics="true" pageId="9" pageNumber="36">D. leucomystax</emphasis>
</taxonomicName>
; very developed in 
<taxonomicName id="4C564D29FFA4EA1BFE8FFE2AFE72FE43" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[300,407,477,503]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="davori" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFE8FFE2AFE72FE43" box="[300,407,477,503]" italics="true" pageId="9" pageNumber="36">D. davori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFE3EFE29FE13FE43" box="[413,502,477,503]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFE3EFE29FE13FE43" bold="true" box="[413,502,477,503]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C564D29FFA4EA1BFDA0FE2AFD75FE43" authorityName="Izecksohn" authorityYear="1994" box="[515,656,477,503]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="carvalhoi">
<emphasis id="B922EAB8FFA4EA1BFDA0FE2AFD75FE43" box="[515,656,477,503]" italics="true" pageId="9" pageNumber="36">D. carvalhoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C564D29FFA4EA1BFD3EFE2AFCD6FE43" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[669,819,477,503]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="izecksohni" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFD3EFE2AFCD6FE43" box="[669,819,477,503]" italics="true" pageId="9" pageNumber="36">D. izecksohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFC9AFE29FC77FE43" box="[825,914,477,503]" pageId="9" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="4C564D29FFA4EA1BFC69FE2AFB89FE43" authorityName="Izecksohn" authorityYear="1994" box="[970,1132,477,503]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="stawiarskyi">
<emphasis id="B922EAB8FFA4EA1BFC69FE2AFB89FE43" box="[970,1132,477,503]" italics="true" pageId="9" pageNumber="36">D. stawiarskyi</emphasis>
</taxonomicName>
); by the fingers not fringed nor webbed, 
<taxonomicName id="4C564D29FFA4EA1BFE84FDF6FE47FDAF" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[295,418,514,539]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFE84FDF6FE47FDAF" box="[295,418,514,539]" italics="true" pageId="9" pageNumber="36">D. imitator</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C564D29FFA4EA1BFD3AFDF6FCE1FDAF" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[665,772,513,539]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="davori" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFD3AFDF6FCE1FDAF" box="[665,772,513,539]" italics="true" pageId="9" pageNumber="36">D. davori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFCA9FDF5FC86FDAF" box="[778,867,513,539]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFCA9FDF5FC86FDAF" bold="true" box="[778,867,513,539]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C564D29FFA4EA1BFC34FDF6FBBAFDAF" authorityName="Jimenez de la Espada" authorityYear="1870" box="[919,1119,513,539]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA4EA1BFC34FDF6FBBAFDAF" box="[919,1119,513,539]" italics="true" pageId="9" pageNumber="36">D. brevipollicatus</emphasis>
</taxonomicName>
(fingers slightly fringed and webbed only at base), 
<taxonomicName id="4C564D29FFA4EA1BFE35FDD2FDCAFD8B" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[406,559,549,575]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="izecksohni" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFE35FDD2FDCAFD8B" box="[406,559,549,575]" italics="true" pageId="9" pageNumber="36">D. izecksohni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFD95FDD1FD74FD8B" box="[566,657,549,575]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFD95FDD1FD74FD8B" bold="true" box="[566,657,549,575]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C564D29FFA4EA1BFD6AFDD2FCCBFD8B" authority="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro, 2019" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" box="[713,814,550,575]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="jureia" status="sp. nov.">
<emphasis id="B922EAB8FFA4EA1BFD6AFDD2FCCBFD8B" box="[713,814,550,575]" italics="true" pageId="9" pageNumber="36">D. jureia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157C3FFA4EA1BFC95FDD1FC74FD8B" box="[822,913,549,575]" pageId="9" pageNumber="36" rank="species">
<emphasis id="B922EAB8FFA4EA1BFC95FDD1FC74FD8B" bold="true" box="[822,913,549,575]" pageId="9" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
(fingers fringed and webbed about one third), and 
<taxonomicName id="4C564D29FFA4EA1BFF64FDBEFEA7FDD7" authorityName="Cruz &amp; Fusinatto" authorityYear="2008" box="[199,322,585,611]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="krausae">
<emphasis id="B922EAB8FFA4EA1BFF64FDBEFEA7FDD7" box="[199,322,585,611]" italics="true" pageId="9" pageNumber="36">D. krausae</emphasis>
</taxonomicName>
(fingers not fringed and webbed only at base).
</paragraph>
<paragraph id="8BE936AAFFA4EA1BFF64FD99FF04FD1F" blockId="9.[151,1437,153,1115]" pageId="9" pageNumber="36">
<emphasis id="B922EAB8FFA4EA1BFF64FD99FDBDFD33" box="[199,600,621,647]" italics="true" pageId="9" pageNumber="36">
Original description of 
<typeStatus id="54ED8808FFA4EA1BFE7EFD9AFDE9FD33" box="[477,524,622,647]" pageId="9" pageNumber="36">type</typeStatus>
series
</emphasis>
(
<taxonomicName id="4C564D29FFA4EA1BFDCFFD99FBA4FD33" authority="Miranda-Ribeiro 1920" authorityName="Miranda-Ribeiro" authorityYear="1920" box="[620,1089,621,647]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFDCFFD99FCD6FD33" box="[620,819,621,647]" italics="true" pageId="9" pageNumber="36">Atelopus imitator</emphasis>
<bibRefCitation id="EFC74B5BFFA4EA1BFC9CFD99FBA4FD33" author="Miranda-Ribeiro, A." box="[831,1089,621,647]" pageId="9" pageNumber="36" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro 1920</bibRefCitation>
</taxonomicName>
, freely translated from Portuguese)
</paragraph>
<paragraph id="8BE936AAFFA4EA1BFF64FD41FF08FBA7" blockId="9.[151,1437,153,1115]" pageId="9" pageNumber="36">
Elongated, with the humerus shorter than the transverse diameter of the neck. Snout compressed and depressed, with lateral nostrils near the tip of snout, 
<emphasis id="B922EAB8FFA4EA1BFDDAFD2DFCDDFD47" box="[633,824,729,755]" italics="true" pageId="9" pageNumber="36">canthus rostralis</emphasis>
evident, giving to loreal region a concave shape and continuing by the upper eyelid. Leg carried forward, the tarsal articulation almost reaching the posterior angle of the eye. Fingers as in 
<taxonomicName id="4C564D29FFA4EA1BFE2BFCD5FDE3FC8F" baseAuthorityName="Lichtenstein and Martens" baseAuthorityYear="1856" box="[392,518,801,827]" class="Amphibia" family="Bufonidae" genus="Atelopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="cruciger">
<emphasis id="B922EAB8FFA4EA1BFE2BFCD5FDE3FC8F" box="[392,518,801,827]" italics="true" pageId="9" pageNumber="36">A. cruciger</emphasis>
</taxonomicName>
; toes same, that is, connecting through a rudimentary membrane, between the first extension is in the order: 1, 2, 4 and 3, and between second 1, 2, 3, 5 and 4. Body more granular in larger individuals, smoother in the shorter. Color pale cinereous or yellowish. A wide transverse stripe between the eyes; and from the vertex of the triangle that is posterior leave two divergent arms backwards, one or two further back, a dark stripe, on the flanks, starting from the snout and finishing in the iliac articulation, bordered of white on its sides; members with transversal stripes. Sometimes a narrow vertebral line, white. Abdominal side dark marbled. Body 
<quantity id="4CAE9B4FFFA4EA1BFABDFC21FA8AFC5B" box="[1310,1391,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="36" unit="mm" value="23.0">23 mm</quantity>
, leg 
<quantity id="4CAE9B4FFFA4EA1BFF34FC0DFF0CFBA7" box="[151,233,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="36" unit="mm" value="28.0">28 mm</quantity>
.
</paragraph>
<paragraph id="8BE936AAFFA4EA1BFF64FBEAFD5DFBEF" blockId="9.[151,1437,153,1115]" pageId="9" pageNumber="36">
<emphasis id="B922EAB8FFA4EA1BFF64FBEAFE32FB83" box="[199,471,1053,1079]" italics="true" pageId="9" pageNumber="36">Description of topotype.</emphasis>
<collectionCode id="ED47AE6FFFA4EA1BFE7EFBE9FDDCFB83" box="[477,569,1053,1079]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="36">MZUSP</collectionCode>
103992, adult male collected in Paranapiacaba, State of 
<collectingRegion id="4992F848FFA4EA1BFB11FBE9FAC3FB83" box="[1202,1318,1053,1079]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="36">São Paulo</collectingRegion>
, on 
<date id="FFE8106AFFA4EA1BFAF0FBE9FF1CFBEF" pageId="9" pageNumber="36" value="1963-03-24">March 24, 1963</date>
by W.C.A. Bokermann and A.B. Costa.
</paragraph>
<caption id="DF296622FFA4EA1BFF34F87CFEE7F871" httpUri="https://zenodo.org/record/3353879/files/figure.png" pageId="9" pageNumber="36" startId="9.[151,250,1928,1953]" targetBox="[151,1436,1137,1908]" targetPageId="9">
<paragraph id="8BE936AAFFA4EA1BFF34F87CFEE7F871" blockId="9.[151,1436,1928,1989]" pageId="9" pageNumber="36">
<emphasis id="B922EAB8FFA4EA1BFF34F87CFEF6F815" bold="true" box="[151,275,1928,1953]" pageId="9" pageNumber="36">FIGURE 4.</emphasis>
<taxonomicName id="4C564D29FFA4EA1BFEB9F87DFCCAF815" authority="(Miranda-Ribeiro, 1920)" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[282,815,1928,1953]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="36" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA4EA1BFEB9F87DFDC8F814" box="[282,557,1929,1953]" italics="true" pageId="9" pageNumber="36">Dendrophryniscus imitator</emphasis>
(
<bibRefCitation id="EFC74B5BFFA4EA1BFD9AF87CFCCDF815" author="Miranda-Ribeiro, A." box="[569,808,1928,1953]" pageId="9" pageNumber="36" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
)
</taxonomicName>
, topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and ventral views.
</paragraph>
</caption>
<paragraph id="8BE936AAFFA7EA18FF64FF6DFC48FCEB" blockId="10.[151,1437,153,863]" pageId="10" pageNumber="37">
Body slender, elongated (
<figureCitation id="136D2A2FFFA7EA18FE44FF6DFDC8FF07" box="[487,557,153,179]" captionStart="FIGURE 4" captionStartId="9.[151,250,1928,1953]" captionTargetBox="[151,1436,1137,1908]" captionTargetId="figure@9.[151,1436,1137,1908]" captionTargetPageId="9" captionText="FIGURE 4. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and ventral views." httpUri="https://zenodo.org/record/3353879/files/figure.png" pageId="10" pageNumber="37">Fig. 4</figureCitation>
); head triangular, longer than large, head length 36.4% of SVL; snout mucronate in dorsal view, acute in lateral view (
<figureCitation id="136D2A2FFFA7EA18FDD0FF49FD5DFF63" box="[627,696,188,215]" captionStart="FIGURE 5" captionStartId="10.[151,250,1967,1992]" captionTargetBox="[302,1285,885,1947]" captionTargetId="figure@10.[302,1285,885,1947]" captionTargetPageId="10" captionText="FIGURE 5. Dendrophryniscus imitator (Miranda-Ribeiro, 1920), topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and lateral views of head, ventral views of hand and foot." httpUri="https://zenodo.org/record/3353881/files/figure.png" pageId="10" pageNumber="37">Fig. 5</figureCitation>
); snout 46.7% of head length; presence of a small tubercle under the tip of snout, nostrils not protuberant, small and elliptical, located laterally near the tip of snout, below the 
<emphasis id="B922EAB8FFA7EA18FAC9FF16FECEFEAB" italics="true" pageId="10" pageNumber="37">canthus rostralis</emphasis>
; internarial distance 48.2% of eye to snout distance; eye diameter 58.6% of eye to snout distance and 77.2% of interorbital distance; eye protuberant; 
<emphasis id="B922EAB8FFA7EA18FD1AFEDDFC93FEF7" box="[697,886,297,323]" italics="true" pageId="10" pageNumber="37">canthus rostralis</emphasis>
slightly curved; loreal region vertical; vocal sac indistinct; choanae small, circular, very far from each other; tongue long, narrow, and few enlarged posteriorly; vocal slits present. Arms robust, forearms more robust than upper arms; hand with fingers slender, not fringed nor webbed; distal end of the third and fourth fingers slightly expanded laterally; finger I enlarged, covered by moderate nuptial pad with minuscule horny unpigmented asperities; relative lengths of fingers I&lt;
<collectionCode id="ED47AE6FFFA7EA18FBD6FE4DFB6DFE67" box="[1141,1160,441,467]" pageId="10" pageNumber="37">II</collectionCode>
&lt;
<collectionCode id="ED47AE6FFFA7EA18FB3BFE4DFB52FE67" box="[1176,1207,441,467]" pageId="10" pageNumber="37">IV</collectionCode>
&lt;
<collectionCode id="ED47AE6FFFA7EA18FB64FE4DFB01FE67" box="[1223,1252,441,467]" pageId="10" pageNumber="37">III</collectionCode>
; subarticular tubercles single, rounded; outer metacarpal tubercle large, rounded; inner metacarpal tubercle rounded, slightly small than subarticular tubercles; supernumerary tubercles present, small; skinfold poor developed on the articulation of the first and second phalanges of the fingers II, III, and IV. Thigh length slightly shorter than tibia length; sum of thigh and tibia lengths about 93.5% of snout–vent length. Tarsal length about 23% of the snout–vent length. Foot with toes slender, not fringed; webbing formulae: 
<date id="FFE8106AFFA7EA18FD7AFD99FCE8FD33" box="[729,781,621,647]" pageId="10" pageNumber="37">I1–2</date>
<superScript id="7C239BE2FFA7EA18FCAEFD98FCF3FDCE" attach="right" box="[781,790,620,634]" fontSize="6" pageId="10" pageNumber="37">+</superScript>
II 
<superScript id="7C239BE2FFA7EA18FC8FFD99FCAFFDCE" attach="none" box="[812,842,620,647]" fontSize="6" pageId="10" pageNumber="37">11/2</superScript>
–3 
<date id="FFE8106AFFA7EA18FCC5FD99FC48FD33" box="[870,941,621,647]" pageId="10" pageNumber="37">III2–3</date>
IV 
<superScript id="7C239BE2FFA7EA18FC6FFD99FC08FDCE" attach="none" box="[972,1005,620,647]" fontSize="6" pageId="10" pageNumber="37">31/2</superScript>
– 
<date id="FFE8106AFFA7EA18FC5FFD99FBD1FD33" box="[1020,1076,620,647]" pageId="10" pageNumber="37" value="1921-05-02">
<superScript id="7C239BE2FFA7EA18FC5FFD99FBFBFDCE" attach="right" box="[1020,1054,620,647]" fontSize="6" pageId="10" pageNumber="37">21/2</superScript>
V
</date>
; distal end of toes globose, not expanded laterally, relative lengths of toes I&lt;
<collectionCode id="ED47AE6FFFA7EA18FD37FD65FD42FD1F" box="[660,679,657,683]" pageId="10" pageNumber="37">II</collectionCode>
&lt;
<collectionCode id="ED47AE6FFFA7EA18FD15FD65FD36FD1F" box="[694,723,657,683]" pageId="10" pageNumber="37">III</collectionCode>
&lt;V&lt;
<collectionCode id="ED47AE6FFFA7EA18FCA4FD65FCC3FD1F" box="[775,806,657,683]" pageId="10" pageNumber="37">IV</collectionCode>
; subarticular tubercles single, rounded, the same size of those of the fingers; outer metatarsal tubercle with the same size of subarticular tubercles, rounded; inner metatarsal tubercle large, approximately twice the outer, elongated; supernumerary tubercles present but scarce. Dorsal and lateral surfaces covered by spinulose granules uniformly distributed, with different sizes, ventral surfaces covered by numerous rounded granules; surfaces of upper eyelid with few granules sparse and external margin prominent; presence of two pronounced granules posterior to the corner of mouth.
</paragraph>
<caption id="DF296622FFA7EA18FF34F85BFD7CF858" httpUri="https://zenodo.org/record/3353881/files/figure.png" pageId="10" pageNumber="37" startId="10.[151,250,1967,1992]" targetBox="[302,1285,885,1947]" targetPageId="10">
<paragraph id="8BE936AAFFA7EA18FF34F85BFD7CF858" blockId="10.[151,1436,1967,2028]" pageId="10" pageNumber="37">
<emphasis id="B922EAB8FFA7EA18FF34F85BFEF1F87C" bold="true" box="[151,276,1967,1992]" pageId="10" pageNumber="37">FIGURE 5.</emphasis>
<taxonomicName id="4C564D29FFA7EA18FEB8F844FCD6F87C" authority="(Miranda-Ribeiro, 1920)" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[283,819,1967,1992]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="37" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA7EA18FEB8F844FDCAF87C" box="[283,559,1968,1992]" italics="true" pageId="10" pageNumber="37">Dendrophryniscus imitator</emphasis>
(
<bibRefCitation id="EFC74B5BFFA7EA18FD9EF85BFCC9F87C" author="Miranda-Ribeiro, A." box="[573,812,1967,1992]" pageId="10" pageNumber="37" pagination="307 - 316" refId="ref22184" refString="Miranda-Ribeiro, A. (1920) Os Brachycephalideos do Museu Paulista (com tres especies novas). Revista do Museu Paulista, 12, 307 - 316. https: // doi. org / 10.5962 / bhl. part. 22255" type="journal article" year="1920">Miranda-Ribeiro, 1920</bibRefCitation>
)
</taxonomicName>
, topotype, MZUSP 103992 (SVL 17.0 mm). Dorsal and lateral views of head, ventral views of hand and foot.
</paragraph>
</caption>
<paragraph id="8BE936AAFFA6EA19FF64FF6DFC14FED2" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FF6DFE58FF07" box="[199,445,153,179]" italics="true" pageId="11" pageNumber="38">Color in preservative.</emphasis>
The topotypes present dorsum pale brown; a head blotch extending on the upper eyelids merging with an ‘X’ shaped blotch on the scapular region and an inverse ‘Y’ shaped blotch on the sacral region, extending to the inguinal region; one transversal bar on forearms and one blotch on elbow; one transversal bar on thigh, tibia, and tarsus, and one blotch on knee and heel; a lateral stripe starting at the snout, passing on the eye and reaching the inguinal region, being narrower in the loreal region; all of the bars, blotches, and stripes are dark brown; ventral surface lighter than the dorsum, with disperse small blotches.
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FE84FEB1FE19" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FE84FE10FE3D" box="[199,501,367,393]" italics="true" pageId="11" pageNumber="38">Measurements of topotype.</emphasis>
SVL 17.0; HL 6.2; HW 4.8; IND 1.4; ESD 2.9; ED 1.7; IOD 2.2; THL 7.7; TL 8.2; TAL 3.9; FL 6.8.
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FE4CFD1BFD89" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FE4CFED2FE65" box="[199,311,440,465]" italics="true" pageId="11" pageNumber="38">Variation.</emphasis>
Specimens are congruent with respect to the morphological characters. Variation of measurements and descriptive statistics of males and females are presented in 
<tableCitation id="C6D40311FFA6EA19FCFAFE2EFC53FE41" box="[857,950,474,501]" captionStart="TABLE 2" captionStartId="26.[151,239,746,772]" captionTargetPageId="26" captionText="TABLE 2. Range and mean ± standard deviation (in mm) of the measurements of the species of Dendrophryniscus (abbreviations of the measurements explained in the Material and methods). Males." httpUri="http://table.plazi.org/id/DF296622FFB7EA08FF34FD1EFE98FC9C" pageId="11" pageNumber="38" tableUuid="DF296622FFB7EA08FF34FD1EFE98FC9C">Tables 2</tableCitation>
and 
<tableCitation id="C6D40311FFA6EA19FC4FFE2FFC19FE41" box="[1004,1020,475,501]" captionStart="TABLE 3" captionStartId="27.[151,239,728,754]" captionTargetBox="[165,2017,809,1457]" captionTargetPageId="27" captionText="TABLE 3. Range and mean ± standard deviation (in mm) of the measurements of the species of Dendrophryniscus (abbreviations of the measurements explained in the Material and methods). Females." pageId="11" pageNumber="38">3</tableCitation>
. Sexual dimorphism can be observed on SVL (larger in female), on arms (slightly more robust in male), finger I (robust in male), and nuptial pad (on finger I of male). Nuptial pad can be pigmented or not.
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FDB2FBCEFC82" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FDB2FD75FDD4" box="[199,656,582,608]" italics="true" pageId="11" pageNumber="38">
Distribution. 
<taxonomicName id="4C564D29FFA6EA19FEC3FDB2FD75FDD4" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[352,656,582,608]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="imitator">Dendrophryniscus imitator</taxonomicName>
</emphasis>
is known from the following municipalities (
<figureCitation id="136D2A2FFFA6EA19FB34FDB2FB0FFDD4" box="[1175,1258,582,608]" captionStart="FIGURE 17" captionStartId="28.[151,250,1983,2008]" captionTargetBox="[151,1347,1124,1966]" captionTargetId="figure@28.[151,1347,1124,1966]" captionTargetPageId="28" captionText="FIGURE 17. Map of geographic distribution of species of Dendrophryniscus brevipollicatus group reviewed in this study. RJ, State of Rio de Janeiro; SP, State of São Paulo." httpUri="https://zenodo.org/record/3353905/files/figure.png" pageId="11" pageNumber="38">Fig. 17</figureCitation>
) in the State of 
<collectingRegion id="4992F848FFA6EA19FF34FD9EFEE9FD37" box="[151,268,617,644]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
, 
<collectingCountry id="F341763AFFA6EA19FEBBFD9DFE84FD37" box="[280,353,617,643]" name="Brazil" pageId="11" pageNumber="38">Brazil</collectingCountry>
: Cubatão (
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23 
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51’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FDE0FD9DFD42FD30" box="[579,679,617,644]" degrees="46" direction="west" minutes="27" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.45">
46 
<superScript id="7C239BE2FFA6EA19FDFCFD9DFD82FDC3" attach="none" box="[607,615,617,631]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
27’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FCA4FD9EFCA0FD30" box="[775,837,618,644]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Juquitiba (
<geoCoordinate id="EE62506DFFA6EA19FC5BFD9EFBBDFD30" box="[1016,1112,617,644]" degrees="23" direction="south" minutes="55" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.916666">
23 
<superScript id="7C239BE2FFA6EA19FBB6FD9DFBF8FDC3" attach="left" box="[1045,1053,617,631]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
55’S
</geoCoordinate>
, 47 
<superScript id="7C239BE2FFA6EA19FB21FD9DFB7CFD30" attach="right" box="[1154,1177,617,644]" fontSize="6" pageId="11" pageNumber="38">oo</superScript>
3’W, Datum 
<collectionCode id="ED47AE6FFFA6EA19FA89FD9EFA8DFD30" box="[1322,1384,618,644]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Itanhaém, Parque Estadual da Serra do Mar Núcleo Curucutu (
<geoCoordinate id="EE62506DFFA6EA19FCF9FD79FC53FD13" box="[858,950,652,679]" degrees="23" direction="south" minutes="59" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.983334">
23 
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59’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FC60FD79FBC3FD13" box="[963,1062,652,679]" degrees="46" direction="west" minutes="44" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.733334">
46 
<superScript id="7C239BE2FFA6EA19FC7CFD78FC02FD2E" attach="none" box="[991,999,652,666]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
44’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FB25FD79FB21FD13" box="[1158,1220,653,679]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Natividade da Serra (
<geoCoordinate id="EE62506DFFA6EA19FF41FD45FEDBFD7F" box="[226,318,688,715]" degrees="23" direction="south" minutes="22" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.366667">
23 
<superScript id="7C239BE2FFA6EA19FF5CFD44FEE2FD0A" attach="none" box="[255,263,688,702]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
22’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FEE9FD44FE4BFD7F" box="[330,430,688,715]" degrees="45" direction="west" minutes="26" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-45.433334">
45 
<superScript id="7C239BE2FFA6EA19FEC5FD44FE8BFD0A" attach="none" box="[358,366,688,702]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
26’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FDAEFD45FDAEFD7F" box="[525,587,689,715]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Salesópolis (
<geoCoordinate id="EE62506DFFA6EA19FCB6FD45FC94FD7F" box="[789,881,688,715]" degrees="23" direction="south" minutes="38" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.633333">
23 
<superScript id="7C239BE2FFA6EA19FC91FD44FCDFFD0A" attach="none" box="[818,826,688,702]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
38’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FCDEFD44FC05FD7F" box="[893,992,688,715]" degrees="45" direction="west" minutes="52" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-45.866665">
45 
<superScript id="7C239BE2FFA6EA19FC3AFD44FC44FD0A" attach="none" box="[921,929,688,702]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
52’W
</geoCoordinate>
, Datum WGS 
<quantity id="4CAE9B4FFFA6EA19FB26FD45FB13FD7F" box="[1157,1270,689,715]" metricMagnitude="4" metricUnit="m" metricValue="8.49" pageId="11" pageNumber="38" unit="m" value="84900.0">84, 900 m</quantity>
altitude), Santo André, Paranapiacaba (
<geoCoordinate id="EE62506DFFA6EA19FE1EFD21FDFCFD5B" box="[445,537,724,751]" degrees="23" direction="south" minutes="46" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.766666">
23 
<superScript id="7C239BE2FFA6EA19FE79FD20FE07FD56" attach="none" box="[474,482,724,738]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
46’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FD84FD21FD6EFD5B" box="[551,651,724,751]" degrees="46" direction="west" minutes="17" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.283333">
46 
<superScript id="7C239BE2FFA6EA19FDE0FD20FDAEFD56" attach="none" box="[579,587,724,738]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
17’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FD4EFD21FCCEFD5B" box="[749,811,725,751]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Santos (
<geoCoordinate id="EE62506DFFA6EA19FC65FD21FBC7FD5B" box="[966,1058,724,751]" degrees="23" direction="south" minutes="56" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.933332">
23 
<superScript id="7C239BE2FFA6EA19FC40FD20FC0EFD56" attach="none" box="[995,1003,724,738]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
56’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FB93FD21FB71FD5B" box="[1072,1172,724,751]" degrees="46" direction="west" minutes="19" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.316666">
46 
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19’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FB55FD21FAD1FD5B" box="[1270,1332,725,751]" pageId="11" pageNumber="38">WGS</collectionCode>
84), São Bernardo do Campo (
<geoCoordinate id="EE62506DFFA6EA19FE28FD0CFE02FCA7" box="[395,487,760,787]" degrees="23" direction="south" minutes="45" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.75">23º45’S</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FE50FD0CFDBDFCA6" box="[499,600,760,786]" degrees="46" direction="west" minutes="33" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.55">46º33’W</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FD15FD0DFD11FCA7" box="[694,756,761,787]" pageId="11" pageNumber="38">WGS</collectionCode>
84), 
<collectingRegion id="4992F848FFA6EA19FC8CFD0DFC46FCA6" box="[815,931,760,787]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
, Represa Billings (
<geoCoordinate id="EE62506DFFA6EA19FBDBFD0DFB31FCA7" box="[1144,1236,759,787]" degrees="23" direction="south" minutes="48" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.8">
23 
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48’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FB43FD0CFAA1FCA7" box="[1248,1348,759,787]" degrees="46" direction="west" minutes="37" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.616665">
46 
<superScript id="7C239BE2FFA6EA19FB5FFD03FAE1FCB1" attach="none" box="[1276,1284,759,773]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
37’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FF34FCE8FF30FC82" box="[151,213,796,822]" pageId="11" pageNumber="38">WGS</collectionCode>
84), Cotia, Reserva do Morro Grande (
<geoCoordinate id="EE62506DFFA6EA19FD2CFCE8FD0EFC82" box="[655,747,795,822]" degrees="23" direction="south" minutes="43" orientation="latitude" pageId="11" pageNumber="38" precision="925" value="-23.716667">
23 
<superScript id="7C239BE2FFA6EA19FD0FFCEFFD51FC9D" attach="none" box="[684,692,795,809]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
43’S
</geoCoordinate>
, 
<geoCoordinate id="EE62506DFFA6EA19FD54FCE8FCBEFC82" box="[759,859,795,822]" degrees="46" direction="west" minutes="57" orientation="longitude" pageId="11" pageNumber="38" precision="925" value="-46.95">
46 
<superScript id="7C239BE2FFA6EA19FCB0FCEFFCFEFC9D" attach="none" box="[787,795,795,809]" fontSize="6" pageId="11" pageNumber="38">o</superScript>
57’W
</geoCoordinate>
, Datum 
<collectionCode id="ED47AE6FFFA6EA19FC1AFCE8FC12FC82" box="[953,1015,796,822]" pageId="11" pageNumber="38">WGS</collectionCode>
84).
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FCB4FBACFCCA" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FCB4FE9CFCEE" box="[199,377,832,858]" italics="true" pageId="11" pageNumber="38">Natural history.</emphasis>
This species is associated to forested areas, from 
<quantity id="4CAE9B4FFFA6EA19FC05FCB4FC08FCEE" box="[934,1005,832,858]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="38" unit="m" value="100.0">100 m</quantity>
to 
<quantity id="4CAE9B4FFFA6EA19FBB3FCB4FB81FCEE" box="[1040,1124,832,858]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="38" unit="m" value="1100.0">1100 m</quantity>
above sea level, and breeds in bromeliads, where its larvae develop (
<bibRefCitation id="EFC74B5BFFA6EA19FDFDFC90FCA3FCCA" author="Malagoli, L. R. &amp; Trevine, V. &amp; Condez, T. H. &amp; Centeno, F. C. &amp; Berneck, B. M. &amp; Haddad, C. F. B." box="[606,838,868,894]" pageId="11" pageNumber="38" pagination="31 - 39" refId="ref21946" refString="Malagoli, L. R., Trevine, V., Condez, T. H., Centeno, F. C., Berneck, B. M. &amp; Haddad, C. F. B. (2017) Notes on the breeding behaviour of the Neotropical toadlet Dendrophryniscus brevipollicatus (Anura: Bufonidae), a bromeliad phytotelmata specialist. Herpetology Notes, 10, 31 - 39." type="journal article" year="2017">
Malagoli 
<emphasis id="B922EAB8FFA6EA19FD6FFC90FCE1FCCA" box="[716,772,868,894]" italics="true" pageId="11" pageNumber="38">et al.</emphasis>
2017
</bibRefCitation>
, as 
<taxonomicName id="4C564D29FFA6EA19FCCCFC90FBDDFCCA" authorityName="Jimenez de la Espada" authorityYear="1870" box="[879,1080,868,894]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA6EA19FCCCFC90FBDDFCCA" box="[879,1080,868,894]" italics="true" pageId="11" pageNumber="38">D. brevipollicatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FC73FD19FB50" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
<emphasis id="B922EAB8FFA6EA19FF64FC73FED4FC15" box="[199,305,903,929]" italics="true" pageId="11" pageNumber="38">Remarks.</emphasis>
All individuals of the 
<typeStatus id="54ED8808FFA6EA19FD9FFC7CFD89FC16" box="[572,620,904,930]" pageId="11" pageNumber="38">type</typeStatus>
series have tip of snout kneaded. All skin is unpigmented and surface granulation and crenulations are few evident because time of preservation. For this reason, we presented the original description and made a new description based on a topotype from Paranapiacaba, State of 
<collectingRegion id="4992F848FFA6EA19FB75FC3BFAA9FC5D" box="[1238,1356,975,1001]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
. In the original description the author refers to five specimens composing the 
<typeStatus id="54ED8808FFA6EA19FC16FC07FC00FBB9" box="[949,997,1011,1037]" pageId="11" pageNumber="38">type</typeStatus>
series: two from Santos, Cubatão, 
<collectingRegion id="4992F848FFA6EA19FAD1FC07FF32FB84" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
(
<collectionCode id="ED47AE6FFFA6EA19FF46FBE2FEA6FB84" box="[229,323,1046,1072]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
477–485) collected by Bicego on 
<date id="FFE8106AFFA6EA19FD60FBE2FC9DFB84" box="[707,888,1046,1072]" pageId="11" pageNumber="38" value="1897-12">December 1897</date>
; one from Alto da Serra (
<collectionCode id="ED47AE6FFFA6EA19FB32FBE2FB0AFB84" box="[1169,1263,1046,1072]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
847), collected by Bicego in 1899; one from Rio Grande, State of 
<collectingRegion id="4992F848FFA6EA19FD40FBCEFCB3FBE0" box="[739,854,1082,1108]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
(
<collectionCode id="ED47AE6FFFA6EA19FCC5FBCEFC21FBE0" box="[870,964,1082,1108]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
835), collected by Wacket on 
<date id="FFE8106AFFA6EA19FA86FBCEFA72FBE0" box="[1317,1431,1082,1108]" pageId="11" pageNumber="38" value="1902-07">July 1902</date>
; and one from Campo Grande, State of 
<collectingRegion id="4992F848FFA6EA19FDE8FBAAFD58FBCC" box="[587,701,1118,1144]" country="Brazil" name="Sao Paulo" pageId="11" pageNumber="38">São Paulo</collectingRegion>
(
<collectionCode id="ED47AE6FFFA6EA19FD68FBAAFCCCFBCC" box="[715,809,1118,1144]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
639), collected by Wacket on 
<date id="FFE8106AFFA6EA19FBDCFBAAFB10FBCC" box="[1151,1269,1118,1144]" pageId="11" pageNumber="38" value="1902-05">May 1902</date>
. The specimen 
<collectionCode id="ED47AE6FFFA6EA19FF34FB76FF16FB28" box="[151,243,1154,1180]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
835 from Rio Grande was lost and the tag number of the specimen from Campo Grande is 
<collectionCode id="ED47AE6FFFA6EA19FAA4FB76FA86FB28" box="[1287,1379,1154,1180]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
634. We concluded that body measures presented by Miranda-Ribeiro in the original description were based on specimen 
<collectionCode id="ED47AE6FFFA6EA19FF34FB3DFF16FB57" box="[151,243,1225,1251]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="38">MZUSP</collectionCode>
847 from Alto da Serra (now, Paranapiacaba).
</paragraph>
<paragraph id="8BE936AAFFA6EA19FF64FB19FE9EFA9F" blockId="11.[151,1437,153,1323]" pageId="11" pageNumber="38">
The specimens referred by 
<bibRefCitation id="EFC74B5BFFA6EA19FE5BFB19FD2BFAB3" author="Heyer, W. R. &amp; Rand, A. S. &amp; Cruz, C. A. G. &amp; Peixoto, O. L. &amp; Nelson, C. E." box="[504,718,1261,1287]" pageId="11" pageNumber="38" pagination="231 - 410" refId="ref21468" refString="Heyer, W. R., Rand, A. S., Cruz, C. A. G., Peixoto, O. L. &amp; Nelson, C. E. (1990) Frogs of Boraceia. Arquivos de Zoologia, 31, 231 - 410." type="journal article" year="1990">
Heyer 
<emphasis id="B922EAB8FFA6EA19FDE7FB1AFD99FAB3" box="[580,636,1261,1287]" italics="true" pageId="11" pageNumber="38">et al.</emphasis>
(1990)
</bibRefCitation>
and 
<bibRefCitation id="EFC74B5BFFA6EA19FCA6FB19FC1EFAB3" author="Malagoli, L. R. &amp; Trevine, V. &amp; Condez, T. H. &amp; Centeno, F. C. &amp; Berneck, B. M. &amp; Haddad, C. F. B." box="[773,1019,1261,1287]" pageId="11" pageNumber="38" pagination="31 - 39" refId="ref21946" refString="Malagoli, L. R., Trevine, V., Condez, T. H., Centeno, F. C., Berneck, B. M. &amp; Haddad, C. F. B. (2017) Notes on the breeding behaviour of the Neotropical toadlet Dendrophryniscus brevipollicatus (Anura: Bufonidae), a bromeliad phytotelmata specialist. Herpetology Notes, 10, 31 - 39." type="journal article" year="2017">
Malagoli 
<emphasis id="B922EAB8FFA6EA19FCD2FB1AFC4CFAB3" box="[881,937,1261,1287]" italics="true" pageId="11" pageNumber="38">et al.</emphasis>
(2017)
</bibRefCitation>
as 
<taxonomicName id="4C564D29FFA6EA19FB83FB19FA79FAB3" authorityName="Jimenez de la Espada" authorityYear="1870" box="[1056,1436,1261,1287]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="brevipollicatus">
<emphasis id="B922EAB8FFA6EA19FB83FB19FA79FAB3" box="[1056,1436,1261,1287]" italics="true" pageId="11" pageNumber="38">Dendrophryniscus brevipollicatus</emphasis>
</taxonomicName>
are now 
<taxonomicName id="4C564D29FFA6EA19FF5BFAE6FE91FA9F" authorityName="Cruz &amp; Caramaschi &amp; Fusinatto &amp; Brasileiro" authorityYear="2019" baseAuthorityName="Miranda-Ribeiro" baseAuthorityYear="1920" box="[248,372,1298,1323]" class="Amphibia" family="Bufonidae" genus="Dendrophryniscus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="38" phylum="Chordata" rank="species" species="imitator">
<emphasis id="B922EAB8FFA6EA19FF5BFAE6FE91FA9F" box="[248,372,1298,1323]" italics="true" pageId="11" pageNumber="38">D. imitator</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>