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<document id="D932397D54B1AA83F62B461F4AF95C19" ID-DOI="10.11646/zootaxa.3852.3.1" ID-GBIF-Dataset="957e5d0f-c452-4d45-85cf-24f138ab2359" ID-ISSN="1175-5326" ID-Zenodo-Dep="229938" ID-ZooBank="13B15A57-576A-4617-9D8A-7F27D970315A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460883855845" checkinUser="plazi" docAuthor="Couri, Márcia &amp; Pont, Adrian C." docDate="2014" docId="03FF87844606FFD7FF5EFDEA6BC4B52D" docLanguage="en" docName="zt03852p320.pdf" docOrigin="Zootaxa 3852 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helina harrisorum Couri &amp; Pont, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="309" masterDocId="FFC6FFFC4600FFDFFFC9FFCB6E55B616" masterDocTitle="New species of Afrotropical Muscidae (Diptera: Muscoidea)" masterLastPageNumber="320" masterPageNumber="301" pageNumber="307" updateTime="1698358705595" updateUser="plazi">
<mods:mods id="0099A6CD0DEDC0BCF1EF1100B78DBE48" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DC684698262ADBC40B43FC08E7FC800">
<mods:title id="530072A002E7AF35138F75191149A6AA">New species of Afrotropical Muscidae (Diptera: Muscoidea)</mods:title>
</mods:titleInfo>
<mods:name id="D922A3C26E6217FAF72F0D74CFB5B20E" type="personal">
<mods:role id="6DCAE256D675B4B80A1B8B069AE8F3CA">
<mods:roleTerm id="5C89BE535A6C451EFB9E1D14DFE3882E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24C5598B51B349FD16C6CB878D755F38">Couri, Márcia</mods:namePart>
</mods:name>
<mods:name id="FB5D5FC3FE653731EE0FE8887CB7567E" type="personal">
<mods:role id="8C2B131CC7B46E3B758B49DD4E881F33">
<mods:roleTerm id="8CFCF9C66B4344E47AAC741DD3ABE5A4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8A6A638ABA8C15F72035220607C39CE">Pont, Adrian C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A6364748C1B4A3AE84964B1AE0564AFA">text</mods:typeOfResource>
<mods:relatedItem id="757829F03D3ACA589F958A5A18232270" type="host">
<mods:titleInfo id="F7511AA0116DFE950AA1139E4AB82BAD">
<mods:title id="EDA0A0504017BA97B1EECF665CE702C3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7BA75FCEF6348A95E2D812C4AA7633B3">
<mods:date id="052C91F2F3B93FA7B2D84690F8604FFC">2014</mods:date>
<mods:detail id="07D95BADC79D49B3A9DA3A01540C81AA" type="volume">
<mods:number id="BF8B0322C05833ED92615D003A638D91">3852</mods:number>
</mods:detail>
<mods:detail id="038D713D707B6DAA830B96D314F2089D" type="issue">
<mods:number id="CFFC2B7C4E153F8C4E53ADDFF67E5E49">3</mods:number>
</mods:detail>
<mods:extent id="5BC9F5DC43020B44869E3DA983B528AC" unit="page">
<mods:start id="71139913FB59BA702E06084C1FEB44B8">301</mods:start>
<mods:end id="FADEDD975E9F8F54FA640916C905FF92">320</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="604CA819E7676075CF810E0688FDB41B">journal article</mods:classification>
<mods:identifier id="52E9486E7C89A651B282C64606B6321B" type="DOI">10.11646/zootaxa.3852.3.1</mods:identifier>
<mods:identifier id="2AB4780F0623A8BC02EF23EE8AF3529C" type="GBIF-Dataset">957e5d0f-c452-4d45-85cf-24f138ab2359</mods:identifier>
<mods:identifier id="C7E639AA37FC879D36EBA7B3C1B52F70" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="37ACC775F20BD11BB170AD369B4E8DF0" type="Zenodo-Dep">229938</mods:identifier>
<mods:identifier id="0657022C272FD5678ABB1726F3F22F56" type="ZooBank">13B15A57-576A-4617-9D8A-7F27D970315A</mods:identifier>
</mods:mods>
<treatment id="03FF87844606FFD7FF5EFDEA6BC4B52D" ID-DOI="http://doi.org/10.5281/zenodo.6137880" ID-GBIF-Taxon="119585837" ID-Zenodo-Dep="6137880" LSID="urn:lsid:plazi:treatment:03FF87844606FFD7FF5EFDEA6BC4B52D" httpUri="http://treatment.plazi.org/id/03FF87844606FFD7FF5EFDEA6BC4B52D" lastPageId="8" lastPageNumber="309" pageId="6" pageNumber="307">
<subSubSection id="C34C65194606FFD9FF5EFDEA6F62B44A" pageId="6" pageNumber="307" type="nomenclature">
<paragraph id="8BE936924606FFD9FF5EFDEA6FB5B42D" blockId="6.[151,480,545,605]" box="[151,480,545,571]" pageId="6" pageNumber="307">
<heading id="D0A181FE4606FFD9FF5EFDEA6FB5B42D" bold="true" box="[151,480,545,571]" fontSize="11" level="1" pageId="6" pageNumber="307" reason="1">
<emphasis id="B922EA804606FFD9FF5EFDEA6FB5B42D" bold="true" box="[151,480,545,571]" pageId="6" pageNumber="307">
<taxonomicName id="4C564D114606FFD9FF5EFDEA6F29B42D" box="[151,380,545,571]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="harrisorum" status="sp. nov.">
<emphasis id="B922EA804606FFD9FF5EFDEA6F29B42D" bold="true" box="[151,380,545,571]" italics="true" pageId="6" pageNumber="307">Helina harrisorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4606FFD9FE4AFDEA6FB5B42D" box="[387,480,545,571]" pageId="6" pageNumber="307" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936924606FFD9FF5EFD8F6F62B44A" blockId="6.[151,480,545,605]" box="[151,311,580,605]" pageId="6" pageNumber="307">
<figureCitation id="136D2A174606FFD9FF5EFD8F6E88B44A" box="[151,221,580,605]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,264,1956,1979]" captionTargetBox="[196,1429,467,1915]" captionTargetId="figure@2.[173,1458,445,1939]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1. Coenosia duomaculata sp. nov., notum, dorsal view; 2. Coenosia nigromaculata sp. nov., notum, dorsal view; 3. Helina harrisorum sp. nov., notum, dorsal view; 4. Helina ferfriniorum sp. nov., notum, dorsal view." httpUri="https://zenodo.org/record/229939/files/figure.png" pageId="6" pageNumber="307">Figs 3</figureCitation>
,
<figureCitation id="136D2A174606FFD9FF23FD8F6F66B44A" box="[234,307,580,604]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,264,1648,1671]" captionTargetBox="[151,1414,197,1571]" captionTargetId="figure@7.[151,1436,193,1628]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Helina harrisorum sp. nov.: 13. cercal plate and surstyli, dorsal view; 14. epandrium, cercal plate and surstylus, lateral view; 15. aedeagus, lateral view; 16. ovipositor, dorsal view; 17. ovipositor, ventral view and spermathecae. Helina ferfriniorum sp. nov.: 18. ovipositor, lateral view; 19. egg." httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="6" pageNumber="307">13–17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194606FFD9FF5EFD476AF2B31E" pageId="6" pageNumber="307" type="materials_examined">
<paragraph id="8BE936924606FFD9FF5EFD476C65B4FA" blockId="6.[151,1437,652,1939]" pageId="6" pageNumber="307">
<emphasis id="B922EA804606FFD9FF5EFD476F1CB4B3" bold="true" box="[151,329,652,677]" pageId="6" pageNumber="307">
<typeStatus id="54ED88304606FFD9FF5EFD476E87B4B3" box="[151,210,652,677]" pageId="6" pageNumber="307">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54ED88304606FFD9FEAAFD476F9EB4B3" box="[355,459,652,677]" pageId="6" pageNumber="307" type="holotype">Holotype</typeStatus>
♂,
<collectingCountry id="F34176024606FFD9FDC4FD476DECB4B2" box="[525,953,652,676]" name="South Africa" pageId="6" pageNumber="307">REPUBLIC OF SOUTH AFRICA</collectingCountry>
: Free State, Harrismith, Mooihoekkop, 28°10′50.0′S, 29°10′51.1′E,
<emphasis id="B922EA804606FFD9FE11FD7A6FA7B4DE" box="[472,498,689,712]" italics="true" pageId="6" pageNumber="307">ca</emphasis>
.
<quantity id="4CAE9B774606FFD9FDCAFD7B6C09B4D1" box="[515,604,688,712]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="307" unit="m" value="1800.0">1800 m</quantity>
, Malaise traps,
<emphasis id="B922EA804606FFD9FCDBFD646DC3B4DE" box="[786,918,687,712]" italics="true" pageId="6" pageNumber="307">Leucosedea</emphasis>
-dominated scrub,
<date id="FFE810524606FFD9FBA2FD646B58B4DE" box="[1131,1293,687,712]" pageId="6" pageNumber="307" value="2009-09-14" valueMax="2009-09-16" valueMin="2009-09-14">14–16.ix.2009</date>
(A.H. Kirk- Spriggs),
<collectionCode id="ED47AE574606FFD9FECDFD1F6F03B4FB" box="[260,342,724,749]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)08761 (
<collectionCode id="ED47AE574606FFD9FE06FD1F6C71B4FA" box="[463,548,724,748]" pageId="6" pageNumber="307">BMSA</collectionCode>
).
</paragraph>
<paragraph id="8BE936924606FFD9FF0EFD336AF2B31E" blockId="6.[151,1437,652,1939]" pageId="6" pageNumber="307">
<typeStatus id="54ED88304606FFD9FF0EFD336F62B506" box="[199,311,760,784]" pageId="6" pageNumber="307" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D50FD1B4606FFD9FE85FD3C6F28B506" box="[332,381,759,784]" pageId="6" pageNumber="307" type="generic">56♂</specimenCount>
41♀.
<collectingCountry id="F34176024606FFD9FE1EFD336D2BB506" box="[471,894,760,784]" name="South Africa" pageId="6" pageNumber="307">REPUBLIC OF SOUTH AFRICA</collectingCountry>
:
<specimenCount id="9D50FD1B4606FFD9FC5AFD336D91B506" box="[915,964,760,784]" pageId="6" pageNumber="307" type="generic">48♂</specimenCount>
33♀, same data as
<typeStatus id="54ED88304606FFD9FB18FD3C6B63B506" box="[1233,1334,759,784]" pageId="6" pageNumber="307" type="holotype">holotype</typeStatus>
, except:
<collectionCode id="ED47AE574606FFD9FF5EFCD76EBCB523" box="[151,233,796,821]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)08760, 0 8763, 0 8764, 0 8782, 0 8788, 0 8797, 0 8799, 0 8800, 0 8806, 0 8808, 0 8818, 0 8819, 08821–08824, 0 8828, 0 8832, 0 8834, 0 8839, 0 8863, 0 8865, 0 8866, 0 8868, 0 8871, 0 8883, 0 8887, 0 8922, 0 8981, 0 9101, 0 9105, 0 9106, 0 9109, 0 9110, 0 9113, 0 9135, 0 9138, 0 9139, 0 9141, 0 9145, 0 9151, 0 9156, 09158–09165, 0 9167, 09169–09176, 0 9178, 0 9179, 09181–09183, 0 9189, 0 9190, 0 9195, 0 9201, 0 9202, 0 9206, 0 9208, 0 9215, 09225–09229, 10026, 10044, 10057, 10066” (all
<collectionCode id="ED47AE574606FFD9FD01FC676D49B5D2" box="[712,796,940,964]" pageId="6" pageNumber="307">BMSA</collectionCode>
);
<specimenCount id="9D50FD1B4606FFD9FCF8FC676D00B5D2" box="[817,853,940,964]" pageId="6" pageNumber="307" type="generic">1♂</specimenCount>
, 2♀, same, except:
<collectionCode id="ED47AE574606FFD9FBF1FC676ADFB5D3" box="[1080,1162,940,965]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)08762, 0 9111, 0 9180 (all
<collectionCode id="ED47AE574606FFD9FF05FC1B6F49B5FE" box="[204,284,976,1000]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="307">MNRJ</collectionCode>
);
<specimenCount id="9D50FD1B4606FFD9FEF0FC1B6F08B5FE" box="[313,349,976,1000]" pageId="6" pageNumber="307" type="generic">1♂</specimenCount>
, same, except:
<collectionCode id="ED47AE574606FFD9FDEFFC046C2DB5FE" box="[550,632,975,1000]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)10265 (
<collectionCode id="ED47AE574606FFD9FD33FC1B6D01B5FE" box="[762,852,976,1000]" pageId="6" pageNumber="307">BMNH</collectionCode>
);
<collectingCountry id="F34176024606FFD9FCB8FC1B6DF9B5FE" box="[881,940,976,1000]" name="South Africa" pageId="6" pageNumber="307">RSA</collectingCountry>
: Free State, Harrismith, Nesshurst farm, 28°26′53.3′S, 29°09′14.2′E, Malaise traps,
<emphasis id="B922EA804606FFD9FD4AFC3F6D52B21B" box="[643,775,1012,1037]" italics="true" pageId="6" pageNumber="307">Leucosedea</emphasis>
-scrub &amp; stream bed,
<date id="FFE810524606FFD9FBCDFC3F6AF3B21B" box="[1028,1190,1012,1037]" pageId="6" pageNumber="307" value="2009-09-17" valueMax="2009-09-18" valueMin="2009-09-17">17–18.ix.2009</date>
(A.H. Kirk-Spriggs),
<specimenCount id="9D50FD1B4606FFD9FF5EFBD36EE9B226" box="[151,188,1048,1072]" pageId="6" pageNumber="307" type="generic">2♂</specimenCount>
, 2♀,
<collectionCode id="ED47AE574606FFD9FECAFBD36F03B226" box="[259,342,1048,1072]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)10739, 10760, 10795, 10849 (all
<collectionCode id="ED47AE574606FFD9FCCEFBD36D0EB226" box="[775,859,1048,1072]" pageId="6" pageNumber="307">BMSA</collectionCode>
);
<collectingCountry id="F34176024606FFD9FCBAFBD36DFBB226" box="[883,942,1048,1072]" name="South Africa" pageId="6" pageNumber="307">RSA</collectingCountry>
: Free State, Harrismith,
<collectingCountry id="F34176024606FFD9FB1BFBDC6B61B226" box="[1234,1332,1047,1072]" name="United Kingdom" pageId="6" pageNumber="307">Scotland</collectingCountry>
farm at: 27°58′59.5′S, 29°37′09.8′E, Malaise traps, dense
<emphasis id="B922EA804606FFD9FD3BFBF76D23B243" box="[754,886,1084,1109]" italics="true" pageId="6" pageNumber="307">Leucosedea</emphasis>
-dominated scrub,
<date id="FFE810524606FFD9FB95FBF76AA8B243" box="[1116,1277,1084,1109]" pageId="6" pageNumber="307" value="2009-11-10" valueMax="2009-11-12" valueMin="2009-11-10">10–12.xi.2009</date>
(A.H. Kirk- Spriggs),
<specimenCount id="9D50FD1B4606FFD9FECDFBAB6F7FB26E" box="[260,298,1120,1144]" pageId="6" pageNumber="307" type="generic">4♂</specimenCount>
, 5♀,
<collectionCode id="ED47AE574606FFD9FEAEFB946FEFB26E" box="[359,442,1119,1144]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)11453, 11504, 11542, 11561, 11607, 11616, 11660, 11687, 11740 (all
<collectionCode id="ED47AE574606FFD9FB3AFBAB6B12B26E" box="[1267,1351,1120,1144]" pageId="6" pageNumber="307">BMSA</collectionCode>
); same, except:
<collectionCode id="ED47AE574606FFD9FF27FB4F6F14B28B" box="[238,321,1156,1181]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)11648,
<specimenCount id="9D50FD1B4606FFD9FE7EFB4F6F8FB28A" box="[439,474,1156,1180]" pageId="6" pageNumber="307" type="generic">1♂</specimenCount>
(
<collectionCode id="ED47AE574606FFD9FE20FB4F6C6FB28A" box="[489,570,1156,1180]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="307">MNRJ</collectionCode>
);
<collectingCountry id="F34176024606FFD9FD84FB4F6CDDB28A" box="[589,648,1156,1180]" name="South Africa" pageId="6" pageNumber="307">RSA</collectingCountry>
: Free State, Harrismith, Geluksdal farm at: 27°54′7.05′S,29°23′31.9′E, Malaise traps, sparse
<emphasis id="B922EA804606FFD9FE56FB6C6C76B2D6" box="[415,547,1191,1216]" italics="true" pageId="6" pageNumber="307">Leucosedea</emphasis>
-dominated scrub,
<date id="FFE810524606FFD9FCCCFB6C6DCDB2D6" box="[773,920,1191,1216]" pageId="6" pageNumber="307" value="2009-11-09" valueMax="2009-11-10" valueMin="2009-11-09">9–10.xi.2009</date>
(A.H. Kirk-Spriggs),
<specimenCount id="9D50FD1B4606FFD9FB6EFB636A9EB2D6" box="[1191,1227,1192,1216]" pageId="6" pageNumber="307" type="generic">1♂</specimenCount>
,
<collectionCode id="ED47AE574606FFD9FB29FB636B67B2D6" box="[1248,1330,1192,1216]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)13230 (
<collectionCode id="ED47AE574606FFD9FF56FB076EA0B2F2" box="[159,245,1228,1252]" pageId="6" pageNumber="307">BMSA</collectionCode>
);
<collectingCountry id="F34176024606FFD9FECEFB076F17B2F2" box="[263,322,1228,1252]" name="South Africa" pageId="6" pageNumber="307">RSA</collectingCountry>
: Eastern Cape, Tsitsikamma N.P., below Sleepkloof hut at:
<geoCoordinate id="EE6250554606FFD9FC10FB076A33B2F3" box="[985,1126,1228,1253]" direction="south" orientation="latitude" pageId="6" pageNumber="307" precision="1" value="-33.949566">33°56.974′S</geoCoordinate>
,
<geoCoordinate id="EE6250554606FFD9FBB8FB076AAAB2F3" box="[1137,1279,1228,1253]" direction="east" orientation="longitude" pageId="6" pageNumber="307" precision="1" value="23.915434">23°54.926′E</geoCoordinate>
, Malaise trap, indigenous forest,
<date id="FFE810524606FFD9FEAFFB246FAEB31E" box="[358,507,1263,1288]" pageId="6" pageNumber="307" value="2009-01-20" valueMax="2009-01-22" valueMin="2009-01-20">20–22.i.2009</date>
(A. Kirk-Spriggs &amp; S. Otto), 1♀,
<collectionCode id="ED47AE574606FFD9FCB3FB246D9EB31E" box="[890,971,1263,1288]" pageId="6" pageNumber="307">BMSA</collectionCode>
(D)03159 (
<collectionCode id="ED47AE574606FFD9FB8FFB3B6ACEB31E" box="[1094,1179,1264,1288]" pageId="6" pageNumber="307">BMSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C65194606FFD9FF0EFADF6B44B38E" pageId="6" pageNumber="307" type="diagnosis">
<paragraph id="8BE936924606FFD9FF0EFADF6B44B38E" blockId="6.[151,1437,652,1939]" pageId="6" pageNumber="307">
<emphasis id="B922EA804606FFD9FF0EFADF6F14B33B" bold="true" box="[199,321,1300,1325]" pageId="6" pageNumber="307">Diagnosis.</emphasis>
Distinguishable by the colour patterning of the notum, the bare meron and presence of 2 postsutural intra-alars. Morphologically similar to
<taxonomicName id="4C564D114606FFD9FD80FAF26D37B346" authority="Seguy, 1937" authorityName="Seguy" authorityYear="1937" box="[585,866,1336,1360]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="icterica">
<emphasis id="B922EA804606FFD9FD80FAF26C08B346" box="[585,605,1337,1360]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B922EA804606FFD9FDA3FAF26C95B346" box="[618,704,1337,1360]" italics="true" pageId="6" pageNumber="307">icterica</emphasis>
(Séguy, 1937)
</taxonomicName>
and
<taxonomicName id="4C564D114606FFD9FC51FAF26A96B346" authority="Zielke, 1971" authorityName="Zielke" authorityYear="1971" box="[920,1219,1335,1360]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="B922EA804606FFD9FC51FAF26DF9B346" box="[920,940,1337,1360]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B922EA804606FFD9FC70FAFC6A7BB346" box="[953,1070,1335,1360]" italics="true" pageId="6" pageNumber="307">scutellaris</emphasis>
<bibRefCitation id="EFC74B634606FFD9FBFDFAFC6A96B346" author="Zielke" box="[1076,1219,1335,1360]" pageId="6" pageNumber="307" refString="Zielke, E. (1971) New species of Muscidae from the Ethiopian region (Diptera). Journal of the Entomological Society of South Africa, 34, 289 - 304." type="journal article" year="1971">Zielke, 1971</bibRefCitation>
</taxonomicName>
, but
<taxonomicName id="4C564D114606FFD9FB30FAF26B24B346" box="[1273,1393,1337,1360]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="icterica">
<emphasis id="B922EA804606FFD9FB30FAF26B58B346" box="[1273,1293,1337,1360]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B922EA804606FFD9FAD2FAF26B24B346" box="[1307,1393,1337,1360]" italics="true" pageId="6" pageNumber="307">icterica</emphasis>
</taxonomicName>
has virtually clear crossveins and the dark abdominal maculae obsolescent, whilst
<taxonomicName id="4C564D114606FFD9FBDBFA966AFEB363" box="[1042,1195,1372,1397]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="B922EA804606FFD9FBDBFA966A73B362" box="[1042,1062,1373,1396]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B922EA804606FFD9FBFFFA976AFEB363" box="[1078,1195,1372,1397]" italics="true" pageId="6" pageNumber="307">scutellaris</emphasis>
</taxonomicName>
differs from the new species, according to Zielke’s (1971) description, by the wings having a strong yellowish brown tinge.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194606FFD7FF0EFA6F6CE3B4E5" lastPageId="8" lastPageNumber="309" pageId="6" pageNumber="307" type="description">
<paragraph id="8BE936924606FFD9FF0EFA6F6D31B0C9" blockId="6.[151,1437,652,1939]" pageId="6" pageNumber="307">
<emphasis id="B922EA804606FFD9FF0EFA6F6F0CB3AB" bold="true" box="[199,345,1444,1469]" pageId="6" pageNumber="307">Description.</emphasis>
<emphasis id="B922EA804606FFD9FEAAFA6F6C19B3AB" box="[355,588,1444,1469]" italics="true" pageId="6" pageNumber="307">General colouration</emphasis>
. Ground colour brown and yellow, with grey to whitish grey pruinosity; head with frons black; fronto-orbital plate, parafacial and face silvery pruinose; gena grey; antenna brown, scape with yellow areas, arista brown; palpus yellow; notum with patterning as in
<figureCitation id="136D2A174606FFD9FC2FFA276A7CB012" box="[998,1065,1516,1541]" captionStart="FIGURES 1 – 4. 1" captionStartId="2.[151,264,1956,1979]" captionTargetBox="[196,1429,467,1915]" captionTargetId="figure@2.[173,1458,445,1939]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1. Coenosia duomaculata sp. nov., notum, dorsal view; 2. Coenosia nigromaculata sp. nov., notum, dorsal view; 3. Helina harrisorum sp. nov., notum, dorsal view; 4. Helina ferfriniorum sp. nov., notum, dorsal view." httpUri="https://zenodo.org/record/229939/files/figure.png" pageId="6" pageNumber="307">Fig. 3</figureCitation>
; pleura mostly yellow, with large grey-brown areas on anepisternum, anepimeron and meron; spiracles yellow; wing with black infuscations over
<emphasis id="B922EA804606FFD9FF5EF9FE6E91B05A" box="[151,196,1589,1612]" italics="true" pageId="6" pageNumber="307">r–m</emphasis>
and
<emphasis id="B922EA804606FFD9FF35F9FF6F12B05B" box="[252,327,1588,1613]" italics="true" pageId="6" pageNumber="307">dm–cu</emphasis>
crossveins, the latter more intense at its ends; bifurcation of veins
<emphasis id="B922EA804606FFD9FBF2F9FE6A30B047" box="[1083,1125,1589,1617]" italics="true" pageId="6" pageNumber="307">R2+3</emphasis>
and
<emphasis id="B922EA804606FFD9FB54F9FE6A9DB047" box="[1181,1224,1589,1617]" italics="true" pageId="6" pageNumber="307">R4+5</emphasis>
dark brown; veins otherwise yellow; calypters white; haltere light yellow, knob white; legs yellow, tarsi brown; abdomen brown, with grey pruinosity and some yellow marks, especially in female; syntergite 1 + 2 of male mostly yellow; syntergite 1 + 2 and tergites 3–4 with light brown lateral infuscations; tergite 5 with median fascia, not reaching the posterior margin, bases of setae in female with small brown round marks.
</paragraph>
<paragraph id="8BE936924606FFD9FF0EF9206DDDB114" blockId="6.[151,1437,652,1939]" box="[199,904,1770,1795]" pageId="6" pageNumber="307">
♂.
<emphasis id="B922EA804606FFD9FF23F9276FD9B115" box="[234,396,1772,1795]" italics="true" pageId="6" pageNumber="307">Measurements</emphasis>
. Body length: 6.0 mm; wing length:
<quantity id="4CAE9B774606FFD9FCE2F9216DD1B114" box="[811,900,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="307" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
<paragraph id="8BE936924606FFD9FF0EF8C56DD8B185" blockId="6.[151,1437,652,1939]" pageId="6" pageNumber="307">
<emphasis id="B922EA804606FFD9FF0EF8C56F51B131" box="[199,260,1806,1831]" italics="true" pageId="6" pageNumber="307">Head</emphasis>
. Eye bare; eyes separated at level of anterior ocellus by distance equal to ¼ head-width; frons narrowed medially; 3 pairs of frontal setae close to lunule; medial vertical seta long, lateral vertical short; ocellar setae short; antenna inserted at mid-level of eye, postpedicel
<emphasis id="B922EA804606FFD9FD2DF8936CAAB179" box="[740,767,1880,1903]" italics="true" pageId="6" pageNumber="307">ca</emphasis>
. 3 × length of pedicel; arista long-plumose, the longest combined plumosity equal to length of postpedicel; palpus filiform.
</paragraph>
<caption id="DF29661A4607FFD8FF5EF9BB6D64B0D2" httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="7" pageNumber="308" targetBox="[151,1414,197,1571]" targetPageId="7">
<paragraph id="8BE936924607FFD8FF5EF9BB6D64B0D2" blockId="7.[151,1437,1648,1732]" pageId="7" pageNumber="308">
<emphasis id="B922EA804607FFD8FF5EF9BB6F02B090" bold="true" box="[151,343,1648,1671]" pageId="7" pageNumber="308">FIGURES 13–19.</emphasis>
<taxonomicName id="4C564D114607FFD8FEA8F9BA6C77B091" box="[353,546,1649,1671]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="harrisorum" status="sp. nov.">
<emphasis id="B922EA804607FFD8FEA8F9BA6C77B091" box="[353,546,1649,1671]" italics="true" pageId="7" pageNumber="308">Helina harrisorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4607FFD8FDE5F9BA6CD4B090" box="[556,641,1649,1670]" pageId="7" pageNumber="308" rank="species">
<emphasis id="B922EA804607FFD8FDE5F9BA6CD4B090" bold="true" box="[556,641,1649,1670]" pageId="7" pageNumber="308">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="B922EA804607FFD8FD5BF9BA6CFEB090" bold="true" box="[658,683,1649,1670]" pageId="7" pageNumber="308">13</emphasis>
. cercal plate and surstyli, dorsal view;
<emphasis id="B922EA804607FFD8FB99F9BA6A3CB090" bold="true" box="[1104,1129,1649,1670]" pageId="7" pageNumber="308">14</emphasis>
. epandrium, cercal plate and surstylus, lateral view;
<emphasis id="B922EA804607FFD8FE4CF95B6FCBB0B3" bold="true" box="[389,414,1680,1701]" pageId="7" pageNumber="308">15</emphasis>
. aedeagus, lateral view;
<emphasis id="B922EA804607FFD8FD57F95B6CE2B0B3" bold="true" box="[670,695,1680,1701]" pageId="7" pageNumber="308">16</emphasis>
. ovipositor, dorsal view;
<emphasis id="B922EA804607FFD8FC77F95B6D82B0B3" bold="true" box="[958,983,1680,1701]" pageId="7" pageNumber="308">17</emphasis>
. ovipositor, ventral view and spermathecae.
<taxonomicName id="4C564D114607FFD8FF5EF9656F0AB0D2" box="[151,351,1710,1732]" class="Insecta" family="Lonchaeidae" genus="Helina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="ferfriniorum" status="sp. nov.">
<emphasis id="B922EA804607FFD8FF5EF9656F0AB0D2" box="[151,351,1710,1732]" italics="true" pageId="7" pageNumber="308">Helina ferfriniorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157FB4607FFD8FEACF9646FE3B0D2" box="[357,438,1711,1732]" pageId="7" pageNumber="308" rank="species">
<emphasis id="B922EA804607FFD8FEACF9646FE3B0D2" bold="true" box="[357,438,1711,1732]" pageId="7" pageNumber="308">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="B922EA804607FFD8FE0DF9656F88B0D5" bold="true" box="[452,477,1710,1731]" pageId="7" pageNumber="308">18</emphasis>
. ovipositor, lateral view;
<emphasis id="B922EA804607FFD8FD29F9656CACB0D5" bold="true" box="[736,761,1710,1731]" pageId="7" pageNumber="308">19</emphasis>
. egg.
</paragraph>
</caption>
<paragraph id="8BE936924607FFD8FF0EF93A6A55B1F4" blockId="7.[151,1436,1777,2018]" pageId="7" pageNumber="308">
<emphasis id="B922EA804607FFD8FF0EF93A6F4DB11C" box="[199,280,1777,1802]" italics="true" pageId="7" pageNumber="308">Thorax</emphasis>
. Dorsocentrals 2 + 3; acrostichal setulae in 3–4 irregular rows presuturally, arranged in 5–6 irregular rows postsuturally, prescutellar pair fine; 2 postpronotals; 1 presutural and 2 postsutural intra-alars; 1 presutural and 2 postsutural supra-alars, posterior one short; 2 postalars; prealar present, but short, barely 3 × as long as ground-setulae; notopleuron with 2 setae, the posterior slightly shorter than anterior, otherwise bare; anepisternum with series of 4 long setae, with finer one between third and fourth, and setula in upper anterior corner; katepisternals 1 + 2 (some specimens with short, but distinct, lower anterior seta). Scutellum with 1 long basal and 1 long apical pair of setae, similar in length; margins and ventral surface bare.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFF526F63B7BB" blockId="8.[151,1436,151,827]" pageId="8" pageNumber="309">
<emphasis id="B922EA804608FFD7FF0EFF526EA9B6A6" box="[199,252,153,176]" italics="true" pageId="8" pageNumber="309">Legs</emphasis>
. Fore femur with rows of posteroventral, posterior and posterodorsal setae; fore tibia with posterior median seta and dorsal preapical seta; apical posteroventral long; mid femur with series of anterodorsal setae in basal ½ and 1 preapical, 3 posteroventral setae in basal ⅓, but no anteroventrals; 3 posterior preapicals, 1 short; mid tibia with 2–3 posterodorsal setae, without anterodorsals; anterior, anteroventral, ventral and posteroventral apical setae, the ventral one longer; hind femur with complete row of anterodorsal setae and well-spaced row of 4–6 anteroventrals, anterodorsal, dorsal and posterodorsal surfaces with preapical setae; hind tibia with 1–2 anterodorsal setae on median ⅓, 1 submedian anteroventral, 1 dorsal and 1 anterodorsal preapical, and 1 long ventral apical.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFE726FFCB7E1" blockId="8.[151,1436,151,827]" pageId="8" pageNumber="309">
<emphasis id="B922EA804608FFD7FF0EFE726EAAB7C6" box="[199,255,441,464]" italics="true" pageId="8" pageNumber="309">Wing</emphasis>
. Veins bare; veins
<emphasis id="B922EA804608FFD7FE12FE726C50B7C2" box="[475,517,441,468]" italics="true" pageId="8" pageNumber="309">R4+5</emphasis>
and
<emphasis id="B922EA804608FFD7FDF6FE726C03B7C6" box="[575,598,441,464]" italics="true" pageId="8" pageNumber="309">M</emphasis>
parallel at wing-tip; costal spine developed; lower calypter large,
<emphasis id="B922EA804608FFD7FA85FE726B33B7C6" box="[1356,1382,441,464]" italics="true" pageId="8" pageNumber="309">ca</emphasis>
. 2 × length of upper calypter.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFDC96FD1B429" blockId="8.[151,1436,151,827]" pageId="8" pageNumber="309">
<emphasis id="B922EA804608FFD7FF0EFDC96F64B40D" box="[199,305,514,539]" italics="true" pageId="8" pageNumber="309">Abdomen</emphasis>
. Syntergite 1 + 2 and tergite 3 with lateral setae; tergites 4–5 each with marginal and discal row of setae; sternite 1 bare.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFD816B11B475" blockId="8.[151,1436,151,827]" box="[199,1348,586,611]" pageId="8" pageNumber="309">
<taxonomicName id="4C564D114608FFD7FF0EFD816F14B475" box="[199,321,586,611]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="309" phylum="Tracheophyta" rank="genus">
<emphasis id="B922EA804608FFD7FF0EFD816F14B475" box="[199,321,586,611]" italics="true" pageId="8" pageNumber="309">Terminalia</emphasis>
</taxonomicName>
. Cercal plate virtually as high as wide; surstylus large (
<figureCitation id="136D2A174608FFD7FC67FD816A65B475" box="[942,1072,586,611]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,264,1648,1671]" captionTargetBox="[151,1414,197,1571]" captionTargetId="figure@7.[151,1436,193,1628]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Helina harrisorum sp. nov.: 13. cercal plate and surstyli, dorsal view; 14. epandrium, cercal plate and surstylus, lateral view; 15. aedeagus, lateral view; 16. ovipositor, dorsal view; 17. ovipositor, ventral view and spermathecae. Helina ferfriniorum sp. nov.: 18. ovipositor, lateral view; 19. egg." httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="8" pageNumber="309">Figs 13–14</figureCitation>
); aedeagus as in
<figureCitation id="136D2A174608FFD7FB24FD816B15B475" box="[1261,1344,586,611]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,264,1648,1671]" captionTargetBox="[151,1414,197,1571]" captionTargetId="figure@7.[151,1436,193,1628]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Helina harrisorum sp. nov.: 13. cercal plate and surstyli, dorsal view; 14. epandrium, cercal plate and surstylus, lateral view; 15. aedeagus, lateral view; 16. ovipositor, dorsal view; 17. ovipositor, ventral view and spermathecae. Helina ferfriniorum sp. nov.: 18. ovipositor, lateral view; 19. egg." httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="8" pageNumber="309">Fig. 15</figureCitation>
.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFDBB6DDDB491" blockId="8.[151,1436,151,827]" box="[199,904,622,647]" pageId="8" pageNumber="309">
♀.
<emphasis id="B922EA804608FFD7FF23FDBB6FD9B491" box="[234,396,624,647]" italics="true" pageId="8" pageNumber="309">Measurements</emphasis>
. Body length:
<quantity id="4CAE9B774608FFD7FDFAFDA56CD9B491" box="[563,652,622,647]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="309" unit="mm" value="6.2">6.2 mm</quantity>
; wing length:
<quantity id="4CAE9B774608FFD7FCE2FDA56DD1B491" box="[811,900,622,647]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="8" pageNumber="309" unit="mm" value="5.9">5.9 mm</quantity>
.
</paragraph>
<paragraph id="8BE936924608FFD7FF0EFD596CE3B4E5" blockId="8.[151,1436,151,827]" pageId="8" pageNumber="309">
Similar to male, differing in the following respects: frons with 5 pairs of setae (frontals and orbitals) along entire length (not confined to area close to lunule as in males); ovipositor long, with microtrichia on both sides (
<figureCitation id="136D2A174608FFD7FF56FD116F74B4E5" box="[159,289,730,755]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,264,1648,1671]" captionTargetBox="[151,1414,197,1571]" captionTargetId="figure@7.[151,1436,193,1628]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Helina harrisorum sp. nov.: 13. cercal plate and surstyli, dorsal view; 14. epandrium, cercal plate and surstylus, lateral view; 15. aedeagus, lateral view; 16. ovipositor, dorsal view; 17. ovipositor, ventral view and spermathecae. Helina ferfriniorum sp. nov.: 18. ovipositor, lateral view; 19. egg." httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="8" pageNumber="309">Figs 16–17</figureCitation>
); spermathecae round, as in
<figureCitation id="136D2A174608FFD7FD96FD116CE7B4E5" box="[607,690,730,755]" captionStart="FIGURES 13 – 19" captionStartId="7.[151,264,1648,1671]" captionTargetBox="[151,1414,197,1571]" captionTargetId="figure@7.[151,1436,193,1628]" captionTargetPageId="7" captionText="FIGURES 13 – 19. Helina harrisorum sp. nov.: 13. cercal plate and surstyli, dorsal view; 14. epandrium, cercal plate and surstylus, lateral view; 15. aedeagus, lateral view; 16. ovipositor, dorsal view; 17. ovipositor, ventral view and spermathecae. Helina ferfriniorum sp. nov.: 18. ovipositor, lateral view; 19. egg." httpUri="https://zenodo.org/record/229941/files/figure.png" pageId="8" pageNumber="309">Fig. 17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C65194608FFD7FF0EFD356DE0B501" box="[199,949,766,791]" pageId="8" pageNumber="309" type="distribution">
<paragraph id="8BE936924608FFD7FF0EFD356DE0B501" blockId="8.[151,1436,151,827]" box="[199,949,766,791]" pageId="8" pageNumber="309">
<emphasis id="B922EA804608FFD7FF0EFD356F0CB501" bold="true" box="[199,345,766,791]" pageId="8" pageNumber="309">Distribution</emphasis>
.
<collectingCountry id="F34176024608FFD7FEAEFD356CD4B501" box="[359,641,766,791]" name="South Africa" pageId="8" pageNumber="309">Republic of South Africa</collectingCountry>
(Free State, Eastern Cape).
</paragraph>
</subSubSection>
<subSubSection id="C34C65194608FFD7FF0EFCE96BC4B52D" box="[199,1425,802,827]" pageId="8" pageNumber="309" type="etymology">
<paragraph id="8BE936924608FFD7FF0EFCE96BC4B52D" blockId="8.[151,1436,151,827]" box="[199,1425,802,827]" pageId="8" pageNumber="309">
<emphasis id="B922EA804608FFD7FF0EFCE96F13B52D" bold="true" box="[199,326,802,827]" pageId="8" pageNumber="309">Etymology</emphasis>
. The species is dedicated to Sandra and John Harris and is a Latinized noun in the genitive plural.
</paragraph>
</subSubSection>
</treatment>
</document>
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