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<document id="9757EA66D7F644A2EF8CFA8E1500717E" ID-DOI="10.11646/zootaxa.5020.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5223173" ID-ZooBank="4FF882DF-334F-49C8-A576-4192B5F2654C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629388374973" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2021" docId="FA08E75DAA338053FDC03575958A699D" docLanguage="en" docName="zootaxa.5020.2.1.pdf" docOrigin="Zootaxa 5020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habetia pedala Ingrisch 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="214" masterDocId="06319F25AA3F805EFD57366B91686D58" masterDocTitle="Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="213" updateTime="1699071125630" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2937A06334F866ECBADE153947CE5DF5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BD64E9077DE867EF0AECCE95F1CA912D">
<mods:title id="E5132606AA72F9EDC17C15536C90DC4E">Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)</mods:title>
</mods:titleInfo>
<mods:name id="D4E8F985E61BBA7A5313C208C4048C74" type="personal">
<mods:role id="6B1E5BD1B4CCA7A44CC434E0A13DA5E4">
<mods:roleTerm id="F0ABD8B06854976C21DC9CF2697DDDF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE219A5FF1C6C89B22D4B187DB4A43A8">Ingrisch, Sigfrid</mods:namePart>
</mods:name>
<mods:typeOfResource id="4A2AF53912860CDEB5EA7E5C511E0231">text</mods:typeOfResource>
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<mods:titleInfo id="C836159DE79191A1A043CA3DD8DCB335">
<mods:title id="B56635FC747D6AF0E5D61363854A8BF7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5D769CEC0D22E24B1F323C664C71419A">
<mods:date id="8B96AE5F852BF4C8119ED452418F19E2">2021</mods:date>
<mods:detail id="5E2FC8AF65DF76002F97E2E441A7BDC1" type="pubDate">
<mods:number id="4B5AC64141F60AAD89BE86C0B7B2574C">2021-08-12</mods:number>
</mods:detail>
<mods:detail id="023078F654EB040DD9B2A0B7DF3BF9F7" type="volume">
<mods:number id="380D828BFCEA66275CA2A194AC19EE51">5020</mods:number>
</mods:detail>
<mods:detail id="87F95389A7BDD3387E5D19F5DB4CAEA7" type="issue">
<mods:number id="49D60872BC414EDFC9293E31762CDE53">2</mods:number>
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<mods:classification id="0CDE63AF96F2BAB2A5826B2D4D5C2A42">journal article</mods:classification>
<mods:identifier id="714A6F596C32CAED9BC06D01924383D3" type="DOI">10.11646/zootaxa.5020.2.1</mods:identifier>
<mods:identifier id="AECE88147A28E9D1D8A67C02858E77AD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EABD328C020747A32B9F7CE88F98A046" type="Zenodo-Dep">5223173</mods:identifier>
<mods:identifier id="7866CBF74B03DD436F6A0F2DEA602FCE" type="ZooBank">4FF882DF-334F-49C8-A576-4192B5F2654C</mods:identifier>
</mods:mods>
<treatment id="FA08E75DAA338053FDC03575958A699D" LSID="urn:lsid:plazi:treatment:FA08E75DAA338053FDC03575958A699D" httpUri="http://treatment.plazi.org/id/FA08E75DAA338053FDC03575958A699D" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213">
<subSubSection id="3ABB05C0AA338052FDC0357590DB6E61" box="[151,435,798,825]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="721E564BAA338052FDC0357590DB6E61" blockId="12.[151,756,798,897]" box="[151,435,798,825]" pageId="12" pageNumber="213">
<heading id="2956E127AA338052FDC0357590DB6E61" bold="true" box="[151,435,798,825]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="40D58A59AA338052FDC0357590DB6E61" bold="true" box="[151,435,798,825]" pageId="12" pageNumber="213">
<taxonomicName id="B5A12DC8AA338052FDC0357590276E61" ID-CoL="8QJWW" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[151,335,798,825]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA338052FDC0357590276E61" bold="true" box="[151,335,798,825]" italics="true" pageId="12" pageNumber="213">Habetia pedala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BE63722AA338052FC01357490DB6E61" box="[342,435,799,825]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA338052FDC0352893F76ED9" pageId="12" pageNumber="213" type="description">
<paragraph id="721E564BAA338052FDC03528939C6E05" blockId="12.[151,756,798,897]" box="[151,756,834,861]" pageId="12" pageNumber="213">
<figureCitation id="EA9A4ACEAA338052FDC0352891986E05" box="[151,240,835,861]" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="12" pageNumber="213">Figs 1C</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FDAA3528907E6E05" box="[253,278,835,861]" captionStart="FIGURE 3" captionStartId="17.[151,250,1860,1884]" captionTargetBox="[177,1409,182,1837]" captionTargetId="figure-15@17.[177,1409,181,1837]" captionTargetPageId="17" captionText="FIGURE 3. A, Habetia pilleata sp. nov. male (Wau 1200m); B, H. pallida sp. nov. female (Hollandia); C, H. tuta sp. nov. male (Quelllager); D–F, H. curvata sp. nov. (D, male Albatros Bivak, E–F, female Prauwenbivak); G, H. spada (Brunner, 1898) female holotype (Konstantinhafen); H, H. multispinulosa Griffini, 1908 female holotype (Dilo); I, H. pedala sp. nov. female (Toricelli Mountains).—Frons (A–E, G–I), female dorsal view (F). Scales 5 mm (A–E, G–I), 10 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5223179" httpUri="https://zenodo.org/record/5223179/files/figure.png" pageId="12" pageNumber="213">3I</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FC753529902F6E05" box="[290,327,834,861]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="12" pageNumber="213">5D</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FC04352890FD6E05" box="[339,405,835,861]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="12" pageNumber="213">8C–E</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FCF7352890B86E05" box="[416,464,835,861]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (A–E), 1 mm (F–K)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="12" pageNumber="213">11C</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FC8B352893466E05" box="[476,558,835,861]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: A–C, H. tuta sp. nov. (A Quelllager, B–C Töpferfluss); D–E, H. pedala sp. nov. (Toricelli Mts); F–I, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="12" pageNumber="213">12D–E</figureCitation>
,
<figureCitation id="EA9A4ACEAA338052FF6E352893036E05" box="[569,619,835,861]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); G–H, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="12" pageNumber="213">18C</figureCitation>
, Map 1(15).
</paragraph>
<paragraph id="721E564BAA338052FDC0350C93F76ED9" blockId="12.[151,756,798,897]" box="[151,671,871,897]" pageId="12" pageNumber="213">
<taxonomicName id="B5A12DC8AA338052FDC0350C90B26ED9" authority="Karny (1912)" authorityName="Karny" authorityYear="1912" box="[151,474,871,897]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052FDC0350C90536ED9" box="[151,315,871,897]" italics="true" pageId="12" pageNumber="213">Habetia spada</emphasis>
Karny (1912)
</taxonomicName>
misidentification
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA338052FDC035C490166B75" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="721E564BAA338052FDC035C493056949" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="213">
<materialsCitation id="C2C95C16AA338052FDC035C493056949" collectingDate="1910-01-01" collectingDateMax="1910-01-31" collectingDateMin="1910-01-01" collectionCode="SMTD" collectorName="Kais. Wilhelmsland Toricelli Gebirge Dr. Schlaginhaufen &amp; Senckenberg Museum &amp; Tierkunde Dresden" country="Papua New Guinea" latitude="-3.5" location="Toricelli Mountains" longLatPrecision="1307" longitude="142.0" pageId="12" pageNumber="213" specimenCount="1" stateProvince="West Sepik" typeStatus="holotype">
<emphasis id="40D58A59AA338052FDC035C490086E90" bold="true" box="[151,352,942,969]" pageId="12" pageNumber="213">
<typeStatus id="AD1AE8E9AA338052FDC035C4906C6E91" box="[151,260,943,969]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="0AB616DBAA338052FC3F35C4932C6E91" box="[360,580,943,969]" name="Papua New Guinea" pageId="12" pageNumber="213">Papua New Guinea</collectingCountry>
:
<collectingRegion id="B06598A9AA338052FF1835C493B86E91" box="[591,720,943,969]" country="Papua New Guinea" name="West Sepik" pageId="12" pageNumber="213">West Sepik</collectingRegion>
,
<location id="777E0090AA338052FF8C35C492D36E91" LSID="urn:lsid:plazi:treatment:FA08E75DAA338053FDC03575958A699D:777E0090AA338052FF8C35C492D36E91" box="[731,955,943,969]" country="Papua New Guinea" latitude="-3.5" longLatPrecision="1307" longitude="142.0" name="Toricelli Mountains" pageId="12" pageNumber="213" stateProvince="West Sepik">Toricelli Mountains</location>
, (
<geoCoordinate id="1795308CAA338052FE9835C495776E91" box="[975,1055,943,969]" degrees="3" direction="south" minutes="30" orientation="latitude" pageId="12" pageNumber="213" precision="925" value="-3.5">3°30’S</geoCoordinate>
,
<geoCoordinate id="1795308CAA338052F97B35C495E46E91" box="[1068,1164,943,969]" degrees="142" direction="east" minutes="0" orientation="longitude" pageId="12" pageNumber="213" precision="925" value="142.0">142°0’E</geoCoordinate>
),
<date id="061F708BAA338052F9F635C494476E91" box="[1185,1327,943,969]" pageId="12" pageNumber="213" value="1910-01-01" valueMax="1910-01-31" valueMin="1910-01-01">
<collectingDate id="165B8963AA338052F9F635C494476E91" box="[1185,1327,943,969]" pageId="12" pageNumber="213" value="1910-01-01" valueMax="1910-01-31" valueMin="1910-01-01">1.–31.i.1910</collectingDate>
</date>
, [original labels: “
<collectorName id="DF54339DAA338052FC5135B892CE6EB5" box="[262,934,979,1005]" pageId="12" pageNumber="213">Kais. Wilhelmsland Toricelli Gebirge Dr. Schlaginhaufen</collectorName>
“ and “1910 1“]—depository:
<collectorName id="DF54339DAA338052F85B35B891916949" pageId="12" pageNumber="213">Senckenberg Museum</collectorName>
für
<collectorName id="DF54339DAA338052FC70359C90956949" box="[295,509,1015,1041]" pageId="12" pageNumber="213">Tierkunde Dresden</collectorName>
(
<collectionCode id="14B0CE8EAA338052FF59359C93096949" box="[526,609,1015,1041]" pageId="12" pageNumber="213">SMTD</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="721E564BAA338052FD90327195A1696D" blockId="12.[151,1437,942,2013]" box="[199,1225,1050,1077]" pageId="12" pageNumber="213">
<materialsCitation id="C2C95C16AA338052FD90327195AD696D" box="[199,1221,1050,1077]" collectingDate="1910-01-01" collectingDateMax="1910-01-31" collectingDateMin="1910-01-01" collectionCode="SMTD" collectorName="Kais. Wilhelmsland Toricelli Gebirge Dr. Schlaginhaufen &amp; Senckenberg Museum &amp; Tierkunde Dresden" country="Papua New Guinea" latitude="-3.5" location="Toricelli Mountains" longLatPrecision="1307" longitude="142.0" pageId="12" pageNumber="213" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="West Sepik" typeStatus="paratype">
<emphasis id="40D58A59AA338052FD903271909E696C" bold="true" box="[199,502,1050,1076]" pageId="12" pageNumber="213">Other specimens studied:</emphasis>
same data as holotype—
<specimenCount id="64A79DC2AA338052FE4632709217696D" box="[785,895,1051,1077]" pageId="12" pageNumber="213" type="female">2 females</specimenCount>
,
<specimenCount id="64A79DC2AA338052FEDD327092BD696D" box="[906,981,1051,1077]" pageId="12" pageNumber="213" type="male">1 male</specimenCount>
(
<typeStatus id="AD1AE8E9AA338052FEB33270953B696D" box="[996,1107,1051,1077]" pageId="12" pageNumber="213" type="paratype">paratypes</typeStatus>
) (
<collectionCode id="14B0CE8EAA338052F93E327095D5696D" box="[1129,1213,1051,1077]" pageId="12" pageNumber="213">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="721E564BAA338052FD90325590166B75" blockId="12.[151,1437,942,2013]" pageId="12" pageNumber="213">
<emphasis id="40D58A59AA338052FD90325590E36901" bold="true" box="[199,395,1086,1113]" pageId="12" pageNumber="213">Taxonomic note.</emphasis>
When Karny studied the
<specimenCount id="64A79DC2AA338052FFFF325492276901" box="[680,847,1087,1113]" pageId="12" pageNumber="213" type="generic">five specimens</specimenCount>
of
<taxonomicName id="B5A12DC8AA338052FE24325492A56901" authorityName="Kirby" authorityYear="1906" box="[883,973,1087,1113]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="40D58A59AA338052FE24325492A56901" box="[883,973,1087,1113]" italics="true" pageId="12" pageNumber="213">Habetia</emphasis>
</taxonomicName>
in SMTD Dresden, he had just published the volume on
<taxonomicName id="B5A12DC8AA338052FC14320890D16925" authorityName="Redtenbacher" authorityYear="1891" box="[323,441,1123,1149]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="tribe" tribe="Agraeciini">Agraeciini</taxonomicName>
in Genera Insectorum (
<bibRefCitation id="16302BBAAA338052FF9E3208920B6925" author="Karny, H. H." box="[713,867,1123,1149]" pageId="12" pageNumber="213" pagination="1 - 47" refId="ref43063" refString="Karny, H. H. (1912 a) Orthoptera Fam. Locustidae Subfam. Agraeciinae. Genera Insectorum, 141, 1 - 47, pls. 1 - 8." type="journal article" year="1912">Karny 1912a</bibRefCitation>
). At the time when he prepared the manuscript to that work, he could not have seen any true male of the genus
<taxonomicName id="B5A12DC8AA338052FE1432EC92F569F9" authorityName="Kirby" authorityYear="1906" box="[835,925,1159,1185]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="40D58A59AA338052FE1432EC92F569F9" box="[835,925,1159,1185]" italics="true" pageId="12" pageNumber="213">Habetia</emphasis>
</taxonomicName>
, since he based the description of the male of that genus exclusively on the single male of “
<taxonomicName id="B5A12DC8AA338052FFCD32C092D2699D" authority="Karny, 1911" authorityName="Karny" authorityYear="1911" box="[666,954,1195,1221]" class="Insecta" family="Tettigoniidae" genus="Parahabetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pictifrons">
<emphasis id="40D58A59AA338052FFCD32C0924F699D" box="[666,807,1195,1221]" italics="true" pageId="12" pageNumber="213">H. pictifrons</emphasis>
<bibRefCitation id="16302BBAAA338052FE7832C092D2699D" author="Karny, H. H." box="[815,954,1195,1221]" pageId="12" pageNumber="213" pagination="334 - 347" refId="ref43037" refString="Karny, H. H. (1911) Descriptiones Conocephalidarum novarum. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 61, 334 - 347." type="journal article" year="1911">Karny, 1911</bibRefCitation>
</taxonomicName>
” and explicitly mentions in a footnote that “the description of the male end of abdomen refers only to
<taxonomicName id="B5A12DC8AA338052FE6A32A492A269B1" authorityName="Karny" authorityYear="1911" box="[829,970,1231,1257]" class="Insecta" family="Tettigoniidae" genus="Parahabetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pictifrons">
<emphasis id="40D58A59AA338052FE6A32A492A269B1" box="[829,970,1231,1257]" italics="true" pageId="12" pageNumber="213">H. pictifrons</emphasis>
</taxonomicName>
” (translated from German). That species is moved below to the genus
<taxonomicName id="B5A12DC8AA338052FCBD3298931E6855" authority="The" authorityName="Ingrisch" authorityYear="2021" box="[490,630,1267,1293]" class="Insecta" family="Tettigoniidae" genus="Parahabetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="40D58A59AA338052FCBD3298931E6855" box="[490,630,1267,1293]" italics="true" pageId="12" pageNumber="213">Parahabetia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BE63722AA338052FF283298938F6855" box="[639,743,1267,1293]" pageId="12" pageNumber="213" rank="genus">gen. nov.</taxonomicNameLabel>
The species introduced here as
<taxonomicName id="B5A12DC8AA338052F90F329895A06855" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[1112,1224,1267,1293]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA338052F90F329895A06855" box="[1112,1224,1267,1293]" italics="true" pageId="12" pageNumber="213">H. pedala</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA338052F986329894456855" bold="true" box="[1233,1325,1267,1293]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="5BE63722AA338052F986329894456855" box="[1233,1325,1267,1293]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
had been reported by
<bibRefCitation id="16302BBAAA338052FC75337C90A2686A" author="Karny, H. H." box="[290,458,1303,1330]" pageId="12" pageNumber="213" pagination="3 - 23" refId="ref43096" refString="Karny, H. H. (1912 b) Conocephaliden (Orthoptera Locustoidea) aus Neuguinea hauptsachlich gesammelt von Dr. O Schlaginhaufen. Abhandlungen und Berichte des zoologischen und anthropologisch-ethnographischen Museums zu Dresden, 14, 3 - 23." type="journal article" year="1912">Karny (1912b)</bibRefCitation>
as
<taxonomicName id="B5A12DC8AA338052FCA4337C923E6869" authority="(Brunner, 1898)" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[499,854,1303,1329]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052FCA4337C93F16869" box="[499,665,1303,1329]" italics="true" pageId="12" pageNumber="213">Habetia spada</emphasis>
(
<bibRefCitation id="16302BBAAA338052FFFE337C92266869" author="Brunner von Wattenwyl, C." box="[681,846,1303,1329]" pageId="12" pageNumber="213" pagination="193 - 288" refId="ref42428" refString="Brunner von Wattenwyl, C. (1898) Orthopteren des Malayischen Archipels, gesammelt von Prof. Dr. W. Kukenthal in den Jahren 1893 und 1894. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, Frankfurt, M 24, 193 - 288, pls. 16 - 20." type="journal article" year="1898">Brunner, 1898</bibRefCitation>
)
</taxonomicName>
. The SMTD holds
<specimenCount id="64A79DC2AA338052F961337C95886869" box="[1078,1248,1303,1329]" pageId="12" pageNumber="213" type="generic">five specimens</specimenCount>
of
<taxonomicName id="B5A12DC8AA338052F85F337C940A6869" authorityName="Kirby" authorityYear="1906" box="[1288,1378,1303,1329]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="40D58A59AA338052F85F337C940A6869" box="[1288,1378,1303,1329]" italics="true" pageId="12" pageNumber="213">Habetia</emphasis>
</taxonomicName>
,
<specimenCount id="64A79DC2AA338052F825337C91B0680D" pageId="12" pageNumber="213" type="male">two males</specimenCount>
and
<specimenCount id="64A79DC2AA338052FC46335090F1680D" box="[273,409,1339,1365]" pageId="12" pageNumber="213" type="female">two females</specimenCount>
from the Toricelli Mountains and a single male labeled Bongu. The male from Bongu was described as
<taxonomicName id="B5A12DC8AA338052FC70333493556821" authority="Karny, 1912" authorityName="Karny" authorityYear="1912" box="[295,573,1375,1401]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA338052FC70333490C16820" box="[295,425,1375,1400]" italics="true" pageId="12" pageNumber="213">H. imitatrix</emphasis>
Karny, 1912
</taxonomicName>
, although its locality lies close to the type locality of
<taxonomicName id="B5A12DC8AA338052F9C73334959D6821" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[1168,1269,1375,1401]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052F9C73334959D6821" box="[1168,1269,1375,1401]" italics="true" pageId="12" pageNumber="213">H. spada</emphasis>
</taxonomicName>
, while the pair from the Toricelli Mountains had been identified as
<taxonomicName id="B5A12DC8AA338052FFBA33E8923C68C5" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[749,852,1411,1437]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052FFBA33E8923C68C5" box="[749,852,1411,1437]" italics="true" pageId="12" pageNumber="213">H. spada</emphasis>
</taxonomicName>
by
<bibRefCitation id="16302BBAAA338052FED533E8954568C6" author="Karny, H. H." box="[898,1069,1411,1438]" pageId="12" pageNumber="213" pagination="3 - 23" refId="ref43096" refString="Karny, H. H. (1912 b) Conocephaliden (Orthoptera Locustoidea) aus Neuguinea hauptsachlich gesammelt von Dr. O Schlaginhaufen. Abhandlungen und Berichte des zoologischen und anthropologisch-ethnographischen Museums zu Dresden, 14, 3 - 23." type="journal article" year="1912">Karny (1912b)</bibRefCitation>
. However, the females from the Toricelli Mountains show distinct differences in the shape of the subgenital plate compared to that of the
<typeStatus id="AD1AE8E9AA338052F84A33CC94176899" box="[1309,1407,1447,1473]" pageId="12" pageNumber="213" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B5A12DC8AA338052FDC033A0919368BD" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[151,251,1483,1509]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052FDC033A0919368BD" box="[151,251,1483,1509]" italics="true" pageId="12" pageNumber="213">H. spada</emphasis>
</taxonomicName>
, and the males do not show the striking modifications of the subgenital plate found in the male from Bongu but is closer to
<taxonomicName id="B5A12DC8AA338052FC15338490DB6B51" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[322,435,1519,1545]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida" status="sp. nov.">
<emphasis id="40D58A59AA338052FC15338490DB6B51" box="[322,435,1519,1545]" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA338052FCED3384937C6B51" bold="true" box="[442,532,1519,1545]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="5BE63722AA338052FCED3384937C6B51" box="[442,532,1519,1545]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Thus, the
<taxonomicName id="B5A12DC8AA338052FFDE3384938B6B51" authorityName="Kirby" authorityYear="1906" box="[649,739,1519,1545]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="40D58A59AA338052FFDE3384938B6B51" box="[649,739,1519,1545]" italics="true" pageId="12" pageNumber="213">Habetia</emphasis>
</taxonomicName>
specimens from the Torricelli Mountains represent a formerly undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA338053FD90305D91846DA1" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="721E564BAA338053FD90305D91846DA1" blockId="12.[151,1437,942,2013]" lastBlockId="13.[151,1437,151,1221]" lastPageId="13" lastPageNumber="214" pageId="12" pageNumber="213">
<emphasis id="40D58A59AA338052FD90305D902A6B08" bold="true" box="[199,322,1590,1616]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
The new species has a dorsal (inner) crest at the end of the male subgenital plate (
<figureCitation id="EA9A4ACEAA338052F84D305C941F6B09" box="[1306,1399,1591,1617]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="12" pageNumber="213">Fig. 8D</figureCitation>
), a character that it shares with
<taxonomicName id="B5A12DC8AA338052FC80303093226B2D" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[471,586,1627,1653]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida" status="sp. nov.">
<emphasis id="40D58A59AA338052FC80303093226B2D" box="[471,586,1627,1653]" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA338052FF05303093C56B2D" bold="true" box="[594,685,1627,1653]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="5BE63722AA338052FF05303093C56B2D" box="[594,685,1627,1653]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, in
<taxonomicName id="B5A12DC8AA338052FE10303092DE6B2D" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[839,950,1627,1653]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA338052FE10303092DE6B2D" box="[839,950,1627,1653]" italics="true" pageId="12" pageNumber="213">H. pedala</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA338052FEE8303095716B2D" bold="true" box="[959,1049,1627,1653]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="5BE63722AA338052FEE8303095716B2D" box="[959,1049,1627,1653]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
that crest is higher, roughly twice as high as the diameter of the rounded ends of the subgenital plate and only faintly incised in middle, while in
<taxonomicName id="B5A12DC8AA338052F8D63014918E6BE5" authorityName="Ingrisch" authorityYear="2021" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="40D58A59AA338052F8D63014918E6BE5" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
it is only as high as one diameter of the rounded ends of the subgenital plate and more distinctly incised. Moreover, the incised apical area of the subgenital plate is markedly longer than in
<taxonomicName id="B5A12DC8AA338052F91130AC95DF6BB9" authorityName="Ingrisch" authorityYear="2021" box="[1094,1207,1735,1761]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="40D58A59AA338052F91130AC95DF6BB9" box="[1094,1207,1735,1761]" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
. Further differences are the longer male cerci of
<taxonomicName id="B5A12DC8AA338052FC813080932D6A5D" authorityName="Ingrisch" authorityYear="2021" box="[470,581,1771,1797]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pedala">
<emphasis id="40D58A59AA338052FC813080932D6A5D" box="[470,581,1771,1797]" italics="true" pageId="12" pageNumber="213">H. pedala</emphasis>
</taxonomicName>
, that have the internal projection inserted little more proximad instead of at end and the acute apical teeth of that process are longer and of different shape. The titillators are similar to those of
<taxonomicName id="B5A12DC8AA338052FDC0315890616A15" authorityName="Ingrisch" authorityYear="2021" box="[151,265,1843,1869]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="40D58A59AA338052FDC0315890616A15" box="[151,265,1843,1869]" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
and
<taxonomicName id="B5A12DC8AA338052FC17315890D16A14" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[320,441,1843,1868]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="curvata" status="sp. nov.">
<emphasis id="40D58A59AA338052FC17315890D16A14" box="[320,441,1843,1868]" italics="true" pageId="12" pageNumber="213">H. curvata</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA338052FCE8315893716A15" bold="true" box="[447,537,1843,1869]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="5BE63722AA338052FCE8315893716A15" box="[447,537,1843,1869]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but have the apical area narrow, not widened as in those species, and the dorsodistal margin is markedly constricted in sub-apical area. From the latter species,
<taxonomicName id="B5A12DC8AA338052F94C313C95E16A29" authorityName="Ingrisch" authorityYear="2021" box="[1051,1161,1879,1905]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pedala">
<emphasis id="40D58A59AA338052F94C313C95E16A29" box="[1051,1161,1879,1905]" italics="true" pageId="12" pageNumber="213">H. pedala</emphasis>
</taxonomicName>
also differs by the small and narrow styli and by the bi-spinose tip of the internal process of the cerci. The female subgenital plate of the new species is similar to that of
<taxonomicName id="B5A12DC8AA338052FF5B31F493176AE1" authorityName="Ingrisch" authorityYear="2021" box="[524,639,1951,1977]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="40D58A59AA338052FF5B31F493176AE1" box="[524,639,1951,1977]" italics="true" pageId="12" pageNumber="213">H. pallida</emphasis>
</taxonomicName>
but has the apical margin more strongly upcurved between the apical spine-like projections which thus appear stouter at base and shorter than in the latter species. From
<taxonomicName id="B5A12DC8AA338052F85431A894026A85" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[1283,1386,1987,2013]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA338052F85431A894026A85" box="[1283,1386,1987,2013]" italics="true" pageId="12" pageNumber="213">H. spada</emphasis>
</taxonomicName>
that looks otherwise similar,
<taxonomicName id="B5A12DC8AA328053FCF136FC937B6DE9" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[422,531,151,177]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA328053FCF136FC937B6DE9" box="[422,531,151,177]" italics="true" pageId="13" pageNumber="214">H. pedala</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA328053FF4E36FC931A6DE9" bold="true" box="[537,626,151,177]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="5BE63722AA328053FF4E36FC931A6DE9" box="[537,626,151,177]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs by the female subgenital plate that is markedly convex with a pair of stout, at tip spinose projections while in
<taxonomicName id="B5A12DC8AA328053FF2836D0938C6D8D" baseAuthorityName="Brunner" baseAuthorityYear="1898" box="[639,740,187,213]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="spada">
<emphasis id="40D58A59AA328053FF2836D0938C6D8D" box="[639,740,187,213]" italics="true" pageId="13" pageNumber="214">H. spada</emphasis>
</taxonomicName>
the subgenital plate is flattened and the apical projections are narrow.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA328053FD903769922B69F9" pageId="13" pageNumber="214" type="description">
<paragraph id="721E564BAA328053FD90376994276C19" blockId="13.[151,1437,151,1221]" pageId="13" pageNumber="214">
<emphasis id="40D58A59AA328053FD90376990316C44" bold="true" box="[199,345,258,284]" pageId="13" pageNumber="214">Description.</emphasis>
Coloration of face uniformly pale, of general color; mandibles fully black. Femora with the following number of spines on ventral margins: (1) a 7, p 9; (2) a 6–8, p 3–4; (3) a 12–14, p about 14 (n=2).
</paragraph>
<paragraph id="721E564BAA328053FD90372093386EFD" blockId="13.[151,1437,151,1221]" pageId="13" pageNumber="214">
Male. Stridulatory file on underside of left tegmen
<quantity id="B559FBAEAA328053FE573721920C6C3D" box="[768,868,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="13" pageNumber="214" unit="mm" value="2.55">2.55 mm</quantity>
long with 74 teeth and with a distinct step in distal area: curved narrow basal area ca
<quantity id="B559FBAEAA328053FF59370493196CD1" box="[526,625,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="214" unit="mm" value="0.37">0.37 mm</quantity>
(broken) with 12 teeth, central area 1,55 mm with 35 teeth, step 0,17 mm with 6 teeth, narrow apical area 0,51 mm with 23 teeth of which 11 clearly separated and 12 of decreasing size at end (
<figureCitation id="EA9A4ACEAA328053FDF737DC906E6C89" box="[160,262,439,465]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); G–H, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="13" pageNumber="214">Fig. 18C</figureCitation>
). Tenth abdominal tergite nearly parallel-sided with down-curved lateral areas, with medial furrow and a narrow apical incision in middle, internal margin of that area down-curved and strongly setose (
<figureCitation id="EA9A4ACEAA328053F99937B0944F6CAD" box="[1230,1319,475,501]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="13" pageNumber="214">Fig. 8C</figureCitation>
). Epiproct little widening toward sub-truncate apical margin and with a fine medial furrow. Cerci narrow elongate, markedly curved in apical half, behind basal area with an internal carina that delimits an excavation of the internal surface on dorsal side; that area with a wide compressed internal projection that divides in apical area into two branches that are little curved proximad, the ventro-proximal branch little longer than the dorso-apical branch; both terminate into an acute spine (
<figureCitation id="EA9A4ACEAA328053FC2F34E490906FF1" box="[376,504,655,681]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="13" pageNumber="214">Figs 8C–D</figureCitation>
). Subgenital plate with roundly incised basal margin, with lateral rims along dorsal margins; ventral disc with convex lateral margins, that become at end slightly diverging; from apex deep and widely incised; styli rather short and thin (
<figureCitation id="EA9A4ACEAA328053FF4A34BC93E96FA9" box="[541,641,727,753]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (A–E), 1 mm (F–K)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="13" pageNumber="214">Fig. 11C</figureCitation>
); at dorsal side of subgenital plate, just before the incision of the apical margin, with a rather high lamellar elevation with slightly bilobate dorsal margin (
<figureCitation id="EA9A4ACEAA328053F965349095E56E4D" box="[1074,1165,763,789]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="13" pageNumber="214">Fig. 8D</figureCitation>
). Titillators at base little widened and curved ventro-distad, in central area nearly parallel-sided, straight, at transition to apical area, the dorso-proximal margin is bent posteriorly with nearly straight dorso-distal and concave ventro-proximal margin, about in mid-length the dorsal margin is constricted and afterward convex, at end titillators curved mediad and narrowed into obtuse tip (
<figureCitation id="EA9A4ACEAA328053FCE035E0932C6EFD" box="[439,580,907,933]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: A–C, H. tuta sp. nov. (A Quelllager, B–C Töpferfluss); D–E, H. pedala sp. nov. (Toricelli Mts); F–I, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="13" pageNumber="214">Figs 12D–E</figureCitation>
).
</paragraph>
<paragraph id="721E564BAA328053FD9035C491A0696D" blockId="13.[151,1437,151,1221]" pageId="13" pageNumber="214">
Female. Subgenital plate with a fine membranous medial furrow and with strongly up-bent lateral and apical margins; at up-bent apico-lateral angles terminating into a pair of stout cones with acute tips (
<figureCitation id="EA9A4ACEAA328053F9E135B894786EB5" box="[1206,1296,978,1005]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="13" pageNumber="214">Fig. 5D</figureCitation>
). Ovipositor about twice as long as body; behind basal constriction straight with dorsal valves faintly convex but not raised (
<figureCitation id="EA9A4ACEAA328053F838359C91D2696D" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="13" pageNumber="214">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="721E564BAA328053FD903254922B69F9" blockId="13.[151,1437,151,1221]" pageId="13" pageNumber="214">
Measurements (
<specimenCount id="64A79DC2AA328053FC2C325490BE6901" box="[379,470,1087,1113]" pageId="13" pageNumber="214" type="male">2 males</specimenCount>
,
<specimenCount id="64A79DC2AA328053FCB43254933B6902" box="[483,595,1087,1114]" pageId="13" pageNumber="214" type="female">2 females</specimenCount>
).—Body w/o wings: male 34–35, female 30–32; pronotum: male 9.0–9.6, female 9.6–9.8; tegmen: male 32.0–32.5, female 36.5–38.0; hind femur: male 22.5–24.0, female 26.5; ovipositor length: female 67–68; ovipositor height: female
<quantity id="B559FBAEAA328053FFE332ED925669F9" box="[692,830,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="13" pageNumber="214" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA328053FD9032C0958A699D" box="[199,1250,1195,1221]" pageId="13" pageNumber="214" type="etymology">
<paragraph id="721E564BAA328053FD9032C0958A699D" blockId="13.[151,1437,151,1221]" box="[199,1250,1195,1221]" pageId="13" pageNumber="214">
<emphasis id="40D58A59AA328053FD9032C09026699D" bold="true" box="[199,334,1195,1221]" pageId="13" pageNumber="214">Etymology.</emphasis>
Named for the large size and long ovipositor; from Latin
<emphasis id="40D58A59AA328053FE8132C0954F699D" box="[982,1063,1195,1221]" italics="true" pageId="13" pageNumber="214">pedalis</emphasis>
= one foot long.
</paragraph>
</subSubSection>
</treatment>
</document>
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