277 lines
74 KiB
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277 lines
74 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.6.1.10" ID-ISSN="2624-2834" ID-Zenodo-Dep="7754820" ID-ZooBank="B822ED53-D559-4DD8-8803-3F2850A00445" approvalRequired="14" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="11" approvalRequired_for_treatments="1" checkinTime="1679380739061" checkinUser="plazi" docAuthor="XUAN, QIANG, CAI, CHEN-YANG & HUANG, DI-YING" docDate="2023" docId="F308E17F4358FFEAFCCD4B45FA701CC2" docLanguage="en" docName="palaeoentomology.6.1.10.pdf" docOrigin="Palaeoentomology 6 (1)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.1.10" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Betaburmesebuthus bellus Lourenco 2016" docType="treatment" docVersion="2" lastPageNumber="73" masterDocId="0F3199074350FFE3FFE74E07FFFE192E" masterDocTitle="Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)" masterLastPageNumber="101" masterPageNumber="64" pageNumber="72" updateTime="1679381284402" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Revision of palaeoburmesebuthid scorpions in mid-Cretaceous amber from northern Myanmar (Scorpiones: Buthoidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>XUAN, QIANG</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>CAI, CHEN-YANG</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>HUANG, DI-YING</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-28</mods:number>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>64</mods:start>
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<mods:end>101</mods:end>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.6.1.10</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.6.1.10</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7754820</mods:identifier>
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<mods:identifier type="ZooBank">B822ED53-D559-4DD8-8803-3F2850A00445</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7754824" ID-Zenodo-Dep="7754824" LSID="urn:lsid:plazi:treatment:F308E17F4358FFEAFCCD4B45FA701CC2" httpUri="http://treatment.plazi.org/id/F308E17F4358FFEAFCCD4B45FA701CC2" lastPageId="9" lastPageNumber="73" pageId="8" pageNumber="72">
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<subSubSection box="[810,1307,1346,1373]" pageId="8" pageNumber="72" type="nomenclature">
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<paragraph blockId="8.[810,1307,1346,1408]" box="[810,1307,1346,1373]" pageId="8" pageNumber="72">
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<heading bold="true" box="[810,1307,1346,1373]" fontSize="10" level="2" pageId="8" pageNumber="72" reason="3">
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<taxonomicName authority="Lourenco, 2016" authorityName="Lourenco" authorityYear="2016" box="[810,1307,1346,1373]" class="Arachnida" family="Palaeoburmesebuthidae" genus="Betaburmesebuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="72" phylum="Arthropoda" rank="species" species="bellus">
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<emphasis bold="true" box="[810,1307,1346,1373]" pageId="8" pageNumber="72">
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<emphasis bold="true" box="[810,1115,1346,1373]" italics="true" pageId="8" pageNumber="72">Betaburmesebuthus bellus</emphasis>
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Lourenço, 2016
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[810,926,1382,1408]" pageId="8" pageNumber="72" type="description">
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<paragraph blockId="8.[810,1307,1346,1408]" box="[810,926,1382,1408]" pageId="8" pageNumber="72">
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(
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<figureCitation box="[818,917,1382,1408]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="1.[136,235,1411,1435]" captionStartId-1="2.[136,235,1854,1878]" captionStartId-2="3.[136,235,1712,1736]" captionStartId-3="4.[136,235,1982,2006]" captionStartId-4="5.[136,235,1934,1958]" captionStartId-5="6.[136,235,1657,1681]" captionStartId-6="7.[136,235,1951,1975]" captionTargetBox-0="[151,1436,189,1387]" captionTargetBox-1="[164,1423,189,1830]" captionTargetBox-2="[151,1436,189,1689]" captionTargetBox-3="[138,1449,189,1959]" captionTargetBox-4="[174,1413,189,1910]" captionTargetBox-5="[151,1436,189,1633]" captionTargetBox-6="[168,1420,189,1928]" captionTargetId-0="figure-209@1.[151,1436,189,1387]" captionTargetId-1="figure-18@2.[164,1423,189,1830]" captionTargetId-2="figure-114@3.[151,1436,189,1689]" captionTargetId-3="figure-18@4.[138,1449,189,1959]" captionTargetId-4="figure-28@5.[174,1413,189,1910]" captionTargetId-5="figure-113@6.[151,1436,189,1633]" captionTargetId-6="figure-28@7.[168,1420,189,1928]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="7" captionText-0="FIGURE 1. General habitus of Betaburmesebuthus bellus, NIGP200648, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 1 mm." captionText-1="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." captionText-2="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." captionText-3="FIGURE 4. General habitus of Betaburmesebuthus bellus, NIGP200649, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 1 mm." captionText-4="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." captionText-5="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." captionText-6="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi-0="http://doi.org/10.5281/zenodo.7754822" figureDoi-1="http://doi.org/10.5281/zenodo.7754826" figureDoi-2="http://doi.org/10.5281/zenodo.7754828" figureDoi-3="http://doi.org/10.5281/zenodo.7754830" figureDoi-4="http://doi.org/10.5281/zenodo.7754834" figureDoi-5="http://doi.org/10.5281/zenodo.7754836" figureDoi-6="http://doi.org/10.5281/zenodo.7754838" httpUri-0="https://zenodo.org/record/7754822/files/figure.png" httpUri-1="https://zenodo.org/record/7754826/files/figure.png" httpUri-2="https://zenodo.org/record/7754828/files/figure.png" httpUri-3="https://zenodo.org/record/7754830/files/figure.png" httpUri-4="https://zenodo.org/record/7754834/files/figure.png" httpUri-5="https://zenodo.org/record/7754836/files/figure.png" httpUri-6="https://zenodo.org/record/7754838/files/figure.png" pageId="8" pageNumber="72">Figs 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="72" type="materials_examined">
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<paragraph blockId="8.[810,1452,1454,2093]" pageId="8" pageNumber="72">
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<emphasis bold="true" box="[810,921,1454,1480]" pageId="8" pageNumber="72">Material.</emphasis>
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<collectionCode box="[929,996,1454,1480]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="8" pageNumber="72">NIGP</collectionCode>
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200648, one probable juvenile male, the part of right pedipalp finger and metasoma damaged.
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<collectionCode box="[810,877,1526,1552]" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="8" pageNumber="72">NIGP</collectionCode>
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200649, one probable juvenile female, a complete scorpion.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="9" lastPageNumber="73" pageId="8" pageNumber="72" type="diagnosis">
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<paragraph blockId="8.[810,1452,1454,2093]" lastBlockId="9.[136,778,158,2093]" lastPageId="9" lastPageNumber="73" pageId="8" pageNumber="72">
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<emphasis bold="true" box="[863,1122,1598,1624]" pageId="8" pageNumber="72">Diagnosis (emended).</emphasis>
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This species can be clearly distinguished from other congeners by the following set of characters: 1) anterior margin of carapace with a single moderately marked median concavity (
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<figureCitation box="[1257,1353,1706,1732]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2A</figureCitation>
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,
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<figureCitation box="[1367,1405,1706,1732]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6A</figureCitation>
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); 2) spiracles small, oblique and oval to slit-like (
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<figureCitation box="[1309,1398,1742,1768]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2E</figureCitation>
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,
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<figureCitation box="[1409,1447,1742,1768]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6G</figureCitation>
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; Lourenço, 2016: fig. 7); 3) pectines with 17–18 teeth (
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<figureCitation captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="8" pageNumber="72">Figs 2K</figureCitation>
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,
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<figureCitation box="[861,896,1814,1841]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="8" pageNumber="72">5B</figureCitation>
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; Lourenço, 2016: fig. 7); 4) vesicle pear-shaped and very long, and aculeus very long and moderately curved (
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<figureCitation box="[905,999,1886,1913]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="8" pageNumber="72">Figs 5E</figureCitation>
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,
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<figureCitation box="[1015,1041,1886,1912]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="8" pageNumber="72">6I</figureCitation>
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; Lourenço, 2016: fig. 4); 5) dorsal patellar spur carina well-developed with a strong tubercle and three relatively small spinous tubercles (
|
||
<figureCitation box="[1300,1447,1958,1985]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="8" pageNumber="72">Figs 3D, E, F</figureCitation>
|
||
,
|
||
<figureCitation box="[810,879,1994,2020]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="8" pageNumber="72">7C, D</figureCitation>
|
||
); 6) Chela manus with two small spinous tubercles on internal surface (
|
||
<figureCitation box="[1045,1237,2030,2057]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="8" pageNumber="72">Figs 3G, I, J, K</figureCitation>
|
||
,
|
||
<figureCitation box="[1253,1321,2030,2057]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="8" pageNumber="72">7F, G</figureCitation>
|
||
; Lourenço, 2016: figs 5, 9); 7) telotarsus with few ventrosubmedian setae and a ventromedian spinules row (
|
||
<figureCitation box="[592,727,158,185]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2M, N</figureCitation>
|
||
,
|
||
<figureCitation captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6L, M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="73" type="description">
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
<emphasis bold="true" box="[189,335,230,256]" pageId="9" pageNumber="73">Description.</emphasis>
|
||
Carapace. Sparsely covered by fine granules without distinct carinae and furrows (
|
||
<figureCitation box="[676,773,266,292]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[136,174,302,328]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6A</figureCitation>
|
||
); median eyes oval and medium sized, and separated by more than one ocular diameter (
|
||
<figureCitation box="[540,635,338,364]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[649,687,338,364]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6A</figureCitation>
|
||
); PDMi posterodorsal to PLMa, and ADMi dorsal to PLMa (
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation box="[183,219,410,436]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
Coxosternal region.Surface smooth with several setae (
|
||
<figureCitation box="[144,235,482,508]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation box="[247,281,482,508]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6E</figureCitation>
|
||
); anterior margin of coxapophysis I rounded densely covered by fine setae (
|
||
<figureCitation box="[471,561,518,544]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation box="[571,605,518,544]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6E</figureCitation>
|
||
); lateral margin of sternum very long, and posterior margin of sternum slightly incurved and as long as genital operculum (
|
||
<figureCitation captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation box="[182,216,626,652]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6E</figureCitation>
|
||
), posterior depression region small and not obvious (
|
||
<figureCitation box="[144,235,662,688]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">Fig. 6E</figureCitation>
|
||
); genital operculum longitudinally divided and composed of two nearly rounded valves (
|
||
<figureCitation box="[599,684,698,724]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">Fig. 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
Chelicerae.With setae on internal and ventral surface; cheliceral dentition partly visible (
|
||
<figureCitation box="[526,650,770,796]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">Fig. 6C, D</figureCitation>
|
||
), including a long dorsal distal (dd) denticle, a small subdistal (sd) denticle, a stout median (m) denticle, a long ventral distal (vd) denticle and two small ventral accessory (va) denticle in movable finger, and distal (d) and subdistal (sd) denticles in fixed finger.
|
||
</paragraph>
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
Pedipalps. Femur with five carinae (
|
||
<figureCitation box="[598,724,986,1012]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="9" pageNumber="73">Figs 3A–C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">7A, B</figureCitation>
|
||
): internomedian carina well-developed with about five small tubercles; dorsointernal carina costate with several granules; dorsoexternal carina feebly marked with few granules; ventroexternal carina costate with few small tubercles; ventrointernal carina costate and incomplete. Patella with seven carinae (
|
||
<figureCitation box="[461,586,1202,1228]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="9" pageNumber="73">Figs 3D–F</figureCitation>
|
||
,
|
||
<figureCitation box="[603,678,1202,1228]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">7C, D</figureCitation>
|
||
): Dorsal Patellar Spur carina(DPSc)andVentral Patellar Spur carina (VPSc) well-developed and each apophysis or tubercle possessing one microseta; dorsointernal, dorsomedian and dorsoexternal carinae smooth and costate; ventroexternal and ventrointernal carinae incomplete and costate. Carinae on chela not evident; Chela relatively slender (Cl/Cw = 6.19 or 6.23,
|
||
<tableCitation box="[283,364,1454,1480]" captionStart="TABLE 1" captionStartId="10.[136,224,496,522]" captionText="TABLE 1. Measurements (mm) of studied specimens and corresponding holotype. Question marks signify doubtful data; value in brackets signify re-estimation from the images in original papers." pageId="9" pageNumber="73">Table 1</tableCitation>
|
||
); each finger denticle row consisting of about 10–11 (first row with 5) granules and a thick and short seta present beneath each accessory granule.
|
||
</paragraph>
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
Trichobothrial pattern (
|
||
<figureCitation box="[461,538,1562,1588]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="9" pageNumber="73">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation box="[557,573,1562,1588]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">7</figureCitation>
|
||
). Pedipalp femur with 10 trichobothria, 5 dorsal, 3 internal and 2 external trichobothria, trichobothrium
|
||
<emphasis box="[476,498,1634,1667]" italics="true" pageId="9" pageNumber="73">d 2</emphasis>
|
||
straddling dorsointernal carina (
|
||
<figureCitation box="[225,355,1670,1697]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">Fig. 7A, B</figureCitation>
|
||
) and trichobothrium
|
||
<emphasis box="[610,630,1671,1703]" italics="true" pageId="9" pageNumber="73">
|
||
e
|
||
<subScript attach="left" box="[622,630,1689,1703]" fontSize="6" pageId="9" pageNumber="73">1</subScript>
|
||
</emphasis>
|
||
proximal to trichobothrium
|
||
<emphasis box="[315,337,1706,1739]" italics="true" pageId="9" pageNumber="73">
|
||
d
|
||
<subScript attach="left" box="[329,337,1725,1739]" fontSize="6" pageId="9" pageNumber="73">5</subScript>
|
||
</emphasis>
|
||
(
|
||
<figureCitation box="[356,489,1706,1733]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="9" pageNumber="73">Figs 3A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[504,577,1706,1733]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">7A, B</figureCitation>
|
||
). Patella with 13 trichobothria, including 5 dorsal (
|
||
<emphasis box="[541,563,1742,1775]" italics="true" pageId="9" pageNumber="73">
|
||
d
|
||
<subScript attach="left" box="[555,563,1761,1775]" fontSize="6" pageId="9" pageNumber="73">2</subScript>
|
||
</emphasis>
|
||
petite), 1 internal and 7 external trichobothria; trichobothrium
|
||
<emphasis box="[659,681,1778,1811]" italics="true" pageId="9" pageNumber="73">
|
||
d
|
||
<subScript attach="left" box="[673,681,1797,1811]" fontSize="6" pageId="9" pageNumber="73">3</subScript>
|
||
</emphasis>
|
||
internal to dorsomedian carina and trichobothrium
|
||
<emphasis box="[625,647,1814,1847]" italics="true" pageId="9" pageNumber="73">
|
||
d
|
||
<subScript attach="left" box="[639,647,1833,1847]" fontSize="6" pageId="9" pageNumber="73">4</subScript>
|
||
</emphasis>
|
||
external to dorsomedian carina (
|
||
<figureCitation box="[368,489,1850,1877]" captionStart="FIGURE 3" captionStartId="3.[136,235,1712,1736]" captionTargetBox="[151,1436,189,1689]" captionTargetId="figure-114@3.[151,1436,189,1689]" captionTargetPageId="3" captionText="FIGURE 3. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy (B, C, E, F, H–K) and green epifluorescence (A, D, G). A, B, Left femur, dorsal view. C, Right femur, dorsal view. D, E, left patella, dorsal view. F, Right patella, dorsal view. G, Left chela, dorsal view. H, Left fixed finger, dorsal view. I, Right manus, ventral view. J, Left manus, dorsal view. K, Left manus, ventral view. Scale bars = 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7754828" httpUri="https://zenodo.org/record/7754828/files/figure.png" pageId="9" pageNumber="73">Figs 3D, E</figureCitation>
|
||
,
|
||
<figureCitation box="[499,568,1850,1876]" captionStart="FIGURE 7" captionStartId="7.[136,235,1951,1975]" captionTargetBox="[168,1420,189,1928]" captionTargetId="figure-28@7.[168,1420,189,1928]" captionTargetPageId="7" captionText="FIGURE 7. Trichobothrial pattern of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy (B, D, F, G) and green epifluorescence (A, C, E). A–B, Right femur, dorsal view. C–D, Right patella, dorsal view. E–F, Right chela, dorsal view. G, Right manus, ventral view. Scale bars = 200 μm in A, B, C, D, G; 400 μm in E, F." figureDoi="http://doi.org/10.5281/zenodo.7754838" httpUri="https://zenodo.org/record/7754838/files/figure.png" pageId="9" pageNumber="73">7C, D</figureCitation>
|
||
). Fixed finger with 1 internal, 3 external, and 2 dorsal trichobothria. Chela manus with 6 external and 2 ventral trichobothria.
|
||
</paragraph>
|
||
<paragraph blockId="9.[136,778,158,2093]" pageId="9" pageNumber="73">
|
||
Legs. Trochanter bearing a lateral apophysis (
|
||
<figureCitation captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">Fig. 5B</figureCitation>
|
||
). Prolateral and retrolateral pedal spurs present on all legs (
|
||
<figureCitation box="[195,312,2030,2056]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Fig. 2L–O</figureCitation>
|
||
). Ungues moderately long, dactyl pointed (
|
||
<figureCitation box="[144,266,2066,2092]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2L–O</figureCitation>
|
||
,
|
||
<figureCitation box="[280,339,2066,2093]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">5H, I</figureCitation>
|
||
,
|
||
<figureCitation box="[351,426,2066,2092]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6L, M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[810,1452,158,1517]" pageId="9" pageNumber="73">
|
||
Pectines (
|
||
<figureCitation box="[968,1059,158,184]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2K</figureCitation>
|
||
,
|
||
<figureCitation box="[1068,1104,158,185]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">5B</figureCitation>
|
||
). Basal piece inverted trapezoid with an anterior median furrow (
|
||
<figureCitation box="[1188,1276,194,220]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">Fig. 6F</figureCitation>
|
||
). Pectines with 3 marginal lamellae and 8–9 median lamellae; fulcra present, very small. Short and transparent sensory hairs extending from surface of lamellae. Teeth elongated and terminal one ovoid, peg sensillae very short and sensory area occupied about three quarters of teeth.
|
||
</paragraph>
|
||
<paragraph blockId="9.[810,1452,158,1517]" pageId="9" pageNumber="73">
|
||
Mesosoma. Tergites finely granular, median carina evident on posterior half of tergites III–VI (
|
||
<figureCitation box="[1301,1388,446,472]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Figs 2F</figureCitation>
|
||
,
|
||
<figureCitation box="[1400,1438,446,473]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">5A</figureCitation>
|
||
); dorsolateral carinae observed on tergites IV–VI (
|
||
<figureCitation box="[1353,1438,482,508]" captionStart="FIGURE 2" captionStartId="2.[136,235,1854,1878]" captionTargetBox="[164,1423,189,1830]" captionTargetId="figure-18@2.[164,1423,189,1830]" captionTargetPageId="2" captionText="FIGURE 2. Details of Betaburmesebuthus bellus, NIGP200648, under confocal microscopy with depth color-coding. A, Carapace. B, Lateral ocelli. C, Coxapophysis, coxa and trochanter of leg I. D, Sternum and genital operculum. E, Spiracle of sternite VI. F, Tergites I–VI. G, Dentition of movable finger. H, Dentition of fixed finger. I, Tibial spur of leg III. J, Tibial spur of leg IV. K, Left pectine. L–O, Telotarsi of legs I–IV.Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; ts, tibial spur. Scale bars = 200 μm in A, F, G, K; 100 μm in C, D, E, H, I, J, L, M, N, O; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754826" httpUri="https://zenodo.org/record/7754826/files/figure.png" pageId="9" pageNumber="73">Fig. 2F</figureCitation>
|
||
); tergite VII with five costate carinae (
|
||
<figureCitation box="[1258,1355,518,545]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">Fig. 5A</figureCitation>
|
||
): paired dorsolateral and lateral carinae, and one axial carina on anterior half. Sternites covered by some fine granules and sparse setae; posterior edge of sternites I–IV incurved (
|
||
<figureCitation box="[818,906,662,689]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="9.[810,1452,158,1517]" pageId="9" pageNumber="73">
|
||
Metasoma (
|
||
<figureCitation box="[1009,1164,698,725]" captionStart="FIGURE 1" captionStartId="1.[136,235,1411,1435]" captionTargetBox="[151,1436,189,1387]" captionTargetId="figure-209@1.[151,1436,189,1387]" captionTargetPageId="1" captionText="FIGURE 1. General habitus of Betaburmesebuthus bellus, NIGP200648, under incident light. A, Dorsal view. B, Ventral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7754822" httpUri="https://zenodo.org/record/7754822/files/figure.png" pageId="9" pageNumber="73">Figs 1A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[1190,1258,698,725]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">5C–E</figureCitation>
|
||
). All segments longer than width and covered by macrosetae ventrally. Segments I–IV with week depression dorsally; segments I and II with ten well marked carinae (paired ventromedian, ventrolateral, lateral, dorsolateral, and dorsal carinae); segments III and IV with eight well marked carinae (paired ventromedian, ventrolateral, dorsolateral, and dorsal carinae) and lateral carinae reduced on segment III; segment V with five carinae (single ventromedian carina, paired ventrolateral and dorsolateral carinae); dorsal carinae on segments II–IV distinctly serrated, other carinae smooth to serrated.
|
||
</paragraph>
|
||
<paragraph blockId="9.[810,1452,158,1517]" pageId="9" pageNumber="73">
|
||
Telson (
|
||
<figureCitation box="[953,1044,1130,1157]" captionStart="FIGURE 5" captionStartId="5.[136,235,1934,1958]" captionTargetBox="[174,1413,189,1910]" captionTargetId="figure-28@5.[174,1413,189,1910]" captionTargetPageId="5" captionText="FIGURE 5. Details of Betaburmesebuthus bellus, NIGP200649, under green epifluorescence. A, Carapace and tergites. B, Coxosternal region, pectines and sternites. C, Metasomal segments I–IV, ventral view. D, Metasomal segments I–IV, dorsal view. E, Metasomal segment V and telson, lateral view. F, Tibial spur of leg III. G, Tibial spur of leg IV. H, Telotarsus of leg III. I, Telotarsus of leg IV. Abbreviations: ts, tibial spur. Scale bars = 400 μm in A–E; 100 μm in F–I." figureDoi="http://doi.org/10.5281/zenodo.7754834" httpUri="https://zenodo.org/record/7754834/files/figure.png" pageId="9" pageNumber="73">Figs 5E</figureCitation>
|
||
,
|
||
<figureCitation box="[1056,1083,1130,1156]" captionStart="FIGURE 6" captionStartId="6.[136,235,1657,1681]" captionTargetBox="[151,1436,189,1633]" captionTargetId="figure-113@6.[151,1436,189,1633]" captionTargetPageId="6" captionText="FIGURE 6. Details of Betaburmesebuthus bellus, NIGP200649, under confocal microscopy. A, Carapace. B, Lateral ocelli. C, Chelicerae, dorsal view. D, Chelicerae, ventral view. E, Coxosternal region. F, Genital operculum and basal piece. G, Spiracles. H, Dentition of movable finger. I, Telson. J, Tibial spur of leg III. K, Tibial spur of leg IV. L, Telotarsi of leg IV. M, Telotarsi of leg III. Abbreviations:ADMi, anterodorsal minor ocellus; ALMa, anterolateral major ocellus; MLMa, mediolateral major ocellus; PDMi, posterodorsal minor ocellus; PLMa, posterolateral major ocellus; sp, spiracle; st, subaculear tubercle; ts, tibial spur. Scale bars = 200 μm in A, E, G, H; 100 μm in C, D, F, I, J, K, L, M; 50 μm in B." figureDoi="http://doi.org/10.5281/zenodo.7754836" httpUri="https://zenodo.org/record/7754836/files/figure.png" pageId="9" pageNumber="73">6I</figureCitation>
|
||
). Vesicle with ventromedian and ventrosubmedian carinae evident and covered by several macrosetae ventrally, a small subaculear tuberance present on terminal of ventromedian carina and subaculear setal pair significantly evident; vesicle/aculeus juncture moderately wide and shorter than half of vesicle; aculeus with several microsetae proximally and progressively darker towards distal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="9" pageNumber="73" type="discussion">
|
||
<paragraph blockId="9.[810,1452,158,1517]" pageId="9" pageNumber="73">
|
||
<emphasis bold="true" box="[863,979,1418,1444]" pageId="9" pageNumber="73">Remarks.</emphasis>
|
||
We add some additional characters to diagnosis,
|
||
<emphasis box="[929,937,1455,1480]" italics="true" pageId="9" pageNumber="73">i</emphasis>
|
||
.
|
||
<emphasis box="[943,955,1455,1480]" italics="true" pageId="9" pageNumber="73">e</emphasis>
|
||
., the DPS of patella and telotarsus armature, which were not mentioned in the original descriptions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |