305 lines
29 KiB
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305 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1002.58017" ID-GBIF-Dataset="8bedb86f-0c38-4738-b1b5-1dab270c95d2" ID-PMC="PMC7746671" ID-Pensoft-Pub="1313-2970-1002-1" ID-Pensoft-UUID="92722606C9CB50B6A63A1345E68167F9" ID-PubMed="33363429" ID-ZooBank="C9EAF9DCD87D402FAC35CC4983D2E142" ModsDocID="1313-2970-1002-1" checkinTime="1607628106955" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana M. & Gattolliat, Jean-Luc" docDate="2020" docId="C35E46679F8F59C5AA3BD0F58FEF272F" docLanguage="en" docName="ZooKeys 1002: 1-114" docOrigin="ZooKeys 1002" docPubDate="2020-12-10" docSource="http://dx.doi.org/10.3897/zookeys.1002.58017" docTitle="Labiobaetis lachicae Kaltenbach & Garces & Gattolliat 2020, sp. nov." docType="treatment" docUuid="60DC83D6-32F6-4181-B224-1A890EF26245" docUuidSource="ZooBank" docVersion="4" id="92722606C9CB50B6A63A1345E68167F9" lastPageNumber="1" masterDocId="92722606C9CB50B6A63A1345E68167F9" masterDocTitle="The success story of Labiobaetis Novikova & Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species" masterLastPageNumber="114" masterPageNumber="1" pageNumber="1" updateTime="1668146493680" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The success story of Labiobaetis Novikova & Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kaltenbach, Thomas</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Garces, Jhoana M.</mods:namePart>
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<mods:affiliation>Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
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<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland & University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-12-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1002</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>114</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1002.58017</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1002.58017</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1002-1</mods:identifier>
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<mods:identifier type="ZooBank">C9EAF9DCD87D402FAC35CC4983D2E142</mods:identifier>
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<mods:identifier type="Pensoft-UUID">92722606C9CB50B6A63A1345E68167F9</mods:identifier>
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<treatment ID-GBIF-Taxon="170725670" LSID="urn:lsid:zoobank.org:act:60DC83D6-32F6-4181-B224-1A890EF26245" httpUri="http://treatment.plazi.org/id/C35E46679F8F59C5AA3BD0F58FEF272F" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/60DC83D6-32F6-4181-B224-1A890EF26245" authority="Kaltenbach & Garces & Gattolliat, 2020" authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis lachicae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lachicae" status="sp. nov.">Labiobaetis lachicae</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">Figures 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">, 12</figureCitation>
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Habitus, larvae, dorsal view a Labiobaetis camiguinensis sp. nov. b Labiobaetis lachicae sp. nov. c Labiobaetis palawano sp. nov. d Labiobaetis sabordoi sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure42" httpUri="https://binary.pensoft.net/fig/484707" pageId="0" pageNumber="1">, 42b</figureCitation>
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<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, ventral view a Labiobaetis lachicae sp. nov. b Labiobaetis palawano sp. nov. c Labiobaetis sabordoi sp. nov. d Hypopharynx and superlinguae of Labiobaetis camiguinensis sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure47" httpUri="https://binary.pensoft.net/fig/484712" pageId="0" pageNumber="1">, 47a</figureCitation>
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<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Labiobaetis in the Philippines a Philippines, overview b Group Labiobaetis dendrisetis c Group Labiobaetis numeratus." figureDoi="10.3897/zookeys.1002.58017.figure48" httpUri="https://binary.pensoft.net/fig/484713" pageId="0" pageNumber="1">, 48c</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
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Following combination of characters: A) dorsal surface of labrum with submarginal arc of one plus 5-7 long, simple setae; B) labial palp segment II with a thumb-like distomedial protuberance, segment III conical; C) left mandible without setae at apex of mola; D) fore femur rather broad, length ca. 3
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<normalizedToken originalValue="×">x</normalizedToken>
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maximum width, dorsal margin with 10-12 curved, spine-like setae and a partial second row near margin; E) claw with 15-17 denticles; F) paraproct distally not expanded, with ca. 13 stout, marginal spines.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
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(Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12</figureCitation>
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,
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Habitus, larvae, dorsal view a Labiobaetis camiguinensis sp. nov. b Labiobaetis lachicae sp. nov. c Labiobaetis palawano sp. nov. d Labiobaetis sabordoi sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure42" httpUri="https://binary.pensoft.net/fig/484707" pageId="0" pageNumber="1">42b</figureCitation>
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,
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<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, ventral view a Labiobaetis lachicae sp. nov. b Labiobaetis palawano sp. nov. c Labiobaetis sabordoi sp. nov. d Hypopharynx and superlinguae of Labiobaetis camiguinensis sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure47" httpUri="https://binary.pensoft.net/fig/484712" pageId="0" pageNumber="1">47a</figureCitation>
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). Body length 3.6-3.7 mm. Cerci ca. 2/3 of body length. Paracercus ca. 2/3 of cerci length. Antenna approx. twice as long as head length.
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</paragraph>
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<caption doi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
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<taxonomicName authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis lachicae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lachicae">
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<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis lachicae</emphasis>
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</taxonomicName>
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sp. nov., larva morphology
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<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
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labrum
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<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
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right mandible
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<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
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right prostheca
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<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
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left mandible
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<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
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left prostheca
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<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
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hypopharynx and superlinguae
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<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
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maxilla
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<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
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labium
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<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
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apex of paraglossa.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
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.
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</emphasis>
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Head dorsally light brown, thorax and abdomen dorsally brown, with bright pattern as in Fig.
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Habitus, larvae, dorsal view a Labiobaetis camiguinensis sp. nov. b Labiobaetis lachicae sp. nov. c Labiobaetis palawano sp. nov. d Labiobaetis sabordoi sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure42" httpUri="https://binary.pensoft.net/fig/484707" pageId="0" pageNumber="1">42b</figureCitation>
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. Fore protoptera light brown with distinct brown striation. Head, thorax, and abdomen ventrally brown with bright pattern as in Fig.
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<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Habitus, larvae, ventral view a Labiobaetis lachicae sp. nov. b Labiobaetis palawano sp. nov. c Labiobaetis sabordoi sp. nov. d Hypopharynx and superlinguae of Labiobaetis camiguinensis sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure47" httpUri="https://binary.pensoft.net/fig/484712" pageId="0" pageNumber="1">47a</figureCitation>
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. Legs ecru, femur with two partly merged distomedial brown spots, apex with brown spot, tibia with distomedial brown area. Caudalii light brown, with a brown band at ca. 1/3 of cerci, cerci brown in distal part.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12f</figureCitation>
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) with scape and pedicel subcylindrical, without distolateral process at scape.
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</paragraph>
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<caption doi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
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<taxonomicName authorityName="Kaltenbach & Garces & Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis lachicae" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lachicae">
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<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis lachicae</emphasis>
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</taxonomicName>
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sp. nov., larva morphology
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<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
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foreleg
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<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
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fore claw
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<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
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tergum IV
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<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
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gill IV
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<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
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paraproct
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<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
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antennal scape
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<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
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metanotum.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11a</figureCitation>
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). Rectangular, length 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of one plus 5-7 long, simple setae, the first two setae after the submedian seta are close together. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid setae; ventral surface with ca. six short, spine-like setae near lateral and anterolateral margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11b, c</figureCitation>
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). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with three denticles, inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola with a pronounced hump. Tuft of setae at apex of mola present and many thin setae distally at base of mola.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11d, e</figureCitation>
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). Incisor and kinetodontium fused. Incisor with three denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola convex. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hypopharynx and superlinguae</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11f</figureCitation>
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). Lingua shorter than superlinguae. Lingua longer than broad; medial tuft of stout setae well developed, short; distal half laterally expanded. Superlinguae distally rounded; lateral margin angulate; fine, long, simple setae along distal margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11g</figureCitation>
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). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and four medium to long, simple setae. Maxillary palp 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as length of galea-lacinia; 2-segmented; palp segment II 1.6
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<normalizedToken originalValue="×">x</normalizedToken>
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length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment slightly pointed, without excavation at inner distolateral margin.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
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(Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Labiobaetis lachicae sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure11" httpUri="https://binary.pensoft.net/fig/484676" pageId="0" pageNumber="1">11h, i</figureCitation>
|
||
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. nine short, stout, spine-like setae plus distalmost one much longer, less robust, spine-like seta; apex with two long and one medium, robust, pectinate setae; outer margin with four spine-like setae increasing in length distally; ventral surface with fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and four medium, simple setae in anteromedial area; outer margin with some long, spine-like setae; dorsally with a row of five long, spine-like, simple setae near inner margin. Labial palp with segment I 0.9
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of segments II and III combined. Segment I ventrally with short, fine, simple setae and one short, stout, simple seta at inner margin. Segment II with thumb-like distomedial protuberance; distomedial protuberance 0.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with a row of 6-8 medium, spine-like, simple setae near outer margin. Segment III conical; apex rounded; length 1.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind protoptera</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12g</figureCitation>
|
||
) minute.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12a, b</figureCitation>
|
||
). Ratio of foreleg segments 1.4:1.0:0.8:0.3.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
|
||
.
|
||
</emphasis>
|
||
Length ca. 3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width. Dorsal margin with 10-12 curved, spine-like setae, proximally and medially a partial second row of spine-like setae and some additional stout setae near margin; length of setae 0.23
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
maximum width of femur. Apex rounded, with a pair of spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch absent.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal margin with a row of short to medium, spine-like setae, on apex one longer, spine-like seta. Ventral margin with a row of medium, curved, spine-like setae, distally of patellotibial suture one long, curved, spine-like seta, on apex some longer, partly bipectinate, spine-like setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal half.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal margin with some short, stout setae. Ventral margin with a row of curved, spine-like setae and some spine-like setae near margin. Claw with one row of 15-17 denticles; distally pointed; with ca. six stripes; subapical setae absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Terga</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12c</figureCitation>
|
||
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae and micropores. Posterior margin of tergum IV with rounded, partly fused spines, wider than long.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gills</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12d</figureCitation>
|
||
). Present on segments II-VII. Margin with small denticles intercalating fine simple setae. Tracheae partly extending from main trunk towards outer and inner margins. Gill IV as long as length of segments V and VI combined. Gill VII as long as length of segments VIII and IX combined.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Labiobaetis lachicae sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure12" httpUri="https://binary.pensoft.net/fig/484677" pageId="0" pageNumber="1">12e</figureCitation>
|
||
). Distally not expanded, with ca. 13 stout, marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with small, marginal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Dedicated to Ms. Maria Kenosis Lachica (Philippines/Japan), friend of the AdMU Biodiversity Laboratory.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Philippines: Mindanao (Fig.
|
||
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Labiobaetis in the Philippines a Philippines, overview b Group Labiobaetis dendrisetis c Group Labiobaetis numeratus." figureDoi="10.3897/zookeys.1002.58017.figure48" httpUri="https://binary.pensoft.net/fig/484713" pageId="0" pageNumber="1">48c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
|
||
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The specimens were collected at altitudes from sea level to 120 m, in root packs, grass bunches or submerged wood in riffles or runs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
.
|
||
</emphasis>
|
||
Philippines • larva; Mindanao, Agusan del Sur, San Francisco, Bayogan, Tagkunayan Creek;
|
||
<geoCoordinate degrees="08" direction="north" minutes="28" orientation="latitude" precision="925" value="8.466666">08°28'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="125" direction="east" minutes="59" orientation="longitude" precision="925" value="125.98333">125°59'E</geoCoordinate>
|
||
; 120 m; 05.II.1998; leg. Mendoza; on slide; GenBank: MT830950; GBIFCH 00654891; PNM.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||
.
|
||
</emphasis>
|
||
Philippines • 2 larvae; same data as holotype; 1 on slide; GBIFCH 00592269; AdMU; 1 on slide; GBIFCH 00592310; ZSM.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Other material.</emphasis>
|
||
Philippines • 1 larva; Mindanao, Surigao del Sur, Tandag, middle Tandag River; 30 m;
|
||
<geoCoordinate degrees="09" direction="north" minutes="03" orientation="latitude" precision="15" seconds="33" value="9.059167">09°03'33"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="126" direction="east" minutes="05" orientation="longitude" precision="15" seconds="57" value="126.09917">126°05'57"E</geoCoordinate>
|
||
; 04.XII.2018; leg. Pangantihon; in alcohol; GBIFCH 00515332; AdMU • 1 larva; Mindanao, Surigao del Sur, Tandag, Pangi, Pangi River;
|
||
<geoCoordinate degrees="09" direction="north" minutes="06" orientation="latitude" precision="15" seconds="18" value="9.1050005">09°06'18"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="126" direction="east" minutes="08" orientation="longitude" precision="15" seconds="53" value="126.148056">126°08'53"E</geoCoordinate>
|
||
; 10 m; 30.XI.2018, leg. Pangantihon; on slide; GBIFCH 515520; ZSM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |