treatments-xml/data/9E/58/A9/9E58A9A6BA8154D594D9F3D7F405ECAB.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1164.103184" ID-Pensoft-Pub="1313-2970-1164-23" ID-Pensoft-UUID="62BD8FF233655E5288FA80F56FEA3299" ID-ZooBank="CC06E1473B074F478AFB08520503A404" ModsDocID="1313-2970-1164-23" checkinTime="1685200798416" checkinUser="pensoft" docAuthor="Shepard, William D. &amp; Barr, Cheryl B." docDate="2023" docId="9E58A9A6BA8154D594D9F3D7F405ECAB" docLanguage="en" docName="ZooKeys 1164: 23-61" docOrigin="ZooKeys 1164" docPubDate="2023-05-26" docSource="http://dx.doi.org/10.3897/zookeys.1164.103184" docTitle="Tychepsephus Waterhouse 1876" docType="treatment" docVersion="1" id="62BD8FF233655E5288FA80F56FEA3299" lastPageNumber="23" masterDocId="62BD8FF233655E5288FA80F56FEA3299" masterDocTitle="A revision of the Chilean water penny genus Tychepsephus Waterhouse, 1876 (Coleoptera, Psephenidae, Eubriinae), with description of a second species and two larval morphotypes, and notes on other Chilean Psephenidae" masterLastPageNumber="61" masterPageNumber="23" pageNumber="23" updateTime="1685200798416" updateUser="pensoft">
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<mods:title id="476923D1BC157CED2F309FE84F91E964">A revision of the Chilean water penny genus Tychepsephus Waterhouse, 1876 (Coleoptera, Psephenidae, Eubriinae), with description of a second species and two larval morphotypes, and notes on other Chilean Psephenidae</mods:title>
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<mods:namePart id="1DDF8C353EE55F32CC5107B17E4EDAF0">Shepard, William D.</mods:namePart>
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<mods:namePart id="FA1D9DEE413F6A01BE7ABB68EB7ED5D0">Barr, Cheryl B.</mods:namePart>
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<mods:affiliation id="F9748AA4042E1F5ED1F6CA8150AFB366">Essig Museum of Entomology, University of California, 1101 Valley Life Sciences Bldg., Berkeley, CA 94720, USA</mods:affiliation>
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<treatment id="9E58A9A6BA8154D594D9F3D7F405ECAB" LSID="urn:lsid:plazi:treatment:9E58A9A6BA8154D594D9F3D7F405ECAB" httpUri="http://treatment.plazi.org/id/9E58A9A6BA8154D594D9F3D7F405ECAB" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection id="5C5DCE7D582C70614385E70A1C26B167" pageId="0" pageNumber="23" type="nomenclature">
<paragraph id="DAC177B4193080ABD92AB0FB2FADEEB8" pageId="0" pageNumber="23">
Subfamily
<taxonomicName id="59971B72F2D04EF551CFE8D7807E0D54" LSID="9E58A9A6-BA81-54D5-94D9-F3D7F405ECAB" authority="Lacordaire, 1857" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubriinae">Eubriinae Lacordaire, 1857</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3BE3B5C1AF355614CF8A9B4D5F89CE11" pageId="0" pageNumber="23" type="type genus">
<paragraph id="EDA0DDB3FA9799F6AF8FB4A1A4FCCF91" pageId="0" pageNumber="23">Type genus.</paragraph>
<paragraph id="3AF67A3C551F10A591D7D11D67F5B528" pageId="0" pageNumber="23">
<taxonomicName id="87BA69415E91FAE3A45808809C7CBD91" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="1DFA14321E28CEDE62E5C86AAAB6B348" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
Latrielle, 1829
</paragraph>
</subSubSection>
<subSubSection id="3369BA173EBC43BCB274EB4DD49A3F02" pageId="0" pageNumber="23" type="diagnosis">
<paragraph id="9AB00D11CADAC789ABB586282278CF3C" pageId="0" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="D87FD0DA0B0684825EC54C7A5D2BF0DF" pageId="0" pageNumber="23">
The following characters, in combination distinguishing the
<taxonomicName id="C18DD93BFE5FE21146AE353B9145E45E" authorityName="Lacordaire" authorityYear="1857" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubriinae">Eubriinae</taxonomicName>
from the other four psephenid subfamilies,
<taxonomicName id="CB0E99BAE0F0067CF3C0C7C78B22DBC2" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
,
<taxonomicName id="D2A94BE149975BEAC6A89295345829BF" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubrianacinae">Eubrianacinae</taxonomicName>
,
<taxonomicName id="EABBEC9FFCB3A946CD22E52729E18AAD" authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
and
<taxonomicName id="93937EF1DE82714B42F7A2FD96A4AA48" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psephenoidinae">Psephenoidinae</taxonomicName>
, are for the most part taken from
<bibRefCitation id="47FED38720D154D92FD6FD56521DA2A0" DOI="https://doi.org/10.1111/j.1365-3113.2006.00374.x" author="Lee, C-F" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="23" pagination="502 - 538" refId="B13" refString="Lee, C-F, Sato, M, Shepard, WD, Jaech, MA, 2007. Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology 32 (3): 502 - 538, DOI: https://doi.org/10.1111/j.1365-3113.2006.00374.x" title="Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters." url="https://doi.org/10.1111/j.1365-3113.2006.00374.x" volume="32" year="2007">Lee et al. (2007</bibRefCitation>
,
<bibRefCitation id="913C24474CC3B2FE6CC766C7AA695544" DOI="https://doi.org/10.1515/9783110373929" author="Lee, C-F" editor="Beutel, RG" journalOrPublisher="2 nd edn. Walter de Gruyter, Berlin" pageId="0" pageNumber="23" pagination="615 - 628" refId="B14" refString="Lee, C-F, Jaech, MA, Beutel, RG, 2016. Psephenidae Lacordaire, 1854. In: Beutel, RG, Leschen, RAB, Eds., Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics. 2nd edn. Walter de Gruyter, Berlin: 615 - 628, DOI: https://doi.org/10.1515/9783110373929" title="Psephenidae Lacordaire, 1854." url="https://doi.org/10.1515/9783110373929" volumeTitle="Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics." year="2016">2016</bibRefCitation>
).
<emphasis id="885DC69F7CD6F89F67EE525A29575FA5" bold="true" pageId="0" pageNumber="23">Adults</emphasis>
: 1) dorsally convex body (flattened in other subfamilies except
<taxonomicName id="5A963AD29A4FEB1D4C2044C0BBC8423F" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
); 2) anterior margin of pronotum truncate or emarginate with an exposed head (
<taxonomicName id="335BC5037ACEF2CBF58551B5A488332D" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubrianacinae">Eubrianacinae</taxonomicName>
with pronotum rounded anteriorly and head entirely concealed); 3) maxillary palpus with apex not tapering (i.e., truncate, rounded or bifurcate) (tapering in
<taxonomicName id="BABB074B1AC12B12A6C299303DD9CD37" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
); 4) apex of the mesosternal process truncate or emarginate (
<taxonomicName id="41EFA9AFCEFE4B09DE7C5F6C59D322FA" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
with process acute;
<taxonomicName id="78054D6A2B872B541694CAF277A2418D" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psephenoidinae">Psephenoidinae</taxonomicName>
with process tapered); and 5) five abdominal ventrites (
<taxonomicName id="CDA1144BBC06BC8532E067210F8C4196" authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
with seven ventrites in males and six in females).
<emphasis id="99CEEEBA0EE885413D1AA4B337E40974" bold="true" pageId="0" pageNumber="23">Larvae</emphasis>
: 1) abdominal paratergites VII not lengthened to reach abdominal segment IX (reaching anterolateral angles of IX in
<taxonomicName id="7B62C989888BC585F12408308E2A2A17" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
and
<taxonomicName id="457EC48343389BA09F049436B1383565" authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
; surrounding IX in
<taxonomicName id="7BDF43765D58094FEBCD7769FF597097" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psephenoidinae">Psephenoidinae</taxonomicName>
); 2) ventral external gills absent (present in
<taxonomicName id="B3BF7BC7B089EC489D4F6AFAA6769AA7" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubrianacinae">Eubrianacinae</taxonomicName>
and
<taxonomicName id="2DB6463379354613310769F4F90B3F70" authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
); and 3) mature larvae metapneustic with a pair of spiracles near bases of abdominal paratergites VIII (
<taxonomicName id="CCF30D165010DFA15BE8D7DCC1DCC81C" authorityName="Lee, Sato, Shepard &amp; Jaech" authorityYear="2007" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Afroeubriinae">Afroeubriinae</taxonomicName>
metapneustic with spiracles at apices of paratergites VIII;
<taxonomicName id="A20821F1CCC4493B51EACDC82F12D939" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubrianacinae">Eubrianacinae</taxonomicName>
and
<taxonomicName id="F233AF1362BEF3C74E5D12E0F5B78B52" authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
amphineustic with exposed ventral gills;
<taxonomicName id="E428BA87491B605E15FD855E8E944B28" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Psephenoidinae">Psephenoidinae</taxonomicName>
apneustic).
</paragraph>
</subSubSection>
<subSubSection id="3B66EF297EBCA3356BD3F8FEAD786E73" pageId="0" pageNumber="23" type="distribution">
<paragraph id="5E3E6F2C009DE8671F9B0991532E3127" pageId="0" pageNumber="23">Geographic distribution.</paragraph>
<paragraph id="CB0920F93B83B6E0B1CBCFDB53E2CA47" pageId="0" pageNumber="23">
The
<taxonomicName id="A919E6D9279950D18FDA5FC0EFB3DAB7" authorityName="Lacordaire" authorityYear="1857" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubriinae">Eubriinae</taxonomicName>
occur almost worldwide except in Antarctica and on some islands, including New Zealand. The subfamily is represented by 15 genera and 144 species (
<bibRefCitation id="BD517217191B706FBA1E9F58DBA4D9DA" DOI="https://doi.org/10.1515/9783110373929" author="Lee, C-F" editor="Beutel, RG" journalOrPublisher="2 nd edn. Walter de Gruyter, Berlin" pageId="0" pageNumber="23" pagination="615 - 628" refId="B14" refString="Lee, C-F, Jaech, MA, Beutel, RG, 2016. Psephenidae Lacordaire, 1854. In: Beutel, RG, Leschen, RAB, Eds., Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics. 2nd edn. Walter de Gruyter, Berlin: 615 - 628, DOI: https://doi.org/10.1515/9783110373929" title="Psephenidae Lacordaire, 1854." url="https://doi.org/10.1515/9783110373929" volumeTitle="Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics." year="2016">Lee et al. 2016</bibRefCitation>
;
<bibRefCitation id="6BB4BA06B2418887AE710E35B96235B4" DOI="https://doi.org/10.1649/0010-065X-71.3.437" author="Barr, CB" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="23" pagination="437 - 444" refId="B3" refString="Barr, CB, Shepard, WD, 2017. Eubria mesoamericana Barr and Shepard (Coleoptera: Psephenidae: Eubriinae), a new psephenid species from Central America with an enigmatic generic distribution. Coleopterists Bulletin 71 (3): 437 - 444, DOI: https://doi.org/10.1649/0010-065X-71.3.437" title="Eubria mesoamericana Barr and Shepard (Coleoptera: Psephenidae: Eubriinae), a new psephenid species from Central America with an enigmatic generic distribution." url="https://doi.org/10.1649/0010-065X-71.3.437" volume="71" year="2017">Barr and Shepard 2017</bibRefCitation>
), with the greatest diversity in Asia. The genera
<taxonomicName id="77CD5E81FFDB914381CBB8327B4A8F76" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="C380B391B8B6DCC17499F547EE1F32B6" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
<normalizedToken id="5850C7E955CA7E27819AE2A6E3334851" originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1861,
<taxonomicName id="C9073E1AED952995F4BD6D35A0A4AE93" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="ACB771AE6A50A91902366562BF6D3DD7" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
,
<taxonomicName id="7A997BC0C629E0F3DA16BD808D33B09E" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="EF8A9D2999EAE7C436A039BE0C27B65A" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
, and
<taxonomicName id="46A6751CB304A48A54852B6F938A87C0" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="2BF4F4C3379915C004AEB2A668C4CB16" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
occur in the Neotropics. Of these, only
<taxonomicName id="ACF6AD08DEC1EA9CE9591ADF8D89F424" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="F579AB931EBF3A4DDC2CD29FA4ADD1B6" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
and
<taxonomicName id="E76B8E2A2418A36FE23F616DBDEAEBF5" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="2E9B1B9136157D866C2566CB53286DC4" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
are known from South America, although
<taxonomicName id="B1A4FAC87556215C560D667721D79BA3" authorityName="Guerin-Meneville" authorityYear="1861" class="Insecta" family="Psephenidae" genus="Dicranopselaphus" kingdom="Animalia" lsidName="Dicranopselaphus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="36A1D918178C0E4B1F2AE9337A7D3583" italics="true" pageId="0" pageNumber="23">Dicranopselaphus</emphasis>
</taxonomicName>
possibly occurs there as well.
</paragraph>
</subSubSection>
<subSubSection id="63187BC906F444A33B53AA4F0787B59F" pageId="0" pageNumber="23" type="biology|habitat">
<paragraph id="E3FBDDF088152ADDF5055BF12182C118" pageId="0" pageNumber="23">Habitat and biology.</paragraph>
<paragraph id="CB411B68DB43E6E78DA34D655604969A" pageId="0" pageNumber="23">
See
<bibRefCitation id="1AB7B884D472A6EEF0122A4E9E26C1E9" DOI="https://doi.org/10.1515/9783110373929" author="Lee, C-F" editor="Beutel, RG" journalOrPublisher="2 nd edn. Walter de Gruyter, Berlin" pageId="0" pageNumber="23" pagination="615 - 628" refId="B14" refString="Lee, C-F, Jaech, MA, Beutel, RG, 2016. Psephenidae Lacordaire, 1854. In: Beutel, RG, Leschen, RAB, Eds., Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics. 2nd edn. Walter de Gruyter, Berlin: 615 - 628, DOI: https://doi.org/10.1515/9783110373929" title="Psephenidae Lacordaire, 1854." url="https://doi.org/10.1515/9783110373929" volumeTitle="Handbook of Zoology - Coleoptera, Beetles (Archostemata, Adephaga, Myxophaga, Polyphaga partim): Morphology and Systematics." year="2016">Lee et al. (2016)</bibRefCitation>
for an overview of the subfamily. See
<bibRefCitation id="064DE171AEE70AB6DD6DBC5590F099EA" DOI="https://doi.org/10.11646/zootaxa.3811.4.7" author="Shepard, WD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="23" pagination="553 - 568" refId="B19" refString="Shepard, WD, Barr, CB, 2014. Neoeubria inbionis Shepard &amp; Barr, a new genus and new species of neotropical water penny beetle (Coleoptera: Psephenidae: Eubriinae), with a key to the adult Eubriinae of the Neotropical Zone. Zootaxa 3811 (4): 553 - 568, DOI: https://doi.org/10.11646/zootaxa.3811.4.7" title="Neoeubria inbionis Shepard &amp; Barr, a new genus and new species of neotropical water penny beetle (Coleoptera: Psephenidae: Eubriinae), with a key to the adult Eubriinae of the Neotropical Zone." url="https://doi.org/10.11646/zootaxa.3811.4.7" volume="3811" year="2014">Shepard and Barr (2014)</bibRefCitation>
and
<bibRefCitation id="13AAF7DA61AE3D1F63A7E4CAB7D5A7C8" DOI="https://doi.org/10.1649/0010-065X-71.3.437" author="Barr, CB" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="23" pagination="437 - 444" refId="B3" refString="Barr, CB, Shepard, WD, 2017. Eubria mesoamericana Barr and Shepard (Coleoptera: Psephenidae: Eubriinae), a new psephenid species from Central America with an enigmatic generic distribution. Coleopterists Bulletin 71 (3): 437 - 444, DOI: https://doi.org/10.1649/0010-065X-71.3.437" title="Eubria mesoamericana Barr and Shepard (Coleoptera: Psephenidae: Eubriinae), a new psephenid species from Central America with an enigmatic generic distribution." url="https://doi.org/10.1649/0010-065X-71.3.437" volume="71" year="2017">Barr and Shepard (2017)</bibRefCitation>
for habitat descriptions of two neotropical species in the genera
<taxonomicName id="FB1F534CC126CB913D7155450C6F49BF" authorityName="Shepard &amp; Barr" authorityYear="2014" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neoeubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B82A757B61EE98354107DF3714C4698" italics="true" pageId="0" pageNumber="23">Neoeubria</emphasis>
</taxonomicName>
and
<taxonomicName id="B81824817F31DDBE5F9CB9A1EE6D836B" class="Insecta" family="Psephenidae" genus="Eubria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eubria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="025F47A02A5E299C659D0614F79A4ADC" italics="true" pageId="0" pageNumber="23">Eubria</emphasis>
</taxonomicName>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="8E7A83C492D785B18880A40993BF8ABA" pageId="0" pageNumber="23" type="remarks">
<paragraph id="ACECC28107AF2ED5EF6F013266A65C77" pageId="0" pageNumber="23">Remarks.</paragraph>
<paragraph id="C7CD1068109040F77C49AB3FD12963FE" pageId="0" pageNumber="23">
In Chile, the only known psephenids are eubriines in the genus
<taxonomicName id="FE205B92B0AC3C7BCBDF3F1F9CE28823" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus" order="Coleoptera" pageId="0" pageNumber="23" rank="genus">
<emphasis id="571E1D366E05CE92BC22F092A214FB61" italics="true" pageId="0" pageNumber="23">Tychepsephus</emphasis>
</taxonomicName>
, and the species currently named
<taxonomicName id="154DE5F1EDAAE4949DAF13D243B0C40A" authorityName="Pic" authorityYear="1947" class="Insecta" family="Psephenidae" genus="Eubrianax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eubrianax luteosignatus" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="luteosignatus">
<emphasis id="BEF008A9E3DA5DF3DD62CE14CF2AB293" italics="true" pageId="0" pageNumber="23">Eubrianax luteosignatus</emphasis>
</taxonomicName>
. The type of
<taxonomicName id="51B4EF2C9DD964EF9B935A7A1729F445" lsidName="Eu. luteosignatus" pageId="0" pageNumber="23" rank="species" species="luteosignatus">
<emphasis id="3CCE0087D9FEA67AA00A320A91DF05F8" italics="true" pageId="0" pageNumber="23">Eu. luteosignatus</emphasis>
</taxonomicName>
appears to be a eubriine, so it is likely misplaced in
<taxonomicName id="9FB60145446E801CFB3556B8D69896DB" authorityName="Kiesenwetter" authorityYear="1874" class="Insecta" family="Psephenidae" genus="Eubrianax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eubrianax" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="8A1D03790BF9E1F0FED29CD69164FFC7" italics="true" pageId="0" pageNumber="23">Eubrianax</emphasis>
</taxonomicName>
which is in the subfamily
<taxonomicName id="998E86D32114703F1311B154795EBE77" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Eubrianacinae">Eubrianacinae</taxonomicName>
.
<bibRefCitation id="5797253EC3F2E0F9EBC93BC38E809A32" author="Elgueta, M" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="23" refId="B8" refString="Elgueta, M, Guerrero, M, 2005. Nuestro conocimiento de Psephenidae (Coleoptera) en Chile &quot;peniques de aqua.&quot; Oral presentation. Sociedad Chilena de Entomologia, 14 Sept. 2005." title="Nuestro conocimiento de Psephenidae (Coleoptera) en Chile &quot; peniques de aqua. &quot; Oral presentation. Sociedad Chilena de Entomologia, 14 Sept. 2005." year="2005">Elgueta and Guerrero (2005)</bibRefCitation>
stated that all known Chilean psephenids are eubriines.
</paragraph>
</subSubSection>
</treatment>
</document>