222 lines
30 KiB
XML
222 lines
30 KiB
XML
<document id="560D250B5C6FF396450D2AF03914D774" ID-CLB-Dataset="264974" ID-DOI="10.5852/ejt.2023.890.2261" ID-GBIF-Dataset="ea7ad0da-111f-4709-819b-6a84832ceec5" ID-ISSN="2118-9773" ID-Zenodo-Dep="8305633" ID-ZooBank="5BA69DED-87A2-4041-886F-E11A6D38D52D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693494219877" checkinUser="felipe" docAuthor="Calcetas, Orlando A." docDate="2023" docId="8B4587A1FFF8596B8D9E24079A2022A2" docLanguage="en" docName="ejt-890_184-203.pdf" docOrigin="European Journal of Taxonomy 890" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucopholis Dejean 1833" docType="key" docVersion="4" lastPageNumber="200" masterDocId="777CFFD9FFF7597B8D2323739F57247C" masterDocTitle="Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896" masterLastPageNumber="203" masterPageNumber="184" pageNumber="199" updateTime="1699623951618" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="FAC89AB0474FE6E50D7814A90F9451CD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="97B828AEE868F93041041441CDCF5606">
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<mods:title id="F8B0EC1C82216A8D87EEFFA413B5F92D">Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896</mods:title>
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<mods:name id="E63A6B64CF005A90F1A020EB0F080F78" type="personal">
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<mods:roleTerm id="84B2BE651EC7C5DFE3FD3F7ECE9C2222">Author</mods:roleTerm>
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<mods:namePart id="FAF657377A9C0E62561F547F8D9015BD">Calcetas, Orlando A.</mods:namePart>
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<mods:nameIdentifier id="B9BD3B608FCE9A6541C176494215E44A" type="LSID">5A1D3F2A-DB38-4170-987C-783749F2E1BD</mods:nameIdentifier>
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<mods:affiliation id="5EE9777212B707F129CF9227DC750B31">Department of Agriculture, IVA-CALABARZON, Regional Crop Protection Center (DA-RCPC) Lipa Agricultural Research and Experiment Station (LARES) Brgy. Maraouy, Lipa City, Batangas 4217, Philippines</mods:affiliation>
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<mods:nameIdentifier id="5DF7590E2072682809CF74D457913C62" type="email">orlando.calcetas@calabarzon.da.gov.ph & orly.calcetas@yahoo.com.ph</mods:nameIdentifier>
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<mods:typeOfResource id="1C5D6FD8D33956FB975874487ED92ADA">text</mods:typeOfResource>
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<mods:title id="788BA1A8558F7323D784AA3C824AF39B">European Journal of Taxonomy</mods:title>
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<mods:part id="ABB189F77845A4A4B10F60AA0BB13348">
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<mods:date id="E2C37F595C0B3914D0FCF78D00D030CA">2023</mods:date>
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<mods:number id="79EAB7CC371BC31E5A867084EECF2178">2023-08-31</mods:number>
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<mods:detail id="E469B73AAF1656D2485BCFA12D19C738" type="volume">
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<mods:number id="3D4D3E924F388033A5817B030A1DCB9F">890</mods:number>
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<mods:classification id="E5B9EE1DBE44DE0B4D2F2C730826B826">journal article</mods:classification>
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<mods:identifier id="43C4C1D1E31B164CC73BDCA023B8D6AC" type="DOI">10.5852/ejt.2023.890.2261</mods:identifier>
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<treatment id="8B4587A1FFF8596B8D9E24079A2022A2" ID-DOI="http://doi.org/10.5281/zenodo.8306352" ID-GBIF-Taxon="212742579" ID-Zenodo-Dep="8306352" LSID="urn:lsid:plazi:treatment:8B4587A1FFF8596B8D9E24079A2022A2" httpUri="http://treatment.plazi.org/id/8B4587A1FFF8596B8D9E24079A2022A2" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199">
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<subSubSection id="4BF6653CFFF859748D9E24079C3523EC" box="[189,866,1907,1936]" pageId="15" pageNumber="199" type="nomenclature">
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<paragraph id="035336B7FFF859748D9E24079C3523EC" blockId="15.[189,866,1907,1936]" box="[189,866,1907,1936]" pageId="15" pageNumber="199">
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<emphasis id="3198EAA5FFF859748D9E24079C3523EC" bold="true" box="[189,866,1907,1936]" italics="true" pageId="15" pageNumber="199">
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Key to the species of
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<taxonomicName id="C4EC4D34FFF859748CF324009D2523EC" ID-CoL="5DWL" ID-ENA="1075168" authorityName="Dejean" authorityYear="1833" box="[464,626,1907,1936]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="199" phylum="Arthropoda" rank="genus">Leucopholis</taxonomicName>
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of the
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<collectingCountry id="7BFB7627FFF859748FEE24009C3523EC" box="[717,866,1907,1936]" name="Philippines" pageId="15" pageNumber="199">Philippines</collectingCountry>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="4BF6653CFFF8596B8D9E24D99A2022A2" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199" type="key">
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<key id="777D6186FFF8596B8D9E24D99A2022A2" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199">
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<keyStep id="B8182E12FFF8596B8D9E24D99A2025AB" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199">
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<paragraph id="035336B7FFF8596B8D9E24D99A20253B" blockId="15.[189,1400,1962,2025]" lastBlockId="16.[189,1400,265,1758]" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199">
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<keyLead id="B81D9582FFF8596B8D9E24D99A20253B" lastPageId="16" lastPageNumber="200" pageId="15" pageNumber="199">
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1. Metaventral process distinctly long, extending in front of prosternal process (
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<figureCitation id="9BD72A32FFF85974895C24D99B9F23B8" box="[1151,1224,1962,1988]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="15" pageNumber="199">Fig. 2</figureCitation>
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); large to very large sized species, length 35.0–
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<quantity id="C4149B52FFF859748F4D24BD9D8B2395" box="[622,732,1998,2025]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="15" pageNumber="199" unit="mm" value="42.5">42.5 mm</quantity>
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; prosternal process mound-shaped, triangulate; pronotal scales variable in shape and size; body scales elongate ovoid, not flattened, slanted at 45° angle, base of scale only attached to surface ....................................................................................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="035336B7FFE7596B8DE222229A2025AB" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8DE222229A2025AB" pageId="16" pageNumber="200">
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– Metaventral process distinctly short not extending in front of prosternal process; moderate to small sized species, length
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<quantity id="C4149B52FFE7596B8CC222079DD825F3" box="[481,655,372,399]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.95" metricValueMin="2.25" pageId="16" pageNumber="200" unit="mm" value="26.0" valueMax="29.5" valueMin="22.5">22.5–29.5 mm</quantity>
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; prosternal process ovoid to spindle-shaped anteriorly; pronotal scales nearly uniform in shape and size; body scales distinctly ovoid, flattened, not slanted at 45° angle, entire scale strongly attached to surface ................................................................................ 5
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="B8182E12FFE7596B8D9E228D9A2026F8" pageId="16" pageNumber="200">
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<paragraph id="035336B7FFE7596B8D9E228D9A202641" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8D9E228D9A202641" pageId="16" pageNumber="200">
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2. Metaventral process lanceolate, constricted subapically (
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<figureCitation id="9BD72A32FFE7596B8EBC228D9CBC2664" box="[927,1003,510,536]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="16" pageNumber="200">Fig. 2</figureCitation>
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); abdomen covered mostly with elongate ovoid scales; length 35.0 mm .................................................................
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<taxonomicName id="C4EC4D34FFE7596B89BA21519A422640" ID-CoL="BSCT6" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1177,1301,546,572]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
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||
<emphasis id="3198EAA5FFE7596B89BA21519A422640" bold="true" box="[1177,1301,546,572]" italics="true" pageId="16" pageNumber="200">L. stainesi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2AAB57DEFFE7596B883F21509A202641" box="[1308,1399,547,573]" pageId="16" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
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||
</paragraph>
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<paragraph id="035336B7FFE7596B8DE221349A2026F8" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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||
<keyLead id="B81D9582FFE7596B8DE221349A2026F8" pageId="16" pageNumber="200">
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||
– Metaventral process not lanceolate, not constricted subapically (
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||
<figureCitation id="9BD72A32FFE7596B8EC621359B66261C" box="[997,1073,582,608]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="16" pageNumber="200">Figs 8</figureCitation>
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,
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<figureCitation id="9BD72A32FFE7596B896321359B08261D" box="[1088,1119,582,609]" captionStart="Fig" captionStartId="11.[189,232,1927,1953]" captionTargetBox="[189,1398,480,1860]" captionTargetId="figure-86@11.[416,1172,1122,1594]" captionTargetPageId="11" captionText="Fig. 15–16. Leucopholis semperi Brenske, 1896. 15. Neotype, ♂ (MNHN), metaventral process, dorsal aspect. 16. Nontypes, ♀♀, dorsal aspect (photograph of Brenske’s non-type series from ZMHB courtesy of Dr Joachim Willers)." figureDoi="http://doi.org/10.5281/zenodo.8306346" httpUri="https://zenodo.org/record/8306346/files/figure.png" pageId="16" pageNumber="200">15</figureCitation>
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,
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<figureCitation id="9BD72A32FFE7596B894E21359BDA261C" box="[1133,1165,582,608]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="16" pageNumber="200">18</figureCitation>
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); abdomen covered mostly with short ovoid scales .......................................................................................................... 3
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="B8182E12FFE7596B8D9E21DF9A20274E" pageId="16" pageNumber="200">
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||
<paragraph id="035336B7FFE7596B8D9E21DF9A202696" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8D9E21DF9A202696" pageId="16" pageNumber="200">
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3. Paramere apical process of male genitalia not distinctly elongate; bird bill-like; apical process of male genitalia tapered or rounded apically (
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<figureCitation id="9BD72A32FFE7596B8F8521A39C472697" box="[678,784,720,747]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,692,1908]" captionTargetPageId="4" captionText="Fig. 4–6. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 4. Elytron, dorsal aspect. 5. Male genitalia, dorsal aspect. 6. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306336" httpUri="https://zenodo.org/record/8306336/files/figure.png" pageId="16" pageNumber="200">Figs 5–6</figureCitation>
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,
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<figureCitation id="9BD72A32FFE7596B8E3E21A39C302696" box="[797,871,720,746]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="16" pageNumber="200">13–14</figureCitation>
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) .................................................................. 4
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</keyLead>
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</paragraph>
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<paragraph id="035336B7FFE7596B8DE221879A20274E" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8DE221879A20274E" pageId="16" pageNumber="200">
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– Paramere apical process of male genitalia distinctly elongate; not bird bill-like; apical process of male genitalia spatulate apically (
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<figureCitation id="9BD72A32FFE7596B8F1220649DED274E" box="[561,698,791,818]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="16" pageNumber="200">Figs 19–20</figureCitation>
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); length
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||
<quantity id="C4149B52FFE7596B8E0320649CDE274E" box="[800,905,791,818]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" pageId="16" pageNumber="200" unit="mm" value="42.5">42.5 mm</quantity>
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||
..................................
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<taxonomicName id="C4EC4D34FFE7596B89AD206B9A42274E" ID-CoL="BSCTG" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1166,1301,791,818]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
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<emphasis id="3198EAA5FFE7596B89AD206B9A42274E" bold="true" box="[1166,1301,791,818]" italics="true" pageId="16" pageNumber="200">L. ratcliffei</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="2AAB57DEFFE7596B883F206B9A20274E" box="[1308,1399,792,818]" pageId="16" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep id="B8182E12FFE7596B8D9E20299A2020CA" pageId="16" pageNumber="200">
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<paragraph id="035336B7FFE7596B8D9E20299A202078" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8D9E20299A202078" pageId="16" pageNumber="200">
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||
4. Paramere apical process of male genitalia tapered apically; paramere basal margin of male genitalia sinuate; paramere lateral margin of male genitalia surface rugose (
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<figureCitation id="9BD72A32FFE7596B893A200E9BCF27E4" box="[1049,1176,893,920]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="16" pageNumber="200">Figs 9–10</figureCitation>
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); medial posterior margin of pronotum slightly extended posteriorly; posterolateral margin of elytron near posterior angle distinctly infleXed laterally; pygidium with vertical flattened margin subposteriorly; length
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<quantity id="C4149B52FFE7596B8DC9209A9E042078" box="[234,339,1001,1028]" metricMagnitude="-2" metricUnit="m" metricValue="3.82" pageId="16" pageNumber="200" unit="mm" value="38.2">38.2 mm</quantity>
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||
.................................................................................................................
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<taxonomicName id="C4EC4D34FFE7596B89B8209A9A42207F" ID-CoL="BSCT9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1179,1301,1000,1027]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
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<emphasis id="3198EAA5FFE7596B89B8209A9A42207F" bold="true" box="[1179,1301,1000,1027]" italics="true" pageId="16" pageNumber="200">L. bezdeki</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="2AAB57DEFFE7596B883F20999A202078" box="[1308,1399,1002,1028]" pageId="16" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
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||
</paragraph>
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||
<paragraph id="035336B7FFE7596B8DE2277E9A2020CA" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
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<keyLead id="B81D9582FFE7596B8DE2277E9A2020CA" pageId="16" pageNumber="200">
|
||
– Paramere apical process of male genitalia rounded apically; paramere basal margin of male genitalia bisinuate; paramere lateral margin of male genitalia surface rugose (
|
||
<figureCitation id="9BD72A32FFE7596B893727429BCA2037" box="[1044,1181,1073,1099]" captionStart="Fig" captionStartId="10.[189,233,1962,1988]" captionTargetBox="[202,1398,500,1918]" captionTargetPageId="10" captionText="Fig. 11–14. Leucopholis semperi Brenske, 1896, neotype, ♂ (MNHN). 11. Habitus, dorsal aspect. 12. Metaventral process, dorsal aspect. 13. Male genitalia, dorsal aspect. 14. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306342" httpUri="https://zenodo.org/record/8306342/files/figure.png" pageId="16" pageNumber="200">Figs 13–14</figureCitation>
|
||
); medial posterior margin of pronotum distinctly extended posteriorly; posterolateral margin of elytron near posterior angle vertically flattened laterally; pygidium without vertical flattened margin subposteriorly, with sloping downward margin. length 36.0 mm ..............................................
|
||
<taxonomicName id="C4EC4D34FFE7596B896427EE9A2020CA" ID-CoL="6Q6HW" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[1095,1399,1180,1207]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="semperi">
|
||
<emphasis id="3198EAA5FFE7596B896427EE9B9120CA" bold="true" box="[1095,1222,1180,1207]" italics="true" pageId="16" pageNumber="200">L. semperi</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B89EE27EF9A2020CA" author="Brenske E." box="[1229,1399,1180,1206]" pageId="16" pageNumber="200" pagination="178 - 205" refId="ref7881" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="B8182E12FFE7596B8D9E27AD9A202161" pageId="16" pageNumber="200">
|
||
<paragraph id="035336B7FFE7596B8D9E27AD9A202084" blockId="16.[189,1400,265,1758]" box="[189,1399,1246,1273]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8D9E27AD9A202084" box="[189,1399,1246,1273]" pageId="16" pageNumber="200">5. Scales on elytra nearly uniform in size ............................................................................................. 6</keyLead>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFE7596B8DE226709A202161" blockId="16.[189,1400,265,1758]" box="[193,1399,1282,1309]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8DE226709A202161" box="[193,1399,1282,1309]" pageId="16" pageNumber="200">
|
||
– Scales on elytra variable in size; length
|
||
<quantity id="C4149B52FFE7596B8F9E26719C3F2161" box="[701,872,1282,1309]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.95" metricValueMin="2.35" pageId="16" pageNumber="200" unit="mm" value="26.5" valueMax="29.5" valueMin="23.5">23.5–29.5 mm</quantity>
|
||
................
|
||
<taxonomicName id="C4EC4D34FFE7596B8EC826709A202161" ID-CoL="724SZ" authority="Burmeister, 1855" authorityName="Burmeister" authorityYear="1855" box="[1003,1399,1282,1309]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="pulverulenta">
|
||
<emphasis id="3198EAA5FFE7596B8EC826709BF32161" bold="true" box="[1003,1188,1282,1309]" italics="true" pageId="16" pageNumber="200">L. pulverulenta</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B898F26719A202161" author="Burmeister H. C. C." box="[1196,1399,1282,1309]" pageId="16" pageNumber="200" refId="ref7902" refString="Burmeister H. C. C. 1855. Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin." type="book" year="1855">Burmeister, 1855</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="B8182E12FFE7596B8D9E26379A2021FE" pageId="16" pageNumber="200">
|
||
<paragraph id="035336B7FFE7596B8D9E26379A202123" blockId="16.[189,1400,265,1758]" box="[189,1399,1348,1375]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8D9E26379A202123" box="[189,1399,1348,1375]" pageId="16" pageNumber="200">6. Clypeus without medial cleft ............................................................................................................ 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFE7596B8DE2261A9A2021FE" blockId="16.[189,1400,265,1758]" box="[193,1399,1384,1411]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8DE2261A9A2021FE" box="[193,1399,1384,1411]" pageId="16" pageNumber="200">– Clypeus with medial cleft ................................................................................................................. 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="B8182E12FFE7596B8D9E26D89A20224C" pageId="16" pageNumber="200">
|
||
<paragraph id="035336B7FFE7596B8D9E26D89A202194" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8D9E26D89A202194" pageId="16" pageNumber="200">
|
||
7. Clypeus strongly convex in males, weakly convex in females, wider than labrum in males, narrow in females; length
|
||
<quantity id="C4149B52FFE7596B8CE626BD9D382195" box="[453,623,1486,1513]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.25" metricValueMin="2.85" pageId="16" pageNumber="200" unit="mm" value="30.5" valueMax="32.5" valueMin="28.5">28.5–32.5 mm</quantity>
|
||
...................................................................
|
||
<taxonomicName id="C4EC4D34FFE7596B894B26BD9A202194" ID-CoL="7254Z" authority="Moser, 1924" authorityName="Moser" authorityYear="1924" box="[1128,1399,1486,1513]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="reflexa">
|
||
<emphasis id="3198EAA5FFE7596B894B26BD9B8B2195" bold="true" box="[1128,1244,1486,1513]" italics="true" pageId="16" pageNumber="200">L. reflexa</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B89C026BC9A202194" author="Moser J." box="[1251,1399,1486,1513]" pageId="16" pageNumber="200" pagination="349 - 357" refId="ref8123" refString="Moser J. 1924. New species of melolonthid beetles from the Philippine Islands. The Philippine Journal of Science 25: 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFE7596B8DE226819A20224C" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8DE226819A20224C" pageId="16" pageNumber="200">
|
||
– Clypeus weakly convex in males, strongly convex in females, nearly as wide as labrum in both sex; length
|
||
<quantity id="C4149B52FFE7596B8C1825659EB2224C" box="[315,485,1558,1585]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" metricValueMax="2.51" metricValueMin="2.25" pageId="16" pageNumber="200" unit="mm" value="23.8" valueMax="25.1" valueMin="22.5">22.5–25.1 mm</quantity>
|
||
.......................................................................................
|
||
<taxonomicName id="C4EC4D34FFE7596B894C25659A20224C" ID-CoL="724TK" authority="Moser, 1924" authorityName="Moser" authorityYear="1924" box="[1135,1399,1557,1584]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="3198EAA5FFE7596B894C25659B8B224C" bold="true" box="[1135,1244,1557,1584]" italics="true" pageId="16" pageNumber="200">L. bakeri</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B89C025659A20224C" author="Moser J." box="[1251,1399,1558,1584]" pageId="16" pageNumber="200" pagination="349 - 357" refId="ref8123" refString="Moser J. 1924. New species of melolonthid beetles from the Philippine Islands. The Philippine Journal of Science 25: 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="B8182E12FFE7596B8D9E252B9A2022A2" pageId="16" pageNumber="200">
|
||
<paragraph id="035336B7FFE7596B8D9E252B9A2022EA" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8D9E252B9A2022EA" pageId="16" pageNumber="200">
|
||
8. Protibial spur long, extending beyond anterior angle of protibia; length
|
||
<quantity id="C4149B52FFE7596B8912252B9B92220E" box="[1073,1221,1624,1651]" metricMagnitude="-2" metricUnit="m" metricValue="2.775" metricValueMax="3.0" metricValueMin="2.55" pageId="16" pageNumber="200" unit="mm" value="27.75" valueMax="30.0" valueMin="25.5">25.5–30 mm</quantity>
|
||
...................... .................................................................................................................
|
||
<taxonomicName id="C4EC4D34FFE7596B8910250F9A2022EA" ID-CoL="6PSTF" authority="Chevrolat, 1841" authorityName="Chevrolat" authorityYear="1841" box="[1075,1399,1660,1686]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="irrorata">
|
||
<emphasis id="3198EAA5FFE7596B8910250F9BE422EA" bold="true" box="[1075,1203,1660,1686]" italics="true" pageId="16" pageNumber="200">L. irrorata</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B899A250F9A2022EA" author="Chevrolat L. A." box="[1209,1399,1660,1686]" pageId="16" pageNumber="200" pagination="221 - 228" refId="ref7998" refString="Chevrolat L. A. 1841. Description de trente quatre especes de Coleopteres de Manille et d'un tricondyle de Ceylan. Revue Zoologique 1841: 221 - 228." type="journal article" year="1841">Chevrolat, 1841</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="035336B7FFE7596B8DE225D39A2022A2" blockId="16.[189,1400,265,1758]" pageId="16" pageNumber="200">
|
||
<keyLead id="B81D9582FFE7596B8DE225D39A2022A2" pageId="16" pageNumber="200">
|
||
– Protibial spur distinctly shorter, not extending beyond anterior angle of protibia; length
|
||
<quantity id="C4149B52FFE7596B886A25D39E0422A2" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.25" metricValueMin="2.1" pageId="16" pageNumber="200" unit="mm" value="21.75" valueMax="22.5" valueMin="21.0">21– 22.5 mm</quantity>
|
||
................................................................................
|
||
<taxonomicName id="C4EC4D34FFE7596B8E8C25B79A2022A2" ID-CoL="6PSTH" authority="Calcetas & Adorada, 2017" authorityName="Calcetas & Adorada" authorityYear="2017" box="[943,1399,1731,1758]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="200" phylum="Arthropoda" rank="species" species="guevarai">
|
||
<emphasis id="3198EAA5FFE7596B8E8C25B79B6D22A1" bold="true" box="[943,1082,1731,1758]" italics="true" pageId="16" pageNumber="200">L. guevarai</emphasis>
|
||
<bibRefCitation id="677D4B46FFE7596B896225B09A2022A2" author="Calcetas O. A. & Adorada J. R." box="[1089,1399,1731,1758]" pageId="16" pageNumber="200" pagination="85 - 103" refId="ref7941" refString="Calcetas O. A. & Adorada J. R. 2017. Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines. Zootaxa 4232 (1): 85 - 103. https: // doi. org / 10.11646 / zootaxa. 4232.1.6" type="journal article" year="2017">Calcetas & Adorada, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |