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<document id="013DD512F0C31AB93100C0A6225100CA" ID-CLB-Dataset="264974" ID-DOI="10.5852/ejt.2023.890.2261" ID-GBIF-Dataset="ea7ad0da-111f-4709-819b-6a84832ceec5" ID-ISSN="2118-9773" ID-Zenodo-Dep="8305633" ID-ZooBank="5BA69DED-87A2-4041-886F-E11A6D38D52D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693494219877" checkinUser="felipe" docAuthor="Calcetas, Orlando A." docDate="2023" docId="8B4587A1FFF5597E8F4E20DC9E8C2395" docLanguage="en" docName="ejt-890_184-203.pdf" docOrigin="European Journal of Taxonomy 890" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucopholis stainesi Calcetas 2023, sp. nov." docType="treatment" docUuid="760739D3-424D-4699-83D0-B7B3A2C5E9B0" docUuidSource="ZooBank" docVersion="6" lastPageNumber="189" masterDocId="777CFFD9FFF7597B8D2323739F57247C" masterDocTitle="Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896" masterLastPageNumber="203" masterPageNumber="184" pageNumber="186" updateTime="1699623951618" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D925DD0AD44FAE2F0B46E515B38F0FFD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E6C7B6210AC2EAFE85A1126D49A4BFBE">
<mods:title id="3098A0BFCD28897797BACC26CAB5CBEA">Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896</mods:title>
</mods:titleInfo>
<mods:name id="8CEC8B16C32662F53AD0ECACF7BC1A97" type="personal">
<mods:role id="29C0F718EBA6A2D158301836347A4F7A">
<mods:roleTerm id="E2AD727E6B4FCBFDFB6F49C86B007D2C">Author</mods:roleTerm>
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<mods:namePart id="70BBEE83A7686B878E3AC7B80871E517">Calcetas, Orlando A.</mods:namePart>
<mods:nameIdentifier id="761C9E2A93DB6B53FDEA7A1F54FA460F" type="LSID">5A1D3F2A-DB38-4170-987C-783749F2E1BD</mods:nameIdentifier>
<mods:affiliation id="E610432863873BC9CD3CB2F2F52C70F2">Department of Agriculture, IVA-CALABARZON, Regional Crop Protection Center (DA-RCPC) Lipa Agricultural Research and Experiment Station (LARES) Brgy. Maraouy, Lipa City, Batangas 4217, Philippines</mods:affiliation>
<mods:nameIdentifier id="4F73FACEDA26765A3EC1F4C3372FB9B9" type="email">orlando.calcetas@calabarzon.da.gov.ph &amp; orly.calcetas@yahoo.com.ph</mods:nameIdentifier>
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<mods:title id="C61B5030487DB8813ACD6638736D739B">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="03D9AB59CE9FF4625139FFF424F0BFDF">
<mods:date id="A67E65EB4D2947D4BFA60D6620330083">2023</mods:date>
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<mods:number id="1D1A201CCE2CECF362F1C422F87A23CF">2023-08-31</mods:number>
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<mods:number id="A54A95A9ACD481C7DA40CB4248DB646B">890</mods:number>
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<mods:classification id="F64F89D3E418BDCDBFDB2B0E68148B6A">journal article</mods:classification>
<mods:identifier id="1221E8D8493D7396360575BA021C9C3C" type="CLB-Dataset">264974</mods:identifier>
<mods:identifier id="B2B40BB2D26FB330F43133586EC094B8" type="DOI">10.5852/ejt.2023.890.2261</mods:identifier>
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<treatment id="8B4587A1FFF5597E8F4E20DC9E8C2395" ID-DOI="http://doi.org/10.5281/zenodo.8306334" ID-GBIF-Taxon="212742584" ID-Zenodo-Dep="8306334" LSID="urn:lsid:zoobank.org:act:760739D3-424D-4699-83D0-B7B3A2C5E9B0" httpUri="http://treatment.plazi.org/id/8B4587A1FFF5597E8F4E20DC9E8C2395" lastPageId="5" lastPageNumber="189" pageId="2" pageNumber="186">
<subSubSection id="4BF6653CFFF559798F4E20DC9C9127B6" box="[621,966,943,970]" pageId="2" pageNumber="186" type="nomenclature">
<paragraph id="035336B7FFF559798F4E20DC9C9127B6" blockId="2.[368,1219,943,1040]" box="[621,966,943,970]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798F4E20DC9C9127B6" box="[621,966,943,970]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="186" reason="2">
<taxonomicName id="C4EC4D34FFF559798F4E20DC9C3327B5" ID-CoL="BSCT6" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[621,868,943,970]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFF559798F4E20DC9C3327B5" bold="true" box="[621,868,943,970]" italics="true" pageId="2" pageNumber="186">Leucopholis stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF559798E4F20C39C9127B6" box="[876,966,944,970]" pageId="2" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF559798C5320A09C1A206C" pageId="2" pageNumber="186" type="description">
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<uri id="777D3AB5FFF559798C5320A09B942792" box="[368,1219,979,1006]" pageId="2" pageNumber="186">
urn:lsid:zoobank.org:act:
<uuid id="774A0C62FFF559798FB020A09B942792" box="[659,1219,979,1006]" pageId="2" pageNumber="186">760739D3-424D-4699-83D0-B7B3A2C5E9B0</uuid>
</uri>
</paragraph>
<paragraph id="035336B7FFF559798FC420859C1A206C" blockId="2.[368,1219,943,1040]" box="[743,845,1014,1040]" pageId="2" pageNumber="186">
<figureCitation id="9BD72A32FFF559798FC420859C1A206C" box="[743,845,1014,1040]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1963,1989]" captionTargetBox-0="[189,1398,556,1908]" captionTargetBox-1="[189,1398,692,1908]" captionTargetId-0="figure-92@3.[347,840,579,1278]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." captionText-1="Fig. 4–6. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 4. Elytron, dorsal aspect. 5. Male genitalia, dorsal aspect. 6. Male genitalia, lateral aspect." figureDoi-0="http://doi.org/10.5281/zenodo.8306332" figureDoi-1="http://doi.org/10.5281/zenodo.8306336" httpUri-0="https://zenodo.org/record/8306332/files/figure.png" httpUri-1="https://zenodo.org/record/8306336/files/figure.png" pageId="2" pageNumber="186">Figs 1–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF559798D9E274B9B3321E2" pageId="2" pageNumber="186" type="diagnosis">
<paragraph id="035336B7FFF559798D9E274B9E91202E" blockId="2.[189,454,1080,1106]" box="[189,454,1080,1106]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798D9E274B9E91202E" bold="true" box="[189,454,1080,1106]" fontSize="11" level="3" pageId="2" pageNumber="186" reason="3">
<emphasis id="3198EAA5FFF559798D9E274B9E91202E" bold="true" box="[189,454,1080,1106]" pageId="2" pageNumber="186">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF559798D9E27149B3321E2" blockId="2.[187,1398,1127,1438]" pageId="2" pageNumber="186">
The new species can be distinguished from all other Philippine
<taxonomicName id="C4EC4D34FFF559798E8527149B6220FD" authorityName="Dejean" authorityYear="1833" box="[934,1077,1127,1153]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFF559798E8527149B6220FD" box="[934,1077,1127,1153]" italics="true" pageId="2" pageNumber="186">Leucopholis</emphasis>
</taxonomicName>
by the metaventral process that has a subapical constriction and is lanceolate subapically while in
<taxonomicName id="C4EC4D34FFF559798ECA27F89B0920D8" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1001,1118,1162,1188]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="3198EAA5FFF559798ECA27F89B0920D8" box="[1001,1118,1162,1188]" italics="true" pageId="2" pageNumber="186">L. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF55979894527F89B9620D9" box="[1126,1217,1163,1189]" pageId="2" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="C4EC4D34FFF5597989E927F89A1220D8" authorityName="Brenske" authorityYear="1896" box="[1226,1349,1163,1188]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="3198EAA5FFF5597989E927F89A1220D8" box="[1226,1349,1163,1188]" italics="true" pageId="2" pageNumber="186">L. semperi</emphasis>
</taxonomicName>
and
<taxonomicName id="C4EC4D34FFF559798D9F27DC9E6A20B4" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[188,317,1198,1224]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
<emphasis id="3198EAA5FFF559798D9F27DC9E6A20B4" box="[188,317,1198,1224]" italics="true" pageId="2" pageNumber="186">L. ratcliffei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF559798C6727DC9ECA20B5" box="[324,413,1199,1225]" pageId="2" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
it is elongate and nearly parallel-sided and without constriction subapically (
<figureCitation id="9BD72A32FFF55979882827DD9A0120B4" box="[1291,1366,1198,1224]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="2" pageNumber="186">Figs 2</figureCitation>
,
<figureCitation id="9BD72A32FFF55979884327DC9A2620B5" box="[1376,1393,1199,1225]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="2" pageNumber="186">7</figureCitation>
,
<figureCitation id="9BD72A32FFF559798D9827A19F8C2090" box="[187,219,1234,1260]" captionStart="Fig" captionStartId="14.[189,231,1963,1989]" captionTargetBox="[228,1398,506,1905]" captionTargetPageId="14" captionText="Fig. 17–20. Leucopholis ratcliffei sp. nov., holotype, ♂ (UNSM). 17. Habitus, dorsal aspect.18. Metaventral process, dorsal aspect. 19. Male genitalia, dorsal aspect. 20. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306348" httpUri="https://zenodo.org/record/8306348/files/figure.png" pageId="2" pageNumber="186">18</figureCitation>
). The abdominal ventrites are covered mostly with elongate ovoid scales in
<taxonomicName id="C4EC4D34FFF55979896027A19BEE2097" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[1091,1209,1234,1259]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFF55979896027A19BEE2097" box="[1091,1209,1234,1259]" italics="true" pageId="2" pageNumber="186">L. stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF5597989E327A19A4F2090" box="[1216,1304,1234,1260]" pageId="2" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
while it is covered mostly with short ovoid scales in all other Philippine
<taxonomicName id="C4EC4D34FFF559798EB427869B722173" authorityName="Dejean" authorityYear="1833" box="[919,1061,1269,1295]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFF559798EB427869B722173" box="[919,1061,1269,1295]" italics="true" pageId="2" pageNumber="186">Leucopholis</emphasis>
</taxonomicName>
with the metaventral process extending in front of the prosternal process. The paramere posterior margin is bisinuate and bowl-shaped in
<taxonomicName id="C4EC4D34FFF559798DFF264E9E02212A" authorityName="Calcetas" authorityYear="2023" box="[220,341,1341,1366]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="stainesi">
<emphasis id="3198EAA5FFF559798DFF264E9E02212A" box="[220,341,1341,1366]" italics="true" pageId="2" pageNumber="186">L. stainesi</emphasis>
</taxonomicName>
and
<taxonomicName id="C4EC4D34FFF559798CAC264E9D5C212A" authorityName="Brenske" authorityYear="1896" box="[399,523,1341,1366]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="3198EAA5FFF559798CAC264E9D5C212A" box="[399,523,1341,1366]" italics="true" pageId="2" pageNumber="186">L. semperi</emphasis>
</taxonomicName>
, but it is nearly straight medially in
<taxonomicName id="C4EC4D34FFF559798E99264E9B64212A" authorityName="Calcetas" authorityYear="2023" box="[954,1075,1341,1366]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="stainesi">
<emphasis id="3198EAA5FFF559798E99264E9B64212A" box="[954,1075,1341,1366]" italics="true" pageId="2" pageNumber="186">L. stainesi</emphasis>
</taxonomicName>
while it is slightly concave medially in
<taxonomicName id="C4EC4D34FFF559798C7326139E9A2105" authorityName="Brenske" authorityYear="1896" box="[336,461,1376,1401]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="3198EAA5FFF559798C7326139E9A2105" box="[336,461,1376,1401]" italics="true" pageId="2" pageNumber="186">L. semperi</emphasis>
</taxonomicName>
. The paramere posterior margin is sinuate, bowl-shaped and slightly concave medially in
<taxonomicName id="C4EC4D34FFF559798C6A26F79E9721E1" authorityName="Calcetas" authorityYear="2023" box="[329,448,1411,1437]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="3198EAA5FFF559798C6A26F79E9721E1" box="[329,448,1411,1437]" italics="true" pageId="2" pageNumber="186">L. bezdeki</emphasis>
</taxonomicName>
while it is concave medially and bisinuate in
<taxonomicName id="C4EC4D34FFF559798EFA26F79B0A21E1" authorityName="Calcetas" authorityYear="2023" box="[985,1117,1411,1437]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="186" phylum="Arthropoda" rank="species" species="ratcliffei">
<emphasis id="3198EAA5FFF559798EFA26F79B0A21E1" box="[985,1117,1411,1437]" italics="true" pageId="2" pageNumber="186">L. ratcliffei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF559798D9E26B69CD8224D" pageId="2" pageNumber="186" type="etymology">
<paragraph id="035336B7FFF559798D9E26B69E1321A3" blockId="2.[189,324,1477,1503]" box="[189,324,1477,1503]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798D9E26B69E1321A3" bold="true" box="[189,324,1477,1503]" fontSize="11" level="3" pageId="2" pageNumber="186" reason="3">
<emphasis id="3198EAA5FFF559798D9E26B69E1321A3" bold="true" box="[189,324,1477,1503]" pageId="2" pageNumber="186">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF559798D9E26879CD8224D" blockId="2.[189,1398,1524,1585]" pageId="2" pageNumber="186">
This new species is named after Dr Charles Staines, world expert on hispines at the Smithsonian Environmental Research Center, Edgewater,
<collectingRegion id="C128F855FFF559798FEF25649C16224D" box="[716,833,1559,1585]" country="United States of America" name="Maryland" pageId="2" pageNumber="186">Maryland</collectingRegion>
,
<collectingCountry id="7BFB7627FFF559798E6E25649CDC224D" box="[845,907,1559,1585]" name="United States of America" pageId="2" pageNumber="186">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF559798D9E252A9A0D22BA" pageId="2" pageNumber="186" type="materials_examined">
<paragraph id="035336B7FFF559798D9E252A9EFD220F" blockId="2.[189,426,1625,1651]" box="[189,426,1625,1651]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798D9E252A9EFD220F" bold="true" box="[189,426,1625,1651]" fontSize="11" level="3" pageId="2" pageNumber="186" reason="3">
<emphasis id="3198EAA5FFF559798D9E252A9EFD220F" bold="true" box="[189,426,1625,1651]" pageId="2" pageNumber="186">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF559798DC925FB9A0D22BA" blockId="2.[189,1370,1672,1734]" pageId="2" pageNumber="186">
<materialsCitation id="B3843CEAFFF559798DC925FB9A0D22BA" ID-GBIF-Occurrence="4402534302" collectingDate="1993-04" collectionCode="PNM" collectorName="V. Samarita" country="Philippines" county="Tiboli" location="Mt. Parker" municipality="Salacapa" pageId="2" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="South Cotabato" typeStatus="holotype">
<emphasis id="3198EAA5FFF559798DC925FB9E0B22DE" bold="true" box="[234,348,1672,1698]" pageId="2" pageNumber="186">
<typeStatus id="DC578815FFF559798DC925FB9E0B22DE" box="[234,348,1672,1698]" pageId="2" pageNumber="186" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7BFB7627FFF559798D9E25D89E3D22B9" box="[189,362,1707,1733]" name="Philippines" pageId="2" pageNumber="186">PHILIPPINES</collectingCountry>
•
<specimenCount id="15EAFD3EFFF559798CA025DF9ECB22BA" box="[387,412,1708,1734]" pageId="2" pageNumber="186" type="male">♂</specimenCount>
;
<collectingRegion id="C128F855FFF559798C8B25D89D3622B9" box="[424,609,1707,1733]" country="Philippines" name="South Cotabato" pageId="2" pageNumber="186">South Cotabato</collectingRegion>
,
<collectingCounty id="EA324E3BFFF559798F4F25D89DE122B9" box="[620,694,1707,1733]" pageId="2" pageNumber="186">Tiboli</collectingCounty>
,
<collectingMunicipality id="E337ACCDFFF559798FE225D89C7B22B9" box="[705,812,1707,1733]" pageId="2" pageNumber="186">Salacapa</collectingMunicipality>
,
<location id="0633606CFFF559798E1B25DF9CEE22B9" LSID="urn:lsid:plazi:treatment:8B4587A1FFF5597E8F4E20DC9E8C2395:0633606CFFF559798E1B25DF9CEE22B9" box="[824,953,1707,1734]" country="Philippines" county="Tiboli" municipality="Salacapa" name="Mt. Parker" pageId="2" pageNumber="186" stateProvince="South Cotabato">Mt. Parker</location>
;
<date id="77521077FFF559798EE725DF9B6B22B9" box="[964,1084,1707,1734]" pageId="2" pageNumber="186" value="1993-04">
<collectingDate id="6716E99FFFF559798EE725DF9B6B22B9" box="[964,1084,1707,1734]" pageId="2" pageNumber="186" value="1993-04">Apr. 1993</collectingDate>
</date>
;
<collectorName id="AE195361FFF55979896B25DF9B8722B9" box="[1096,1232,1707,1734]" pageId="2" pageNumber="186">V. Samarita</collectorName>
leg.;
<collectionCode id="65FDAE72FFF55979883125DF9A0222BA" box="[1298,1365,1708,1734]" collectionName="PNM" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="2" pageNumber="186">PNM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF5597E8D9E259E9E4A2308" lastPageId="5" lastPageNumber="189" pageId="2" pageNumber="186" type="description">
<paragraph id="035336B7FFF559798D9E259E9E0A237B" blockId="2.[189,349,1773,1799]" box="[189,349,1773,1799]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798D9E259E9E0A237B" bold="true" box="[189,349,1773,1799]" fontSize="11" level="3" pageId="2" pageNumber="186" reason="3">
<emphasis id="3198EAA5FFF559798D9E259E9E0A237B" bold="true" box="[189,349,1773,1799]" pageId="2" pageNumber="186">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF559798D9E246F9DC8234B" blockId="2.[189,671,1820,1847]" box="[189,671,1820,1847]" pageId="2" pageNumber="186">
<collectingCountry id="7BFB7627FFF559798D9E246F9E16234A" box="[189,321,1820,1846]" name="Philippines" pageId="2" pageNumber="186">Philippines</collectingCountry>
(Mindanao,
<collectingRegion id="C128F855FFF559798CFB246F9DC4234B" box="[472,659,1820,1847]" country="Philippines" name="South Cotabato" pageId="2" pageNumber="186">South Cotabato</collectingRegion>
).
</paragraph>
<paragraph id="035336B7FFF559798D9E242E9E19230B" blockId="2.[189,334,1885,1911]" box="[189,334,1885,1911]" pageId="2" pageNumber="186">
<heading id="581B81DBFFF559798D9E242E9E19230B" bold="true" box="[189,334,1885,1911]" fontSize="11" level="3" pageId="2" pageNumber="186" reason="3">
<emphasis id="3198EAA5FFF559798D9E242E9E19230B" bold="true" box="[189,334,1885,1911]" pageId="2" pageNumber="186">Description</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF559798D9E24FE9EB323DB" blockId="2.[189,484,1932,1959]" box="[189,484,1932,1959]" pageId="2" pageNumber="186">BODY LENGTH. 35.0 mm.</paragraph>
<paragraph id="035336B7FFF559798D9E24BD9E812395" blockId="2.[189,470,1998,2025]" box="[189,470,1998,2025]" pageId="2" pageNumber="186">BODY WIDTH. 17.0 mm.</paragraph>
<paragraph id="035336B7FFF459788D9E227A9D052515" blockId="3.[188,1399,265,361]" pageId="3" pageNumber="187">
COLOUR. Dorsum dichromatic, head, pronotum, scutellum, elytra and legs blackish; with brownish tinge on posterior margin of elytra; body covered with yellowish white scales. Venter dichromatic blackish with little shade of brown (
<figureCitation id="9BD72A32FFF459788CDF223C9D162515" box="[508,577,335,361]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="3" pageNumber="187">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="035336B7FFF4597F8D9E22E29A3225AF" blockId="3.[187,1399,401,498]" lastBlockId="4.[187,1401,265,467]" lastPageId="4" lastPageNumber="188" pageId="3" pageNumber="187">
HEAD. Clypeus with medial anterior margin nearly straight anteriorly; slightly cleft, slanted at 60° angle laterally; anterior surface lustrous, glabrous, impunctate subanteriorly; above clypeo-labral suture rugose medially; with row of large, rounded to rugose punctures adjacent to posterior margin; each puncture with stiff brownish hair or seta; anterior and lateral margins widely carinate dorsally; with slight medial cleft laterally; anterior angle widely rounded laterally; dorsal surface covered with short, narrow, elongate, parallel-sided, acicular-like, tapered, blunt apically yellowish white scales; scales slightly bent downward. Apical maXillary palpomere rice grain-like, with oval flattened area. Mentum pot-shaped, lustrous, glabrous; with very few hair brushes on each side of medial cleft subanteriorly; with nearly flattened surface medially. Antennal lamellae length
<quantity id="C4149B52FFF3597F8F9A22CA9C4125AF" box="[697,790,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="188" unit="mm" value="3.2">3.2 mm</quantity>
, longer than entire length of antennomeres II–VII.
</paragraph>
<caption id="5793663FFFF459788D9E24D89CB12394" ID-DOI="http://doi.org/10.5281/zenodo.8306332" ID-Zenodo-Dep="8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="3" pageNumber="187" startId="3.[189,232,1963,1989]" targetBox="[189,1398,556,1908]" targetPageId="3">
<paragraph id="035336B7FFF459788D9E24D89CB12394" blockId="3.[189,1398,1963,2025]" pageId="3" pageNumber="187">
<emphasis id="3198EAA5FFF459788D9E24D89E7D23B9" bold="true" box="[189,298,1963,1989]" pageId="3" pageNumber="187">Fig. 1–3.</emphasis>
<taxonomicName id="C4EC4D34FFF459788C1124D89D7223B9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[306,549,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="187" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFF459788C1124D89D7223B9" box="[306,549,1963,1989]" italics="true" pageId="3" pageNumber="187">Leucopholis stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF459788F1324DF9DDA23BA" box="[560,653,1964,1990]" pageId="3" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="DC578815FFF459788FB924D89C5323B9" box="[666,772,1963,1989]" pageId="3" pageNumber="187" type="holotype">holotype</typeStatus>
,
<specimenCount id="15EAFD3EFFF459788E3124DF9C7F23BA" box="[786,808,1964,1990]" pageId="3" pageNumber="187" type="male">♂</specimenCount>
(PNM).
<emphasis id="3198EAA5FFF459788EB724D89CF423B9" bold="true" box="[916,931,1963,1989]" pageId="3" pageNumber="187">1</emphasis>
. Habitus, dorsal aspect.
<emphasis id="3198EAA5FFF4597889E824D89B8F23B9" bold="true" box="[1227,1240,1963,1989]" pageId="3" pageNumber="187">2</emphasis>
. Metaventral process, dorsal aspect.
<emphasis id="3198EAA5FFF459788CE824BD9E8D2394" bold="true" box="[459,474,1998,2024]" pageId="3" pageNumber="187">3</emphasis>
. Elytron, anterior margin and anterior angle.
</paragraph>
</caption>
<paragraph id="035336B7FFF3597E8D9E22889A2F25C7" blockId="4.[187,1399,506,639]" lastBlockId="5.[189,1400,265,443]" lastPageId="5" lastPageNumber="189" pageId="4" pageNumber="188">PRONOTUM. Anterior margin moderately concave, distinctly long nearly straight margin medially; anterior angle obtuse; without lustrous, glabrous, impunctate callosity subapically; rounded, carinate apically; lateral margin widely convex medially; anterolateral margin at ~45° angle, crenulate; posterolateral margin distinctly wedge-shaped, slightly crenulate; posterior angle at 90° angle, strongly sinuate, rounded apically; posterior margin bisinuate; with deep, rounded margin medially; margin distinctly extended posteriorly; surface covered with variable scales; covered mostly with short, narrow, spindle-shaped, tapered to blunt apically scales medio-subanteriorly; with elongate, very narrow, nearly parallel-sided spindle-shaped, tapered apically scales medio-subposteriorly; with short, narrow, lanceolate, tapered apically scales submedially; with short, ovoid, rounded apex scales on each side adjacent to lateral margin.</paragraph>
<caption id="5793663FFFF3597F8D9E24D89D8D2395" ID-DOI="http://doi.org/10.5281/zenodo.8306336" ID-Zenodo-Dep="8306336" httpUri="https://zenodo.org/record/8306336/files/figure.png" pageId="4" pageNumber="188" startId="4.[189,232,1963,1989]" targetBox="[189,1398,692,1908]" targetPageId="4">
<paragraph id="035336B7FFF3597F8D9E24D89D8D2395" blockId="4.[189,1398,1963,2025]" pageId="4" pageNumber="188">
<emphasis id="3198EAA5FFF3597F8D9E24D89E7F23B9" bold="true" box="[189,296,1963,1989]" pageId="4" pageNumber="188">Fig. 4–6.</emphasis>
<taxonomicName id="C4EC4D34FFF3597F8C0D24D89D4823B9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[302,543,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="188" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFF3597F8C0D24D89D4823B9" box="[302,543,1963,1989]" italics="true" pageId="4" pageNumber="188">Leucopholis stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF3597F8F0624DF9DD523BA" box="[549,642,1964,1990]" pageId="4" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="DC578815FFF3597F8FAE24D89DA023B9" box="[653,759,1963,1989]" pageId="4" pageNumber="188" type="holotype">holotype</typeStatus>
,
<specimenCount id="15EAFD3EFFF3597F8E2124DF9C4F23BA" box="[770,792,1964,1990]" pageId="4" pageNumber="188" type="male">♂</specimenCount>
(PNM).
<emphasis id="3198EAA5FFF3597F8EA224D89CC723B9" bold="true" box="[897,912,1963,1989]" pageId="4" pageNumber="188">4</emphasis>
. Elytron, dorsal aspect.
<emphasis id="3198EAA5FFF3597F898924D89BEE23B9" bold="true" box="[1194,1209,1963,1989]" pageId="4" pageNumber="188">5</emphasis>
. Male genitalia, dorsal aspect.
<emphasis id="3198EAA5FFF3597F8C4724BD9E242394" bold="true" box="[356,371,1998,2024]" pageId="4" pageNumber="188">6</emphasis>
. Male genitalia, lateral aspect.
</paragraph>
</caption>
<paragraph id="035336B7FFF2597E8D9E22969BF0269F" blockId="5.[187,1399,485,739]" pageId="5" pageNumber="189">
VENTRAL SIDE OF THORAX. Prosternal process mound-shaped to nearly isosceles triangulate, widely rounded apically; with flattened, lustrous, impunctate area dorsally (
<figureCitation id="9BD72A32FFF2597E8EFF21789B742659" box="[988,1059,523,549]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="5" pageNumber="189">Fig. 2</figureCitation>
). Metaventral process length 5.0 mm; distinctly wide, triangulate basally; elongate apically; constricted subapically; sublateral margin lanceolate; narrow rounded apically; lustrous, glabrous, impunctate dorsally; with straight, narrow, faint, longitudinal dark line medially; faint line much distinct in metasternum (
<figureCitation id="9BD72A32FFF2597E8ED5210E9B6B26EB" box="[1014,1084,637,663]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="5" pageNumber="189">Fig. 2</figureCitation>
). Metasternum with medial depression; depression on each side of metaventral process covered with very thin, acicular-like white scales; metasternum covered mostly with short, spindle-shaped, apically blunt scales.
</paragraph>
<paragraph id="035336B7FFF2597E8D9E207E9BC0275B" blockId="5.[189,1175,781,807]" box="[189,1175,781,807]" pageId="5" pageNumber="189">LEG. Foretibia bidentate, widely rounded apex; posterior metatibia with 20 spicules.</paragraph>
<paragraph id="035336B7FFF2597E8D9E20229A2027ED" blockId="5.[189,1399,849,914]" pageId="5" pageNumber="189">SCUTELLUM. Covered with large, moderate, spindle-shaped to ovoid whitish scales; with elongate, parallel-sided acicular whitish scales; lustrous, impunctate medially, with very few rounded punctures on each side.</paragraph>
<paragraph id="035336B7FFF2597E8D9E20C89D2420A3" blockId="5.[187,1399,955,1247]" pageId="5" pageNumber="189">
ELYTRON. Anterior margin distinctly wedge-shaped towards scutellum; anterior angle widely rounded; thickly carinate, explanate (
<figureCitation id="9BD72A32FFF2597E8CD620929D6E2787" box="[501,569,993,1019]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[189,1398,556,1908]" captionTargetId="figure-92@3.[347,840,579,1278]" captionTargetPageId="3" captionText="Fig. 1–3. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 1. Habitus, dorsal aspect. 2. Metaventral process, dorsal aspect. 3. Elytron, anterior margin and anterior angle." figureDoi="http://doi.org/10.5281/zenodo.8306332" httpUri="https://zenodo.org/record/8306332/files/figure.png" pageId="5" pageNumber="189">Fig. 3</figureCitation>
); with narrow suture towards anterior angle; margin evanescent medially; posterolateral margin vertically flattened towards posterior angle; posterior angle obtuse, widely rounded, minutely explanate; with thin crust-like margin; posterior margin smooth, slightly sinuate towards sutural angle; sutural angle approximately at 90° angle; sutural margin medially and towards sutural angle; distinctly carinate towards scutellum, infleXed, not carinate (
<figureCitation id="9BD72A32FFF2597E8E5A270A9C9720EF" box="[889,960,1145,1171]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,692,1908]" captionTargetPageId="4" captionText="Fig. 4–6. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 4. Elytron, dorsal aspect. 5. Male genitalia, dorsal aspect. 6. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306336" httpUri="https://zenodo.org/record/8306336/files/figure.png" pageId="5" pageNumber="189">Fig. 4</figureCitation>
); each side near sutural angle covered with short, fringed hairs; surface mostly covered with variable sized inverted lanceolate, tapered apically, bent medially yellowish white scales.
</paragraph>
<paragraph id="035336B7FFF2597E8D9E267A9B5621E9" blockId="5.[189,1400,1289,1429]" pageId="5" pageNumber="189">ABDOMEN. Abdominal sternites I–V covered with short, inverted lanceolate, tapered basally, rounded apically scales; sternites III and IV each with small patch of scales medially; patch of scales on sternite III much closer compared to IV; with very few elongated, narrow, parallel-sided scales; scales larger mesally compared to each side; sternite VI with minute to small scales.</paragraph>
<paragraph id="035336B7FFF2597E8D9E26CC9D7E220D" blockId="5.[189,1400,1471,1649]" pageId="5" pageNumber="189">PYGIDIUM. Anterolateral margin thickly carinate towards anterior angle; disappearing medially; with narrow explanate margin medially, much distinct and wide towards posterior margin; posterior margin widely rounded, narrowly explanate, depressed medially; convex laterally; subposterior margin sloping downward laterally; surface rugosely punctured; covered with short, narrow, minute to small spindle-shaped to parallel-sided scales.</paragraph>
<paragraph id="035336B7FFF2597E8D9E25E89E4A2308" blockId="5.[187,1401,1691,1908]" pageId="5" pageNumber="189">
MALE GENITALIA. Genitalia length
<quantity id="C4149B52FFF2597E8F7F25E89DEB22CA" box="[604,700,1691,1718]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="189" unit="mm" value="9.8">9.8 mm</quantity>
. Phallobase with dorsal apical margin wide, bowl-shaped, distinctly convex medially (
<figureCitation id="9BD72A32FFF2597E8F2925B29D0322A0" box="[522,596,1729,1756]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,692,1908]" captionTargetPageId="4" captionText="Fig. 4–6. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 4. Elytron, dorsal aspect. 5. Male genitalia, dorsal aspect. 6. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306336" httpUri="https://zenodo.org/record/8306336/files/figure.png" pageId="5" pageNumber="189">Fig. 5</figureCitation>
). Parameres with dorsal basal margin bowl-shaped, slightly convex medially.Apical process length
<quantity id="C4149B52FFF2597E8F0425949DD6237E" box="[551,641,1767,1794]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="189" unit="mm" value="2.8">2.8 mm</quantity>
, distinctly shorter than phallobase; dog head-like, short, triangulate towards apex, rounded apically; with deep, wide, horse saddle-shaped depression medially; basal margin widely explanate. Lateral margin of paramere with very shallow longitudinal depression on each side (
<figureCitation id="9BD72A32FFF2597E8DE4242A9E582308" box="[199,271,1881,1908]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[189,1398,692,1908]" captionTargetPageId="4" captionText="Fig. 4–6. Leucopholis stainesi sp. nov., holotype, ♂ (PNM). 4. Elytron, dorsal aspect. 5. Male genitalia, dorsal aspect. 6. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306336" httpUri="https://zenodo.org/record/8306336/files/figure.png" pageId="5" pageNumber="189">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF2597E8D9E24EE9E8C2395" box="[189,475,1949,2025]" pageId="5" pageNumber="189" type="distribution">
<paragraph id="035336B7FFF2597E8D9E24EE9E0123CB" blockId="5.[189,342,1949,1975]" box="[189,342,1949,1975]" pageId="5" pageNumber="189">
<heading id="581B81DBFFF2597E8D9E24EE9E0123CB" bold="true" box="[189,342,1949,1975]" fontSize="11" level="3" pageId="5" pageNumber="189" reason="3">
<emphasis id="3198EAA5FFF2597E8D9E24EE9E0123CB" bold="true" box="[189,342,1949,1975]" pageId="5" pageNumber="189">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF2597E8D9E24BD9E8C2395" blockId="5.[189,475,1998,2025]" box="[189,475,1998,2025]" pageId="5" pageNumber="189">
<collectingCountry id="7BFB7627FFF2597E8D9E24BD9E162394" box="[189,321,1998,2024]" name="Philippines" pageId="5" pageNumber="189">Philippines</collectingCountry>
(Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>
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