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<document id="8953581060DC8B3300048A3F377250E5" ID-CLB-Dataset="264974" ID-DOI="10.5852/ejt.2023.890.2261" ID-GBIF-Dataset="ea7ad0da-111f-4709-819b-6a84832ceec5" ID-ISSN="2118-9773" ID-Zenodo-Dep="8305633" ID-ZooBank="5BA69DED-87A2-4041-886F-E11A6D38D52D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693494219877" checkinUser="felipe" docAuthor="Calcetas, Orlando A." docDate="2023" docId="8B4587A1FFF159738F4D227B9E8C2290" docLanguage="en" docName="ejt-890_184-203.pdf" docOrigin="European Journal of Taxonomy 890" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucopholis bezdeki Calcetas 2023, sp. nov." docType="treatment" docUuid="24A67CFE-43C3-41EF-B115-074908EE1C72" docUuidSource="ZooBank" docVersion="6" lastPageNumber="192" masterDocId="777CFFD9FFF7597B8D2323739F57247C" masterDocTitle="Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896" masterLastPageNumber="203" masterPageNumber="184" pageNumber="190" updateTime="1699623951618" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F3C0B64215F7A4F8BC3153855AE193D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="782F7FDD9F4B0B4A474B83EAAC127CCD">
<mods:title id="315E68B9829C9D8F660B107ACA69AFA2">Three new species of the genus Leucopholis Dejean, 1833 (Coleoptera, Scarabaeidae, Melolonthinae, Leucopholini) from the Philippines and designation of a neotype for L. semperi Brenske, 1896</mods:title>
</mods:titleInfo>
<mods:name id="008F0DF815CE06CC07EB7CACB8B5C870" type="personal">
<mods:role id="E6424F049553ACA79F22AF0AC7822E85">
<mods:roleTerm id="18CD004E7943BD4F7346FDE62D112FCE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A70C15DD361CB9131F0319F9F71FA1E7">Calcetas, Orlando A.</mods:namePart>
<mods:nameIdentifier id="3BAC4D096692FF73538EB9D096FAE55F" type="LSID">5A1D3F2A-DB38-4170-987C-783749F2E1BD</mods:nameIdentifier>
<mods:affiliation id="E3CC2E0EF3ED3AEAEE9D8E4424886F02">Department of Agriculture, IVA-CALABARZON, Regional Crop Protection Center (DA-RCPC) Lipa Agricultural Research and Experiment Station (LARES) Brgy. Maraouy, Lipa City, Batangas 4217, Philippines</mods:affiliation>
<mods:nameIdentifier id="9209F72E32DB01CE6A26157F9935EA20" type="email">orlando.calcetas@calabarzon.da.gov.ph &amp; orly.calcetas@yahoo.com.ph</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="751DA4A212ADA600AF315745B00A9E31">text</mods:typeOfResource>
<mods:relatedItem id="1FABD1966D73F8415F627272CC410151" type="host">
<mods:titleInfo id="ECE99FE13F67B766BB2A28F744274865">
<mods:title id="04342D3B0C987ECD64AB44BFB22A54EE">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="DBCD5E2703CA4AC861CEC5457A4B97D9">
<mods:date id="213DEFD27DF514D2EA0942CA86665FD4">2023</mods:date>
<mods:detail id="7C27D38E51E89CD3BEDDD96E580790F0" type="pubDate">
<mods:number id="B68ECFC313E32FC8080E3B9AF2AEF29E">2023-08-31</mods:number>
</mods:detail>
<mods:detail id="73C14369C13D1F83EF0E4083CDCD184D" type="volume">
<mods:number id="C582B26C886CAA6C7F2B60B8F1DA6834">890</mods:number>
</mods:detail>
<mods:extent id="9D21729355F34569A27317F3A035EB1B" unit="page">
<mods:start id="4C8869BE8F1A955C6D6110AB69CC5F13">184</mods:start>
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</mods:extent>
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<mods:classification id="ABCD4E2BAE86AE2E1B5E6F01454F8C41">journal article</mods:classification>
<mods:identifier id="AC792DB29B3434477B77BB497B15FB9B" type="CLB-Dataset">264974</mods:identifier>
<mods:identifier id="C40771E7DAA501F381F656EF170A7CFD" type="DOI">10.5852/ejt.2023.890.2261</mods:identifier>
<mods:identifier id="6214EF82B8CD300FFDF387F42BF2CBCE" type="GBIF-Dataset">ea7ad0da-111f-4709-819b-6a84832ceec5</mods:identifier>
<mods:identifier id="A0646A8388E48036690675858E6B5C1B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="40DEBEC93C545BE10361FE2956566C74" type="Zenodo-Dep">8305633</mods:identifier>
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</mods:mods>
<treatment id="8B4587A1FFF159738F4D227B9E8C2290" ID-DOI="http://doi.org/10.5281/zenodo.8306338" ID-GBIF-Taxon="212742585" ID-Zenodo-Dep="8306338" LSID="urn:lsid:zoobank.org:act:24A67CFE-43C3-41EF-B115-074908EE1C72" httpUri="http://treatment.plazi.org/id/8B4587A1FFF159738F4D227B9E8C2290" lastPageId="8" lastPageNumber="192" pageId="6" pageNumber="190">
<subSubSection id="4BF6653CFFF1597D8F4D227B9C91255F" box="[622,966,264,291]" pageId="6" pageNumber="190" type="nomenclature">
<paragraph id="035336B7FFF1597D8F4D227B9C91255F" blockId="6.[372,1216,264,362]" box="[622,966,264,291]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8F4D227B9C91255F" box="[622,966,264,291]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="190" reason="2">
<taxonomicName id="C4EC4D34FFF1597D8F4D227B9C33255F" ID-CoL="BSCT9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[622,868,264,291]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="3198EAA5FFF1597D8F4D227B9C33255F" bold="true" box="[622,868,264,291]" italics="true" pageId="6" pageNumber="190">Leucopholis bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF1597D8E48227A9C91255F" box="[875,966,265,291]" pageId="6" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF1597D8C57225F9C032516" pageId="6" pageNumber="190" type="description">
<paragraph id="035336B7FFF1597D8C57225F9B97253B" blockId="6.[372,1216,264,362]" box="[372,1216,300,327]" pageId="6" pageNumber="190">
<uri id="777D3AB5FFF1597D8C57225F9B97253B" box="[372,1216,300,327]" pageId="6" pageNumber="190">
urn:lsid:zoobank.org:act:
<uuid id="774A0C62FFF1597D8FB7225F9B97253B" box="[660,1216,300,327]" pageId="6" pageNumber="190">24A67CFE-43C3-41EF-B115-074908EE1C72</uuid>
</uri>
</paragraph>
<paragraph id="035336B7FFF1597D8FFC223C9C032516" blockId="6.[372,1216,264,362]" box="[735,852,335,362]" pageId="6" pageNumber="190">
<figureCitation id="9BD72A32FFF1597D8FFC223C9C032516" box="[735,852,335,362]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="6" pageNumber="190">Figs 7–10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF1597D8D9E22E29A0226D1" pageId="6" pageNumber="190" type="diagnosis">
<paragraph id="035336B7FFF1597D8D9E22E29E9125D7" blockId="6.[189,454,401,427]" box="[189,454,401,427]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8D9E22E29E9125D7" bold="true" box="[189,454,401,427]" fontSize="11" level="3" pageId="6" pageNumber="190" reason="3">
<emphasis id="3198EAA5FFF1597D8D9E22E29E9125D7" bold="true" box="[189,454,401,427]" pageId="6" pageNumber="190">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF1597D8D9E22CC9A0226D1" blockId="6.[187,1400,447,685]" pageId="6" pageNumber="190">
The new species can be distinguished from all other Philippine
<taxonomicName id="C4EC4D34FFF1597D8E9522CC9B1225A5" authorityName="Dejean" authorityYear="1833" box="[950,1093,447,473]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFF1597D8E9522CC9B1225A5" box="[950,1093,447,473]" italics="true" pageId="6" pageNumber="190">Leucopholis</emphasis>
</taxonomicName>
by the paramere with the apical process bird bill-like, distinctly shorter than the phallobase, tapering towards the apex and narrowly rounded apically. The pronotum has the posterior angle obtuse, rounded apically and slightly sinuate in
<taxonomicName id="C4EC4D34FFF1597D8D9E21599E63263F" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[189,308,553,579]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="3198EAA5FFF1597D8D9E21599E63263F" box="[189,308,553,579]" italics="true" pageId="6" pageNumber="190">L. bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF1597D8C1921599EC22638" box="[314,405,554,580]" pageId="6" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
while it is either nearly or slightly above 90° angle and distinctly sinuate in the other six species of Philippine
<taxonomicName id="C4EC4D34FFF1597D8CC4213F9D21261A" authorityName="Dejean" authorityYear="1833" box="[487,630,588,614]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="genus">
<emphasis id="3198EAA5FFF1597D8CC4213F9D21261A" box="[487,630,588,614]" italics="true" pageId="6" pageNumber="190">Leucopholis</emphasis>
</taxonomicName>
with the metaventral process extending in front of the prosternal process. The posterolateral margin of the elytron near the posterior angle is distinctly infleXed in
<taxonomicName id="C4EC4D34FFF1597D882021039A2026F6" authorityName="Calcetas" authorityYear="2023" box="[1283,1399,624,650]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="bezdeki">
<emphasis id="3198EAA5FFF1597D882021039A2026F6" box="[1283,1399,624,650]" italics="true" pageId="6" pageNumber="190">L. bezdeki</emphasis>
</taxonomicName>
and
<taxonomicName id="C4EC4D34FFF1597D8DCC21E79E2426D1" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[239,371,659,685]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="ratcliffei" status="sp. nov.">
<emphasis id="3198EAA5FFF1597D8DCC21E79E2426D1" box="[239,371,659,685]" italics="true" pageId="6" pageNumber="190">L. ratcliffei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF1597D8C5921E09E8226D1" box="[378,469,659,685]" pageId="6" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
while it is vertically flattened laterally in
<taxonomicName id="C4EC4D34FFF1597D8E9C21E79B6F26D1" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[959,1080,660,685]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="stainesi" status="sp. nov.">
<emphasis id="3198EAA5FFF1597D8E9C21E79B6F26D1" box="[959,1080,660,685]" italics="true" pageId="6" pageNumber="190">L. stainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF1597D891D21E09BCE26D1" box="[1086,1177,659,685]" pageId="6" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="C4EC4D34FFF1597D89F121E79A1926D1" authorityName="Brenske" authorityYear="1896" box="[1234,1358,660,685]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="190" phylum="Arthropoda" rank="species" species="semperi">
<emphasis id="3198EAA5FFF1597D89F121E79A1926D1" box="[1234,1358,660,685]" italics="true" pageId="6" pageNumber="190">L. semperi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF1597D8D9E21A69DB2273C" pageId="6" pageNumber="190" type="etymology">
<paragraph id="035336B7FFF1597D8D9E21A69E132693" blockId="6.[189,324,725,751]" box="[189,324,725,751]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8D9E21A69E132693" bold="true" box="[189,324,725,751]" fontSize="11" level="3" pageId="6" pageNumber="190" reason="3">
<emphasis id="3198EAA5FFF1597D8D9E21A69E132693" bold="true" box="[189,324,725,751]" pageId="6" pageNumber="190">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF1597D8D9E20709DB2273C" blockId="6.[189,1399,771,833]" pageId="6" pageNumber="190">
This new species is named after Dr Aleš Bezděk (Biology Centre CAS, České Budějovice,
<collectingCountry id="7BFB7627FFF1597D880F20709E7B273D" name="Czech Republic" pageId="6" pageNumber="190">Czech Republic</collectingCountry>
), a specialist on Asian melolonthines.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF1597D8D9E20149A7A2045" pageId="6" pageNumber="190" type="materials_examined">
<paragraph id="035336B7FFF1597D8D9E20149EFD27FD" blockId="6.[189,426,871,897]" box="[189,426,871,897]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8D9E20149EFD27FD" bold="true" box="[189,426,871,897]" fontSize="11" level="3" pageId="6" pageNumber="190" reason="3">
<emphasis id="3198EAA5FFF1597D8D9E20149EFD27FD" bold="true" box="[189,426,871,897]" pageId="6" pageNumber="190">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF1597D8DC920E59A5627A8" blockId="6.[189,1281,918,980]" pageId="6" pageNumber="190">
<materialsCitation id="B3843CEAFFF1597D8DC920E59A5627A8" ID-GBIF-Occurrence="4402534305" collectingDate="1993-06" collectionCode="PNM" collectorName="V. Samarita" country="Philippines" location="Watershed Camp-II" pageId="6" pageNumber="190" specimenCount="1" specimenCount-male="1" stateProvince="ZamboanGa City" typeStatus="holotype">
<emphasis id="3198EAA5FFF1597D8DC920E59E0B27CC" bold="true" box="[234,348,918,944]" pageId="6" pageNumber="190">
<typeStatus id="DC578815FFF1597D8DC920E59E0B27CC" box="[234,348,918,944]" pageId="6" pageNumber="190" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7BFB7627FFF1597D8D9E20C99E3D27A8" box="[189,362,954,980]" name="Philippines" pageId="6" pageNumber="190">PHILIPPINES</collectingCountry>
•
<specimenCount id="15EAFD3EFFF1597D8CA020C99ECB27A8" box="[387,412,954,980]" pageId="6" pageNumber="190" type="male">♂</specimenCount>
;
<collectingRegion id="C128F855FFF1597D8C8B20CA9D3927AF" box="[424,622,953,979]" pageId="6" pageNumber="190">Zamboanga City</collectingRegion>
,
<location id="0633606CFFF1597D8F5A20CA9C3527A8" LSID="urn:lsid:plazi:treatment:8B4587A1FFF159738F4D227B9E8C2290:0633606CFFF1597D8F5A20CA9C3527A8" box="[633,866,953,980]" country="Philippines" name="Watershed Camp-II" pageId="6" pageNumber="190" stateProvince="ZamboanGa City">Watershed Camp-II</location>
;
<date id="77521077FFF1597D8E4D20C99CB427A8" box="[878,995,954,980]" pageId="6" pageNumber="190" value="1993-06">
<collectingDate id="6716E99FFFF1597D8E4D20C99CB427A8" box="[878,995,954,980]" pageId="6" pageNumber="190" value="1993-06">Jun. 1993</collectingDate>
</date>
;
<collectorName id="AE195361FFF1597D8ECC20C99B2027AF" box="[1007,1143,953,980]" pageId="6" pageNumber="190">V. Samarita</collectorName>
leg.;
<collectionCode id="65FDAE72FFF1597D899A20C99BAB27A8" box="[1209,1276,954,980]" collectionName="PNM" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="6" pageNumber="190">PNM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="035336B7FFF1597D8DC920889A7A2045" blockId="6.[189,1325,1019,1081]" pageId="6" pageNumber="190">
<materialsCitation id="B3843CEAFFF1597D8DC920889A7F2045" ID-GBIF-Occurrence="4402534303" collectingDate="1993-06" collectionCode="PNM" collectorName="V. Samarita" country="Philippines" location="Watershed Camp-II" pageId="6" pageNumber="190" specimenCount="3" specimenCount-female="3" stateProvince="ZamboanGa City" typeStatus="paratype">
<emphasis id="3198EAA5FFF1597D8DC920889E302069" bold="true" box="[234,359,1019,1045]" pageId="6" pageNumber="190">
<typeStatus id="DC578815FFF1597D8DC920889E302069" box="[234,359,1019,1045]" pageId="6" pageNumber="190" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7BFB7627FFF1597D8D9E276D9E3D2044" box="[189,362,1054,1080]" name="Philippines" pageId="6" pageNumber="190">PHILIPPINES</collectingCountry>
•
<specimenCount id="15EAFD3EFFF1597D8CA0276D9E912045" box="[387,454,1054,1081]" pageId="6" pageNumber="190" type="female">3 ♀♀</specimenCount>
;
<collectingRegion id="C128F855FFF1597D8CF7276D9DCD2044" box="[468,666,1054,1080]" pageId="6" pageNumber="190">Zamboanga City</collectingRegion>
,
<location id="0633606CFFF1597D8F86276D9CD92044" LSID="urn:lsid:plazi:treatment:8B4587A1FFF159738F4D227B9E8C2290:0633606CFFF1597D8F86276D9CD92044" box="[677,910,1054,1080]" country="Philippines" name="Watershed Camp-II" pageId="6" pageNumber="190" stateProvince="ZamboanGa City">Watershed Camp-II</location>
;
<date id="77521077FFF1597D8EB9276C9B582044" box="[922,1039,1054,1081]" pageId="6" pageNumber="190" value="1993-06">
<collectingDate id="6716E99FFFF1597D8EB9276C9B582044" box="[922,1039,1054,1081]" pageId="6" pageNumber="190" value="1993-06">Jun. 1993</collectingDate>
</date>
;
<collectorName id="AE195361FFF1597D8938276C9BF42044" box="[1051,1187,1054,1081]" pageId="6" pageNumber="190">V. Samarita</collectorName>
leg.;
<collectionCode id="65FDAE72FFF1597D89C6276C9A7F2045" box="[1253,1320,1055,1081]" collectionName="PNM" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="6" pageNumber="190">PNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFF159738D9E27139DF62207" lastPageId="8" lastPageNumber="192" pageId="6" pageNumber="190" type="description">
<paragraph id="035336B7FFF1597D8D9E27139E0A2005" blockId="6.[189,349,1119,1146]" box="[189,349,1119,1146]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8D9E27139E0A2005" bold="true" box="[189,349,1119,1146]" fontSize="11" level="3" pageId="6" pageNumber="190" reason="3">
<emphasis id="3198EAA5FFF1597D8D9E27139E0A2005" bold="true" box="[189,349,1119,1146]" pageId="6" pageNumber="190">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF1597D8D9E27FD9DFA20D5" blockId="6.[189,685,1166,1193]" box="[189,685,1166,1193]" pageId="6" pageNumber="190">
<collectingCountry id="7BFB7627FFF1597D8D9E27FD9E1620D4" box="[189,321,1166,1192]" name="Philippines" pageId="6" pageNumber="190">Philippines</collectingCountry>
(Mindanao,
<collectingRegion id="C128F855FFF1597D8CFB27FD9D3520D4" box="[472,610,1166,1192]" country="Philippines" name="Zamboanga" pageId="6" pageNumber="190">Zamboanga</collectingRegion>
City).
</paragraph>
<paragraph id="035336B7FFF1597D8D9E27BC9E192095" blockId="6.[189,334,1231,1257]" box="[189,334,1231,1257]" pageId="6" pageNumber="190">
<heading id="581B81DBFFF1597D8D9E27BC9E192095" bold="true" box="[189,334,1231,1257]" fontSize="11" level="3" pageId="6" pageNumber="190" reason="3">
<emphasis id="3198EAA5FFF1597D8D9E27BC9E192095" bold="true" box="[189,334,1231,1257]" pageId="6" pageNumber="190">Description</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFF1597D8D9E278D9EB32165" blockId="6.[189,484,1278,1305]" box="[189,484,1278,1305]" pageId="6" pageNumber="190">
BODY LENGTH.
<quantity id="C4149B52FFF1597D8C50278D9E882165" box="[371,479,1278,1305]" metricMagnitude="-2" metricUnit="m" metricValue="3.82" pageId="6" pageNumber="190" unit="mm" value="38.2">38.2 mm</quantity>
.
</paragraph>
<paragraph id="035336B7FFF1597D8D9E26339E812126" blockId="6.[189,470,1344,1370]" box="[189,470,1344,1370]" pageId="6" pageNumber="190">
BODY WIDTH.
<quantity id="C4149B52FFF1597D8C4526339E852126" box="[358,466,1344,1370]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="6" pageNumber="190" unit="mm" value="19.1">19.1 mm</quantity>
.
</paragraph>
<paragraph id="035336B7FFF1597D8D9E26F29A6521E0" blockId="6.[189,1330,1409,1436]" box="[189,1330,1409,1436]" pageId="6" pageNumber="190">
COLOUR. Dorsum and venter monochromatic black; covered with yellowish white scales (
<figureCitation id="9BD72A32FFF1597D89FF26F29A7321E0" box="[1244,1316,1409,1436]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="6" pageNumber="190">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="035336B7FFF1597D8D9E26B09C8A228B" blockId="6.[187,1400,1475,1783]" pageId="6" pageNumber="190">
HEAD. Clypeus with medial anterior margin nearly straight anteriorly; moderately cleft laterally, slanted at 60° angle; above clypeo-labral suture with row of large, rounded to rugose punctures adjacent to posterior margin; each puncture with stiff brownish hair or seta; surface directly above clypeo-labral suture lustrous, glabrous; anterior and lateral margins carinate dorsally, distinctly concave medially; anterior angle widely rounded dorsally; dorsal surface covered with short or long, narrow, elongate, nearly parallel-sided, rounded, apically scales; scales slightly bent downward. Apical maxillary palpomere rice grain-like with spindle-shaped flattened area. Mentum pot-shaped, lustrous, glabrous; with nearly flattened surface; with very few hair brushes on each side of medial cleft subanteriorly. Antennal lamellae length
<quantity id="C4149B52FFF1597D8D9E25AE9E4D228B" box="[189,282,1757,1783]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="190" unit="mm" value="2.9">2.9 mm</quantity>
, distinctly longer than entire length of antennomeres II–VII.
</paragraph>
<paragraph id="035336B7FFF1597C8D9E246D9DF725AF" blockId="6.[189,1400,1822,2025]" lastBlockId="7.[187,1400,265,467]" lastPageId="7" lastPageNumber="191" pageId="6" pageNumber="190">PRONOTUM. Anterior margin widely concave; with short, nearly straight margin medially; anterior angle obtuse, lustrous, glabrous, triangulate, impunctate subapically; with rounded callosity apically; anterolateral margin wedge-shaped at 45° angle, crenulate; medial margin widely rounded, crenulate; posterolateral margin slightly wedge-shaped, smooth; posterior angle distinctly obtuse; slightly sinuate, rounded, apically; above it with small glabrous, impunctate callosity; posterior margin evenly wedge-shaped downward on each side; widely concave, medial margin slightly extended posteriorly; surface with very narrow impunctate longitudinal medial callosity, disappearing subanteriorly and subposteriorly; surface covered with variable scales; covered mostly with elongate, ovoid, medially stout, rounded apically scales medio-subanteriorly; covered mostly with elongate, spindle-shaped, rounded apically scales medio-subposteriorly; covered mostly with elongate, nearly parallel-sided, inverted-lanceolate, rounded apically scales submedially; covered mostly with elongate, inverted lanceolate, widely rounded apically scales adjacent to lateral margin.</paragraph>
<caption id="5793663FFFF0597C8D9E24D89B9C2395" ID-DOI="http://doi.org/10.5281/zenodo.8306340" ID-Zenodo-Dep="8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="7" pageNumber="191" startId="7.[189,232,1963,1989]" targetBox="[250,1398,568,1908]" targetPageId="7">
<paragraph id="035336B7FFF0597C8D9E24D89B9C2395" blockId="7.[189,1399,1963,2025]" pageId="7" pageNumber="191">
<emphasis id="3198EAA5FFF0597C8D9E24D89E6023B9" bold="true" box="[189,311,1963,1989]" pageId="7" pageNumber="191">Fig. 7–10.</emphasis>
<taxonomicName id="C4EC4D34FFF0597C8C1C24D89D7823B9" authority="Calcetas, 2023" authorityName="Calcetas" authorityYear="2023" box="[319,559,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="191" phylum="Arthropoda" rank="species" species="bezdeki" status="sp. nov.">
<emphasis id="3198EAA5FFF0597C8C1C24D89D7823B9" box="[319,559,1963,1989]" italics="true" pageId="7" pageNumber="191">Leucopholis bezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AAB57DEFFF0597C8F1B24DF9DC223BA" box="[568,661,1964,1990]" pageId="7" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="DC578815FFF0597C8F8124D89C5B23B9" box="[674,780,1963,1989]" pageId="7" pageNumber="191" type="holotype">holotype</typeStatus>
,
<specimenCount id="15EAFD3EFFF0597C8E3B24DF9C7923BA" box="[792,814,1964,1990]" pageId="7" pageNumber="191" type="male">♂</specimenCount>
(PNM).
<emphasis id="3198EAA5FFF0597C8EBA24D89CFF23B9" bold="true" box="[921,936,1963,1989]" pageId="7" pageNumber="191">7</emphasis>
. Habitus, dorsal aspect.
<emphasis id="3198EAA5FFF0597C89E824D89B8E23B9" bold="true" box="[1227,1241,1963,1989]" pageId="7" pageNumber="191">8</emphasis>
. Metaventral process, dorsal aspect.
<emphasis id="3198EAA5FFF0597C8CE824BD9E8D2394" bold="true" box="[459,474,1998,2024]" pageId="7" pageNumber="191">9</emphasis>
. Male genitalia, dorsal aspect.
<emphasis id="3198EAA5FFF0597C8E6524BD9C342394" bold="true" box="[838,867,1998,2024]" pageId="7" pageNumber="191">10</emphasis>
. Male genitalia, lateral aspect.
</paragraph>
</caption>
<paragraph id="035336B7FFFF59738D9E227A9EDF2647" blockId="8.[187,1401,265,571]" pageId="8" pageNumber="192">
VENTRAL SIDE OF THORAX. Prosternal process mound-shaped to nearly isosceles triangulate; wedge-shaped subapically; widely rounded apically; with flattened impunctate area medially and apically; with very few scales and hairy on each side and posteriorly. Metaventral process length
<quantity id="C4149B52FFFF59738975223C9BE32516" box="[1110,1204,335,362]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="192" unit="mm" value="4.6">4.6 mm</quantity>
; distinctly wide, triangulate basally, elongate towards apex; not constricted subapically; subapical lateral margin distinctly long, nearly parallel-sided; moderately rounded apically; lustrous, glabrous, impunctate dorsally; each side covered with scales of variable shapes and sizes along each depression; with very narrow, faint, longitudinal dark line dividing metaventral process and metasternum medially; faint line much distinct in metasternum (
<figureCitation id="9BD72A32FFFF59738C41228D9EF02664" box="[354,423,510,536]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="8" pageNumber="192">Fig. 8</figureCitation>
). Metasternum with shallow medial depression, covered mostly with short, inverted lanceolate scales.
</paragraph>
<paragraph id="035336B7FFFF59738D9E21109A542601" blockId="8.[189,1283,611,637]" box="[189,1283,611,637]" pageId="8" pageNumber="192">LEG. Foretibia bidentate; with narrow rounded apex; posterior metatibia with 22–24 spicules.</paragraph>
<paragraph id="035336B7FFFF59738D9E21D79CB5269D" blockId="8.[189,1397,676,737]" pageId="8" pageNumber="192">SCUTELLUM. Covered with large and small, elongate, inverted lanceolate, ovoid, nearly parallel-sided scales; lustrous, impunctate, without scales around posterior margin.</paragraph>
<paragraph id="035336B7FFFF59738D9E207B9BB527AD" blockId="8.[187,1400,776,978]" pageId="8" pageNumber="192">ELYTRON. Anterior margin distinctly wedge-shaped towards scutellum; anterior angle widely rounded; thickly carinate, explanate; with narrow suture towards anterior angle; margin evanescent medially; posterolateral margin distinctly infleXed towards posterior angle; posterior angle obtuse, widely rounded; posterior margin weakly explanate, not carinate; wedge-shaped laterally; sutural angle widely obtuse; sutural margin towards sutural angle and scutellum distinctly carinate, covered with minute fringed hairs; surface covered mostly with short, inverted lanceolate, rounded to truncate apically scales.</paragraph>
<paragraph id="035336B7FFFF59738D9E208A9A25204A" blockId="8.[187,1398,1017,1078]" pageId="8" pageNumber="192">ABDOMEN.Abdominal sternites I–IV covered with short, ovoid lanceolate, widely rounded basally, rounded apically scales; abdominal sternite V covered mostly with elongate lanceolate to elongate ovoid scales.</paragraph>
<paragraph id="035336B7FFFF59738D9E272E9A26209C" blockId="8.[187,1401,1117,1248]" pageId="8" pageNumber="192">PYGIDIUM. Anterolateral margin thickly carinate towards anterior angle; disappearing medially; with narrow explanate margin medially, very distinct and wide towards posterior margin; posterior margin slightly concave, narrowly explanate, distinctly depressed medially; convex laterally; subposterior margin vertically flattened laterally; surface covered mostly with minute, elongate ovoid, parallel-sided scales.</paragraph>
<paragraph id="035336B7FFFF59738D9E267B9BE721AD" blockId="8.[187,1400,1287,1489]" pageId="8" pageNumber="192">
MALE GENITALIA. Genitalia
<quantity id="C4149B52FFFF59738CD1267B9D01215E" box="[498,598,1288,1314]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="8" pageNumber="192" unit="mm" value="14.9">14.9 mm</quantity>
long. Phallobase dorsal apical margin bisinuate, W-shaped, moderately convex medially, distinctly concave on each side and slightly convex towards lateral margin (
<figureCitation id="9BD72A32FFFF5973883826599A322139" box="[1307,1381,1322,1349]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="8" pageNumber="192">Fig. 9</figureCitation>
). Paramere dorsal basal margin bowl-shaped, widely concave medially; distinctly extended posteriorly; each side sinuate. Apical process length
<quantity id="C4149B52FFFF59738FB926039DA021F7" box="[666,759,1392,1419]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="192" unit="mm" value="3.6">3.6 mm</quantity>
, distinctly shorter than phallobase; distinctly elongate, bird bill-like, tapered towards apex; rounded apically; anterior posterior margin widely rounded. Lateral margin of paramere with wide, shallow longitudinal depression on each side (
<figureCitation id="9BD72A32FFFF5973896826C49BF421AD" box="[1099,1187,1463,1489]" captionStart="Fig" captionStartId="7.[189,232,1963,1989]" captionTargetBox="[250,1398,568,1908]" captionTargetPageId="7" captionText="Fig. 7–10. Leucopholis bezdeki sp. nov., holotype, ♂ (PNM). 7. Habitus, dorsal aspect. 8. Metaventral process, dorsal aspect. 9. Male genitalia, dorsal aspect. 10. Male genitalia, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.8306340" httpUri="https://zenodo.org/record/8306340/files/figure.png" pageId="8" pageNumber="192">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="035336B7FFFF59738D9E268B9DF62207" blockId="8.[187,1399,1528,1659]" pageId="8" pageNumber="192">
FEMALE. Length 41.0–42.0 mm, width
<quantity id="C4149B52FFFF59738F58268B9C70226E" box="[635,807,1528,1555]" metricMagnitude="-2" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.05" metricValueMin="1.98" pageId="8" pageNumber="192" unit="mm" value="20.15" valueMax="20.5" valueMin="19.8">19.8–20.5 mm</quantity>
. Clypeal anterior margin strongly convex medially, anterior angle widely rounded dorsally; medial cleft slanted at 60° angle laterally. Lamellae length
<quantity id="C4149B52FFFF5973886125689E4E2225" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="8" pageNumber="192" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3– 2.5 mm</quantity>
, nearly as long as entire length of antennomeres II–VII. Metaventral process length
<quantity id="C4149B52FFFF597389C0254D9A242225" box="[1251,1395,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="8" pageNumber="192" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5–4.8 mm</quantity>
. Posterior metatibiae with 26–34 spicules.
</paragraph>
</subSubSection>
<subSubSection id="4BF6653CFFFF59738D9E25D19E8C2290" box="[189,475,1698,1772]" pageId="8" pageNumber="192" type="distribution">
<paragraph id="035336B7FFFF59738D9E25D19E0122C0" blockId="8.[189,342,1698,1724]" box="[189,342,1698,1724]" pageId="8" pageNumber="192">
<heading id="581B81DBFFFF59738D9E25D19E0122C0" bold="true" box="[189,342,1698,1724]" fontSize="11" level="3" pageId="8" pageNumber="192" reason="3">
<emphasis id="3198EAA5FFFF59738D9E25D19E0122C0" bold="true" box="[189,342,1698,1724]" pageId="8" pageNumber="192">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="035336B7FFFF59738D9E25A29E8C2290" blockId="8.[189,475,1745,1772]" box="[189,475,1745,1772]" pageId="8" pageNumber="192">
<collectingCountry id="7BFB7627FFFF59738D9E25A29E162297" box="[189,321,1745,1771]" name="Philippines" pageId="8" pageNumber="192">Philippines</collectingCountry>
(Mindanao).
</paragraph>
</subSubSection>
</treatment>
</document>
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